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11 

U563 
CRLSSI 



SMITHSONIAN INSTITUTION 

UNITED STATES NATIONAL MUSEUM 

Bulletin 199 



AN ANNOTATED CHECKLIST AND KEY 

TO THE REPTILES OF MEXICO 

EXCLUSIVE OF THE SNAKES 



BY 

HOBART M. SMITH and EDWARD H. TAYLOR 



UNITED STATES 

GOVERNMENT PRINTING OFFICE 

WASHINGTON : 1950 



For sale by the Superintendent of Documents, U. S. Government Printing Office 
Washington 25, D. C. - Price 75 cents (Paper) 



ADVERTISEMENT 

The scientific publications of the National Museum include two 
series, known, respectively, as Proceedings and Bulletin. 

The Proceedings series, begun in 1878, is intended primarily as a 
medium for the publication of original papers, based on the collections 
of the National Museum, that set forth newly acquired facts in biology, 
anthropology, and geology, with descriptions of new forms and re- 
visions of limited groups. Copies of each paper, in pamphlet form, 
are distributed as published to libraries and scientific organizations 
and to specialists and others interested in the different subjects. The 
dates at which these separate papers are published are recorded in the 
table of contents of each of the volumes. 

The series of Bulletins, the first of which was issued in 1875, con- 
tains separate publications comprising monographs of large zoological 
groups and other general systematic treatises (occasionally in several 
volumes), faunal works, reports of expeditions, catalogs of type 
specimens, special collections, and other material of similar nature. 
The majority of the volumes are octavo in size, but a quarto size has 
been adopted in a few instances in which large plates were regarded 
as indispensable. In the Bulletin series appear volumes under the 
heading Contributions from the United States National Herbarium, in 
octavo form, published by the National Museum since 1902, which 
contain papers relating to the botanical collections of the Museum. 

The present work forms No. 199 of the Bulletin series. 

Alexander Wetmore, 
Secretary, Smithsonian Institution. 

II 



CONTENTS 



Page 

Introduction. 

Itinerary and gazetteer of localities for the Walter Rathbone Bacon 

Expedition, 1938-1941 4 

Class Reptilia ^^ 

Subclass Anapsida 

Order Testudines ^2 

Suborder Atheca 1^ 

Family Dermochelyidae 13 

Genus Dermochelys 13 

Suborder Carettoidea. 14 

Family Cheloniidae 1'^ 

Genus Lepidochelys 14^ 

Genus Caretta IS 

Genus Eretmochelys 16 

Genus Chelonia 1"^ 

Suborder Trionychoidea 1^ 

Family Trionycbidae 18 

Genus Amyda 18 

Suborder Cryptodira 18 

Family Dermatemyidae 19 

Genus Dermatemys 19 

Family Chelydridae 20 

Genus Chelydra 20 

Family Kinosternidae 21 

Genus Kinosternon 21 

Genus Claudius 26 

Genus Staurotypus 27 

Family Testudinidae 27 

Genus Gopherus 28 

Family Emyidae 28 

Genus Geoemyda 29 

Genus Pseudemys 31 

Genus Chrysemys 33 

Genus Terrapene 34 

Genus Malaclemys 36 

Genus Clemmys 37 

Subclass Lepidosauria 37 

Order Squamata.. ^' 

Suborder Amphisbaenia 37 

Family Bipedidae 37 

Genus Bipes 38 

rn 



IV BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Class Reptilia — Continued 

Subclass Lepidosauria — Continued 

Order Squamata — Continued Page 

Suborder Sauria 39 

Family Gekkonidae 40 

Genus Coleonyx 41 

Genus Gonatodes 45 

Genus Phyllodadylus 46 

Genus Thecadadylus 49 

Genus Hemidadylus 49 

Genus Aristelliger 51 

Genus Peropus 51 

Genus Sphaerodadylus 52 

Family Iguanidae 53 

Genus Anolis 55 

Genus Corythophanes 68 

Genus Laemandus 69 

Genus Basiliscus 71 

Genus Iguana 72 

Genus Ctenosaura 73 

Genus Enyaliosaurus 75 

Genus Dipsosaurus 77 

Genus Sauromalus 78 

Genus Holbrookia 81 

Genus Callisaurus 85 

Genus Uma 89 

Genus Petrosaurus 90 

Genus Streptosaurus 91 

Genus Crotaphytus 91 

Genus Gambelia 93 

Genus Phrynosoma 94 

Genus Sceloporus 105 

Genus Sator 139 

Genus Urosaurus 140 

Genus Uta 147 

Family Xantusiidae 151 

Genus Lepidophyma 151 

Genus Gaigeia 153 

Genus Xantusia 154 

Family Scincidae 155 

Genus Mabuya 155 

Genus Scincella 156 

Genus Eumeces 160 

Family Anelytropsidae 170 

Genus Anelytropsis 170 

Family Teiidae 170 

Genus Ameiva 170 

Genus Cnemidophorus 174 

Genus Gymnophthabnus 192 

Family Helodermidae 192 

Genus Heloderma 192 



CONTENTS V 

Class Reptilia — Continued 

Subclass Lepidosauria — Continued 
Order Squamata — Continued 

Suborder Sauria — Continued Page 

Family Anguidae 194 

Genus Celestus 194 

Genus Abronia 196 

Genus Barisia 198 

Genus Gerrhonotus 203 

Genus Elgaria 205 

Family Xenosauridae 207 

Genus Xenosaurus 207 

Family Anniellidae 208 

Genus Anniella 209 

Suborder Serpentes.' 

Subclass Archosauria 209 

Order Loricata 209 

Family Crocodylidae 210 

Genus Crocodylus 210 

Family Alligatoridae 211 

Genus Caiman 212 

Species inquirendae 212 

State lists 215 

Index 231 

' See U. S. Nat. Mus. BuU. 187, 1945, p. 12. 



AN ANNOTATED CHECKLIST AND KEY TO 
THE REPTILES OF MEXICO, EXCLUSIVE 
OF THE SNAKES 



By HoBART M. Smith and Edward H. Taylor 



INTRODUCTION 

This volume constitutes the last of a series of checklists and keys 
to the herpetological fauna of Mexico. The first, ''An Annotated 
Checklist and Key to the Snakes of Mexico" (U. S. Nat. Mus. Bull. 
187), was published on October 5, 1945. The second, "An Annotated 
Checklist and Key to the Amphibia of Mexico" (U. S. Nat. Mus. 
Bull. 194), appeared on June 17, 1948. In this, the third volume, the 
turtles, amphisbaenians, lizards, and crocodilians are treated. The 
preparatory work that preceded these checklists was pursued as time 
permitted from 1932 to the present, a period of about 17 years. It 
has involved a number of expeditions to Mexico at our own expense ^ 
and the collecting of more than 50,000 herpetological specimens. 

Except in rare instances the materials have all passed under the 
scrutiny of one or the other of us, and the detailed results of these 
studies are embodied in more than a hundred short or longer reports. 
Our own ideas have often changed with the acquisition of materials 
not originally available. 

It was hoped that the long-awaited work on turtles by Stejneger 
and Hartweg w^ould see the light before the appearance of the turtle 
checklist, but since there is still no certainty of the completion of this 
work in the immediate future it does not seem wise to delay on this 
account. 

We have refrained from anticipating certain forms not now recog- 
nized that are present even in our own collection; the specimens repre- 
senting them are referred without comment to the recognized form 
they most closely resemble. A number of such forms belong to Anolis, 
no adequate revision of which has yet been made, although the senior 
author is contemplating such a work. Certain other groups, especially 
Cnemidophorus and Uta, merit reinvestigation. 

There are many other unsolved problems in Mexican herpetology; 
much more exploration remains to be performed. Probably no single 

' From 1938 to 1941 1 was financed by the Smithsonian Institution on several expeditions, through tenure 
of a Walter Rathbone Bacon Traveling Scholarship.— H. M. S. 

1 



BULLETIN" 199, UNITED STATES NATIONAL MUSEUM 



problem of distribution is more demanding of explanation than the 
extreme dearth of Plethodontidae in the Guerrero "island" and in the 
high western plateau, and the apparent replacement of that family in 
the latter region, largely by the Ambystomidae. 

The treatment of forms in this volume differs little from that of the 
two preceding volumes, except that the name of the collector of each 
type is added. This name often is as important perhaps as the name 
of the author of a species. Unfortunately, the collector is frequently 
unknown or uncertain. This is true of the collections from Mexico 
studied by Wiegmann, in which case we have given Deppe credit 
for obtaining most of the reptiles. The other two men constituting 
the collecting party, Count Von Sack and Schiede, were, we believe, 
engaged primarily in the collection of orchids. However, it is not 
at all improbable that some forms accredited to Deppe were actually 
taken by one of the others. 

We refrain here from considering the implications that may be 
deduced from a perusal of the combined data of the three volumes, 
such as an investigation of faunal origins, lines of migration, relative 
age of taxonomic groups, and faunal areas of ecological relationships. 
However, we are fully aware that such consideration would doubtless 
yield valuable information. 

The number of forms recognized in the herpetofauna of Mexico, 
including those forms described since the fii'st of these checldists was 
published, is relatively large. The following table represents the 
numbers in the various groups: 



Table 1. — Number of forms recognized 


in the herpetofauna of Mexico 


Order or suborder 


Families 


Genera 


Species 
and sub- 
species 


Gymnophiona __ __ . 


1 
4 

7 

8 

1 

10 

8 
2 


2 

16 
25 
18 

1 
47 
80 

2 


2 


Caudat a _. _ ._ _.. 


65 


Saiientia _- - -_ 


163 


Testudines _ _ _ 


49 


Amphisbaenia 


3 


Sauria __ 


394 


Serpentes _ __ . _- 


486 


Loricata _ 


3 






Total . 


41 


191 


1,165 







That there will in the future be considerable shifting of species to 
the subspecific category, and changing forms we regard as subspecies 
to the species category, is to be anticipated. On occasion we have 
found even ourselves at variance on certain problems of this sort. 
We both feel, however, that the currently and deservedly popular 
trend toward recognition of subspecies where they exist has brought 



CHECKLIST OF REPTILES OF MEXICO 3 

with it, almost inevitably, occasional overzealous reduction of species 
to subspecies. Such changes are made mthout due demand for facts, 
but instead with speculations that often merely serve as an excuse to 
tamper with an accepted arrangement that may be equally as plausible. 

One of the most exasperating tasks has been the unraveling of certain 
purely nomenclatorial knots. Several names, for example, are of 
controversial orthography. Worthy of special mention are Agkistro- 
don vs. Ancistrodon, Cnemidophorus tessellatus vs. C. tesselatus, Cory- 
tophanes vs. Corythophanes, and Kinosternon vs. Cinosternon. Opinion 
3G of the International Commission on Zoological Nomenclature 
interprets the permission given by Article 19 to correct any "error 
of transcription" to mean also "error of transliteration," whether 
the original author was aware of his error or not. By such procedure, 
the names Ancistrodon, tessellatus, Corythophanes, and Cinosternon 
would be recommended. On the other hand, Moore, Weller, and 
Knight (Journ. Paleont., vol. 16, 1942, pp. 250-261) maintain with 
excellent reason that only in very clear-cut and exceptional cases is 
any modification of the original orthography of a generic name 
justified. Their view is strengthened by Blackwelder, Knight, and 
Sabrosky (Science, vol. 108, 1948, pp. 37-38). Maintenance of original 
form would perpetuate Agkistrodon, tesselatus, Kinosternon, and 
Corytophanes. Since the proper procedure is not universally or even 
generally agreed upon, we see little advantage in deviation from 
currently accepted form untU some means of real standardization is 
provided by the International Commission on Zoological Nomen- 
clature. 

For the first time in these checklists we here adopt the practice of 
placing a comma between scientific name and author in citations of all 
references except the original. 

We wish to acknowledge the assistance of W. Leslie Burger in the 
laborious task of checking and rechecking various points in the 
manuscript, and in criticizing certain parts, and of Dr. Doris M. 
Cochran, zoologist, division of reptiles and amphibians. United States 
National Museum, who devoted much time to curatorial duties 
connected with the voluminous collection and in making available for 
study the National Museum herpetological collections. We are grate- 
ful likewise for the corrections and improvements suggested by Dr. 
D. F. Hoffmeister, Karl P. Schmidt, and Dr. Frederick Shannon. The 
Graduate Research Boards of the University of Illinois and the 
University of Kansas have generously provided financial support for 
research and clerical work involved in completion of the present 
project, and the Walter Rathbone Bacon Scholarship of the Smith- 
sonian Institution furnished a substantial sum for travel and research 
in Mexico. 



4 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

ITINERARY AND GAZETTEER OF LOCALITIES FOR THE WALTER 
RATHBONE BACON EXPEDITION, 1938-1941 

As previously stated, the senior author was enabled to collect in 
and study material from Mexico by tenure of a Walter Rathbone 
Bacon Traveling Scholarship from 1938 to 1941. The collection 
obtained during those years has formed an important addition to 
other material from Mexico and merits an elaboration of the routes 
traversed by the expedition and the localities at which specimens 
were secured. Such an itinerary was, as a matter of fact, to have 
accompanied a study of the lizards of the Bacon collection, as stated 
in our introduction to the summary of the amphibians (Proc. U. S. 
Nat. Mus., vol. 95, 1945, p. 521). It now seems unlikely that a com- 
plete survey of the Bacon Hzards can appear for a number of years, 
for a satisfactory completion of such a study involves revision of a 
number of very sketchily understood genera, such as Anolis and 
Cnemidophorus. In the meantime a need for an account of the Bacon 
itinerary and localities continues to exist. We have, accordingly, 
taken this opportunity to present this information in this the final 
checklist. 

The expedition personnel consisted of the senior author and his wife. 
They received hearty support in field work from a number of indi- 
viduals, through whose combined efforts about 22,000 specimens were 
secured during a period of two years. As closel}'' as can now be deter- 
mined (the lizards not yet having been fully studied), 500 species and 
subspecies (146 amphibians, 160 lizards, 1 amphisbaenian, 170 snakes, 
20 turtles, 3 crocodilians) were secured. 

Among those who contributed to the success of the expedition are 
Dr. Alexander Wetmore, who saw to it that the expedition was prop- 
erly planned and could function smoothly; Dr. Doris M. Cocliran, 
who must have nearly equaled the expedition personnel in time ex- 
pended for the collection, inasmuch as the laborious task of cataloging 
the specimens rested with her; Dr. Linton P. Satterthwaite, who 
provided the facilities for our stay at Piedras Negras in the midst of a 
Httle-known, remote, and austere area; Mr. and Mrs. Dyfrig McH. 
Forbes, who provided for nearly two years a base of operations in 
Veracruz and who were a constant source of companionship, inspira- 
tion, and material; Eizi Matuda, who very generously provided facil- 
ities for a two-month stay at his finca and who sympathetically aided 
us in every possible way to sample the herpetofauna of the area as 
thoroughly as possible; Thomas MacDougall, who in Tehuantepec 
secured for us numerous specimens and arranged for our travel into 
areas otherwise difficult of access; Ernest Rateike, of Palenque, 
Chiapas, who accepted us as a guest in his home for a month: and a 



CHECKLIST OF REPTILES OF MEXICO 5 

host of local residents, far too numerous to mention, who materially 
augmented our collections and made our visits pleasant and successful. 

Inasmuch as we traveled chiefly by car, our routes are best followed 
on a road map of Mexico. We entered Mexico on October 5, 1938, at 
Ciudad Juarez, Chihuahua, and followed the Chihuahua-El Paso 
highway southward to Vado (October 9). We then turned back to 
Ahumada and followed a poor trail through Carrizal and Rancho 
Nuevo to Progreso (October 10). We camped beside the Rio Santa 
Maria about a mile southwest of Progreso for 5 days, interrupted only 
by one hurried round trip to Casas Grandes for the benefit of a snake- 
bitten cowboy. At this camp we were aided greatly by 10 or 15 
cowboys who brought in material as fast as it could be preserved. 
Our specimen containers full, we determined to go to the border to 
ship specimens and hkewise to obtain higher wheels for the car (a 
half-ton panel truck), with which we had encountered numerous diffi- 
culties because of deep ruts and high centers. We left the Progreso 
camp on October 15, passing through Casas Grandes and Ascencidn, 
and reached Las Palomas the same day. After two days in Columbus 
and Deming, N. Mex., we retraced our steps (October 20) from Las 
Palomas to Casas Grandes. As the weather had become cool, we 
continued southward along the road through San Buenaventura and 
Carmen to the main El Paso-Chihuahua highway, thence southward 
to Ciudad Chihuahua. We left there on October 27 for Torredn, 
arriving October 29. We collected in the vicinity of Torre6n October 
30 and 31, and on November 1 started for Ciudad Durango. We 
could get no farther than Pedricena, however, and, after exploring 
with little success a side road to Nazas, returned to Torre6n on Novem- 
ber 4, We continued to San Pedro the next day and collected in the 
vicinity of that town until November 10. After a side trip to Parras 
off the Torredn-Saltillo highway, we made an attempt to find Jaral, 
Coahuila, a locality made famous zoologically by Heller and Barber. 
This turned out to be an abandoned ranch near Hip61ito, and, as no 
likely habitat for the montane species recorded from Jaral appeared 
to be nearer than 20 miles or so, we continued to Saltillo (November 
14). We collected in the vicinity of Saltillo, Arteaga, and Mount 
Zapaliname until November 17, when we left for Monterrey. 

On November 18 we arrived at Hacienda La Clementina (near 
Forl6n), and we remained there until November 28, when we drove 
to Laredo to ship another lot of specimens. We returned to La 
Clementina on December 3 and left December 5 for Mexico City. 
We stayed at Huichihuayan for several days, leaving December 9. 
Our route, with brief stops at or near various towns, then led through 
Mexico City to Tehuacan, Puebla, thence through Orizaba to Potrero 
Viejo, Veracruz, where we arrived on December 11. 



6 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Potrero Viejo remained our headquarters for over a month. We 
made numerous short trips during that time; especially noteworthy 
were those to Palma Sola (on the Veracruz-Orizaba highway), to 
Tezonapa, Veracruz, and Cosolapa, Oaxaca, and to Cuautlapan, 
Orizaba, C6rdoba, Acultzingo, and other localities along the C6rdoba- 
Tehuacan road. We left Potrero Viejo on January 16, 1939, and 
after a brief stop in Mexico City made headquarters in Cuernavaca, 
Morelos, for nearly two weeks, with side trips to Puente de Ixtla 
(Morelos), Cacahuamilpa (Guerrero), and Zempoala (Morelos and 
Mexico) . 

Leaving Cuernavaca February 2, we continued toward Acapulco, 
reaching there February 5. We collected in the vicinity of Acapulco, 
with side trips to Coyuca, until February 11. Our route, with fre- 
quent stops, then led northward again to Mexico City, where we 
arrived on March 1. After two days in search of axolotls and other 
ambystomids in the vicinity of Mexico City (with side trips to Texcoco, 
Zumpango, and Chimalhuacan) , we started (March 4) on the road to 
Guadalajara. Our first deviation from this route was on March 9 
and 10, to Patzcuaro. On March 11 we reached Uruapan on another 
side trip and continued southward to Apatzingan, returning to 
Uruapan on March 19. An attempt was made to find Crotalus foly- 
stictus in the marshes of the eastern end of Lake Chapala, but without 
success, since most of the marshes have been drained and are now under 
cultivation. We reached Guadalajara on March 24 and returned to 
Potrero Viejo on March 26. On March 30 we left Potrero Viejo 
enroute to Laredo with another shipment of specimens, arriving at 
Laredo on April 3, 1939. 

After nearly a month in the United States we left Laredo on April 29, 
1939, and arrived in Potrero Viejo on May 2. From there we went 
to Veracruz by train and secured passage for Alvaro Obreg6n, arriving 
on May 13. A river boat was then taken, via Ciudad del Carmen, 
to Tenosique, Tabasco, the head of navigation on the Rio Usumacinta. 
Our final destination, Piedras Negras, Guatemala, was reached on May 
21 after two days by mule. We collected in the immediate vicinity 
of Piedras Negras until June 23, when we left for Tenosique. 

From Tenosique (June 30) we continued downstream to Emiliano 
Zapata and there procured mules for a 2-day trip to San Juanito, a 
ranch half a mile from the village of Palenque. There we remained, 
with a side trip only to the ruins of Palenque several miles distant 
in the hills, until August 6, when we left for Alvaro Obreg6n and, 
immediately thereafter, Potrero Viejo, where we arrived on August 14. 
There we were joined by the junior author and with him left on 
August 18 for Mexico City, where we arrived, via the usual route 
through Tehuac§,n and Puebla, on August 22, after numerous brief 



CHECKLIST OF REPTILES OF MEXICO 7 

stops en route. After a few days spent in the environs of Mexico City 
searching for ambystomids, on August 29 we took the Acapulco road 
and followed it with only one deviation to Tixtla, Guerrero, arriving 
at Acapulco on September 3, We returned to Mexico City by the 
same route, arriving on September 6. Agaui we collected in the 
immediate environs of Mexico City, chiefly investigating montane 
faunas. On September 10 we left for Guadalajara but turned back 
near Sahuayo. We then, on September 15, started northward on the 
Pachuca Road to El Chico National Park in Hidalgo. After one day 
there we returned, September 18, to Mexico City, where Dr. Taylor 
left us. 

We then returned, September 19, to Potrero Viejo. After a few 
local trips we again returned to the Guadalajara road, leaving Sep- 
tember 29. We took the side road to the Nevado de Toluca (October 
2) and then retraced our steps to Mexico City, arriving October 3. 
After a few days spent in local trips, we turned northward to Laredo 
with another load of specimens, arriving at Laredo October 17. Our 
only side trip enroute was to Galeana, Nuevo Le6n, via the road from 
Linares, Tamaulipas (October 13, 14). 

After nearly two months in the United States, we again crossed the 
border at Laredo on December 10, 1939, and headed directly for 
Potrero Viejo, Veracruz, arriving December 15. After a few local 
trips near Potrero Viejo, we left (December 27) for Tehuantepec by 
rail. We arrived on December 30 and remained until January 28, 
1940. A number of side trips were taken, by truck to Cerro Arenal, 
by rail to Matias Romero, Oaxaca, by rail to Salina Cruz, and by rail 
to Tonala. Illness necessitated return to Potrero Viejo and ulti- 
mately to Mexico City. We remained there until March 15, A\hen we 
returned to Potrero Viejo. On March 17 we started on a side trip, 
lasting until March 24, which led by highway through Tehuacan, 
Tecamachalco, thence northeastward on the Jalapa road, to Teziutlan 
(Puebla) from El Limdn, Totalco, and eastward to Puente Nacional. 

Shortly thereafter we left for Tehuantepec by rail again, arriving 
April 1. We stayed only one day and traveled by rail to Acapetahua, 
whence we went by bus to Escuintla. From there pack animals were 
used to transport our equipment to La Esperanza, a finca about 5 
miles northeast of Escuintla, We arrived there on April 4 and re- 
mained until June 5, 1940. Various short side trips were taken to 
nearby fincas. 

On June 5 we returned to Tehuantepec and, after one day, to 
Potrero Viejo, arriving June 9. About one month later, on July 6, we 
crossed the border at Laredo with no deviations from the direct route 
from Potrero Viejo to the border. 



8 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

We returned immediately to Mexico City by way of the Pan 
American Highway and remained there until August 7, except for a 
trip by rail to Guanajuato, Guanajuato, on July 19-21. On August 8 
we again drove to Potrero Viejo. Within a few days we made a brief 
and final foray along the highway from Acultzingo to Tehuacan before 
packing all equipment that had been stored at Potrero Viejo, We 
left our headquarters there for the final time on August 8 and, with 
but a brief stop in Mexico City, traveled slowly noithward along the 
Pan American Highway, with a heavy load, crossing the border at 
Laredo on August 24. 

The following gazetteer includes only those localities represented by 
specimens in the Bacon collection that are not to be found on the 
National Geographic Society's 1939 map of Mexico, Central America, 
and the West Indies, scale 1 : 5,702,400. Most of the localities are 
plotted on other, larger-scale maps, but inasmuch as these are not 
always readily available we include all missing from the Geographic 
Society's map, which is readily available to all investigators. The 
localities are arranged alphabetically within the states, which are 
themselves alphabetically arranged. 

CHIAPAS 

Acacoyagua: 3 miles northwest of Escuintla. 

Ag^uacate: 7 miles north-northeast by north of Palenque. 

Belen: 20 miles southeast of Escuintla. 

Cerro Obando: 3 miles northeast of La Esperanza (which see). 

Colonia Hidalgo: 7 miles northeast of Acacoyagua (which see). 

Colonia Soconusco: 12 miles northeast of La Esperanza (which see). 

Cruz de la Piedra: 2 miles west of La Esperanza (which see). 

Finca JuS.rez: 10 miles northeast of La Esperanza (which see). 

Javarinero: 13 miles east of Palenque. 

La Esperanza: 3 miles east of Acacoyagua (which see). 

La Magnolia: 3 miles northwest of La Esperanza (which see). 

Las Nubes: On Cerro Obando, 2 miles northeast of La Esperanza (which see). 

Motozintla: 25 miles east-northeast by east of Escuintla. 

Bancho Las Gradas: 2 miles west of La Esperanza (which see). 

Salto de Agua: On Cerro Obando, 1 mile northeast of La Esperanza (which see). 

San Juanito: 1 mile east-northeast of Palenque. 

Santa Bosa: Near Comitdn. 

CHIHUAmjA 

Carmen: 32 miles east-northeast by east of Galeana. 

Carrizal: 10 miles west-southwest by south of Ahumada. 

Ciudad Delicias: 5 miles south of Meoqui. 

Ojo de Federico: A small ranch 8 miles southeast of Ascenci6n. 

Progreso: 35 miles northeast of Galeana, near Rfo Santa Maria. 

Rancho Nuevo: A small ranch 10 miles west-northwest by west of Carrizal. 

Samalayuca: 25 miles south of Ciudad Judrez. 



CHECKLIST OF REPTILES OF MEXICO 
COAHUILA. 

Hip61ito: 25 miles north-northeast of General Cepeda. 
Zapaliname: A mountain 3 miles south of Saltillo. 

DISTRITO FEDERAL 

Atzacoaico: 10 miles northeast of Mexico City. 
Cafiada de Contreras: 5 miles southeast of Mexico City. 

DURANGO 

La Qoma: 12 miles west of Lerdo, on the south side of the Rfo Nazas. 
La Loma: 12 miles west of Lerdo, on the north side of the Rfo Nazas. 
Pedricefla: 25 miles east-southeast of Nazas. 

GUATEMALA 

Desempefia: 4 miles southeast of Piedras Negras. 

Pozo de la Jicotea: 2 miles southeast of Piedras Negras. 

GUERRERO 

Agua Bendita: 13 miles south of Taxco. 

Agua del Obispo: 8 miles north of Dos Caminos. 

Cacahuamilpa: 10 miles southeast of Tetecala, Morelos. 

Cojruca: 7 miles northeast of Acapulco. 

El Treinta: 20 miles north of Acapulco. 

Garrapatas: 5 miles southwest of Dos Caminos. 

Julian Blanco: 4 miles north of Dos Caminos. 

Ocotito: 3 miles north of Dos Caminos. 

Omilteme: 3 miles northwest of Chilpancingo. 

Paso del Limonero: 9 miles northeast of Acapulco. 

Tierra Colorada: 3 miles southwest of Garrapatas (which see). 

Xaltianguis: 31 miles northeast of Acapulco. 

HIDALGO 

Barranca de los Horcones: 6 miles south of Durango (which see). 

Chapxilhuacan: 19 miles southwest of Tamazunchale, San Luis Potosl. 

Colonia: 6 miles southeast of Pachuca. 

Durango: 15 miles west-southwest by south of Jacala. 

El Chico Parque Nacional: 15 miles northeast of Pachuca. 

Maguey Verde: 7 miles south of Durango. 

Tianguistengo : 10 miles north-northeast of Zacualtipdn. 

MEXICO 

Chalco: 13 miles northeast of Xochimilco. 

Chimalhuacan: 12 miles east of Mexico City. 

Lerma: 10 miles east of Toluca. 

Nevado de Toluca: 32 miles south-southwest by south of Toluca. 

Rio Frio: 18 miles west-northwest of Texraelucdn, Puebla. 

Santa Magdalena: 15 miles east of Mexico City. 

Zempoala: 6 miles west-southwest of Tres Cumbres, Morelos. 



10 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

michoacAn 

Carapa(n) : 23 miles north of Paracho, 
El Temazcal: 20 miles east of Morelia. 
La Palma: 10 miles north of Sahuayo. 
Puerto Hondo: 23 miles east of Moreha. 
Bancho San Jose: 24 miles east of Morelia. 
Tacicuaro: 5 miles east of Quiroga. 

NUEVO le6n 

Mamulique Pass: 45 miles north of Monterrey. 
Santa Caterina: 7 miles west of Monterrey. 

OAXACA 

Caj6n de Piedra: 12 miles south-southwest of Salina Cruz. 

Cerro Arenal: 16 miles west of Tehuan tepee. 

Cerro de Chipehua: 16 miles southeast of Salina Cruz. 

Cerro de Huamelula: 34 miles west-southwest by south of Tehuantepec. 

Cerro Quengola: 10 miles west-northwest by west of Tehuantepec. 

Cosolapa: 8 miles northwest of AcatMn. 

Coyol: between San Antonio and Las Cruces. 

El Lim.6n: 25 miles southeast of Tehuantepec. 

Escurano: 15 miles west-northwest of Tehuantepec. 

La Concepci6n: 32 miles west of Tehuantepec. 

Lachiguiri: 29 miles northeast of Tehuantepec. 

La Gloria: 8 miles southeast of Chimalapa. 

Las Pilas: 4 miles northwest of Tehuantepec. 

Las Vacas: 40 miles west of Tehuantepec. 

Llano Ocotal: 20 miles west-southwest by west of Tehuantepec. 

Matias Romero: 34 miles north of San Ger6nimo. 

Mixtequilla: 4 miles northwest of Tehuantepec. 

Palmar: 37 miles west-northwest by west of Tehuantepec. 

Pixixi: 6 miles south of Tehuantepec. 

Portillo Quayabo: 16 miles west of Tehuantepec. 

Portillo Las Vacas: 40 miles west of Tehuantepec. 

Portillo Los Nanches: 32 miles northwest of Tehuantepec. 

Rincon San Pedro: 16 miles northeast of Tehuantepec. 

Rio Grande: 20 miles west-southwest by south of Tehuantepec. 

Bio Hondo: 41 miles west-northwest by west of Tehuantepec. 

San Francisco Guichina: 61 miles west-northwest by west of Tehuantepec. 

San Jose Manteca: 61 miles west-northwest by west of Tehuantepec. 

San Mateo del Mar: 14 miles east-northeast by east of Salina Cruz. 

San Pedro Quiechapa: 10 miles west of Yautepec. 

Tenango: 24 miles west-southwest by west of Tehuantepec. 

Tres Cruces: 43 miles west-northwest by west of Tehuantepec. 

Yerba Santa: 10 miles west-northwest by west of Tehuantepec. 

PUEBLA 

Alchichica: 10 miles southwest of Perote, Veracruz. 
Cacaloapam: 8 miles southeast of Tlacotepec. 
El Seco: 27 miles northeast of Tepeaca. 
La Virgin: 2 miles north of Cacaloapam. 



CHECKLIST OF REPTILES OF MEXICO 11 

Ozumbilla: 4 miles east-southeast of Morelos. 

Pajaro Verde: 300 feet west of Puebla- Veracruz state line, about 7 miles south- 
east of Morelos. 
Tecaraachalco: 15 miles northwest of Tlacotepec. 

SAN LUIS POTOSf 

Pujal: 20 miles north of Tancanhuitz. 

TABASCO 

El Retire: 13 miles south of Tenosique. 
Los Rieles: 10 miles south of Tenosique, 
Santo Tomas: 12 miles southeast of Tenosique. 

TAMAULIPAS 

Ciudad Mante (= Villa Judrez): 18 miles east-northeast by north of Antiguo 

Morelos. 
Forlon: 18 miles east-southeast of Llera. 
Hacienda La Clementina: 13 miles east-southeast of Llera. 

TLAXCALA 

Apizaco: 8 miles north-northwest by north of Tlaxcala. 

VERACRUZ 

Acultzingo: 9 miles east-southeast by east of Morelos, Puebla. 
Atoyac: 4 miles east-northeast of Potrero Viejo (which see), 
Cerro Gordo: 20 miles east-southeast of Jalapa. 
Cuautlapan: 4 miles southwest of Fortin (which see). 
Cruz Blanca: 8 miles northeast of Perote. 
El Liraon Totalco: 10 miles southwest of Perote. 

El Maguey: 8 miles east-southeast by east of Potrero Viejo (which see). 
Encero: 8 miles east-southeast of Jalapa. 
Fortin: 5 miles east of C6rdoba. 

Matacabestro: 7 miles north-northwest by north of Joaqufn. 
Mata de Cafia: 18 miles east-southeast of Jalapa. 
Medellin: 10 miles south of Veracruz. 
Metlac : 1 mile west of Fortf n (which see) . 

Ojo de Agua: 2 miles east of Paraje Nuevo (which see), near Potrero Viejo. 
Pan de OUa: 8 miles south of Teziutldn, Puebla. 
Paraje Nuevo: 2 miles east of Penuela (which see). 
Paso del Macho: 5 miles northeast of Atoyac. 
Pefiuela: 4 miles southeast of C6rdoba. 
Potrero Viejo: 2 miles east of Paraje Nuevo (which see). 
Presidio: 22 miles southwest of C6rdoba. 

San Juan de La Punta: 15 miles east-southeast by south of C6rdoba. 
San Jose de Gracia: 13 miles southeast of C6rdoba. 
Palma Sola: 9 miles east-southeast by south of C6rdoba. 
Tequejru tepee: 10 miles northeast of Jalapa. 
Tezonapa: 8 miles northwest of Acatldn. 
Toxtlacuaya: 17 miles northwest of Jalapa. 
Xuchil: 1 mile north of Potrero Viejo (which see). 
861316—60 2 



12 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Class REPTILIA Laurenti 

Reptilia Laurenti, Specimen medicum exhibens synopsin reptilium, 1768, p. 19. 

Subclasses. — Three subclasses are represented by living members. 
All occur in Mexico. 

KEY TO MEXICAN' SUBCLASSES, ORDERS, AND SUBORDERS OF REPTILIA 

1. Provided with a shell encasing body above and below, consisting of plastron and 

carapace; turtles. .subclass Anapsida order Testudines (p. 12)_ 5 

Not provided with a shell 2 

2. Anal slit longitudinal; skull diapsid subclass Archosauria. 

order Loricata (p. 209) 

Anal slit transverse; skull modified diapsid; lower (quadratojugal-quadrate) 

arch interrupted.. subclass Lepidosauria order Squamata (p. 37) 

3. Two limbs only, the forelegs; body ringed with grooves. 

suborder Amphisbaenia (p. 37) 
Four hmbs or none; body not ringed with grooves 4 

4. Limbs present or, if not, movable eyelids present; lizards. 

suborder Sauria (p. 39) 
Limbs absent; movable eyelids absent; snakes suborder Serpentes 

5. No epidermal scutes on shell 6 

Epidermal scutes present 7 

6. Limbs without external evidence of digits, oar-shaped, clawless. 

suborder Atheca (p. 13) 
Limbs with distinct digits, 3 with claws suborder Trionychoidea (p. 18) 

7. Limbs oar-shaped, with 1 to 3 claws suborder Carettoidea (p. 14) 

Limbs not oar-shaped, at least foreleg with 4 or 5 claws. 

suborder Cryptodira (p. 18) 

Subclass Anapsida Williston 

Anapsida Williston, Journ. Geol., vol. 25, 1917, p. 419. 
Orders. — A single living order exists, the Testudines. 

Order TESTUDINES Batsch 

Testudines Batsch, Versuch Anleit. Kenntn. Gesch. Thier. Mineral., vol. 1, 

1788, p. 437. 
Chelonia Macartney, in Ross, Transl. Cuvier's Legons d'anatomie compar^e . . . , 

vol. 1, 1802, tab. 3. 
Testudinata Oppel, Die Ordnungen, Familien und Gattungen der Reptilien . . . , 

1811, p. 3. 

Suborders. — Five living suborders of Testudines are recognized; 
four occur in Mexico, and the fifth (Pleurodira) is restricted to south- 
ern portions of both hemispheres. 

» Characters in all keys apply to Mexican forms but not necessarily to those in other parts of the world. 



CHECKLIST OF REPTILES OF MEXICO 13 

Suborder Atheca Cope 

Athecae Cope, Proc. Amer. Assoc. Adv. Sci., vol. 19, 1870, p. 235. 
Families. — One. 

Family DERMOCHELYIDAE * Fitzinger 

Dermatochelydae Fitzinger, Systema reptilium, 1843, p. 30. 

Genera. — One. 

Range. — Tropical oceans of the world. 

Genus DERMOCHELYS Blalnville 

Dermochelys Blainville, Bull. Soc. Philom., 1816, p. 119. 

Genotype. — Testudo coriacea Linnaeus. 

Range. — World-wide, in tropical oceans and occasionally into tem- 
perate waters. 

Species. — As many as three species (or subspecies) may be valid. 
The species has been recorded in Mexico only from the western coast, 
although Atlantic records are to be expected. 

DERMOCHELYS CORIACEA • (Linnaeus) 

Testudo coriacea Linnaeus, Systema naturae, ed. 12, vol. 1, 1766, p. 350. 
Dermochelys coriacea, Blainville, Bull. Soc. Philom., 1816, p. 119. — Boulenger, 

Catalogue of the chelonians . . ., 1889, p. 10. — Deraniyagala, Tetrapod 

reptiles of Ceylon, vol. 1, 1939, pp. 38-102, figs. 12-34.— Schmidt, Marine 

Life Occ. Pap., vol. 1, No. 3, 1945, pp. 7-10. 
Sphargis coriacea schlegelii Garman, U. S. Nat. Mus. Bull. 25, 1884, p. 303 (type 

unknown; type locality, tropical Pacific and Indian Ocean, here restricted to 

Guaymas, Sonora). 
Dermochelys schlegelii, Stejneger, U. S. Nat. Mus. Bull. 58, 1907, p. 485. 

Type . — Unknown . 

Type locality. — Palermo, Sicily, by present restriction. 

Range. — Pacific coast and probably the tropical Atlantic coast. 
Recorded definitely from Sonora: Guaymas; Baja California: Los 
Coronados Islands. 

' Actually the first family name proposed for this group was Sphargidae Gray (Ami. Philos., vol. 10, 1825, 
p. 212), based upon the genus Sphargis Merrem, 1820, a junior ssmonym of Dermochelys Blainville, 1816; 
both genera are based upon the same tjT)e, Testudo coriacea Linnaeus. Many taxonomists retain the oldest 
family name regardless of the status of its type. We believe there is a certain degree of error courted by 
such a procedure, however, inasmuch as a family name based upon a generic name later found to be a junior 
homonym of an earlier name in another family would, very unfortunately, be preserved; and what if the 
senior homonym had also served as a type for its family? It seems to us that only currently valid generic 
names should be available as types for family names, and that all synonyms and homonyms should be 
regarded unavailable for that purpose. 

• A common arrangement restricts coriacea to the Atlantic, schlegelii to the Pacific, either as species or .sub- 
species. We are unable to find that anything more than geographic probability has led to such arrangement* 



14 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Suborder Carettoidea^ Fitzinger 

Carettoidea Fitzinger, Neue Classification der Reptilien . . ., 1826, p. 5. 
Families .■ — One. 

Family CHELONIIDAE Gray 

Cheloniadae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 212. 
Genera. — Four. 
Range. — Tropical oceans, world-wide. 

KEY TO GENERA OF THE FAMH^Y CHELONIIDAE ' 

1. One pair of prefrontal scutes; costal scutes 4; horny cutting edge of lower jaw 

coarsely dentate, that of upper jaw strongly ribbed vertically; bony alveolar 
ridge of upper jaw with a low but regularly raised auxiliary ridge behind 
anterior ridge, which is very strong and terminates anteriorly in a tooth at 

posterolateral corner of premaxillary pit Chelonia (p. 17) 

Two pairs of prefrontals; costal scutes 4-9; horny cutting edge of lower jaw 
smooth or feebly denticulate, that of upper jaw without markedly elevated 
vertical ribbing on its inner surface; bony alveolar surface of upper jaw 
smooth or with a single ridge; this ridge not terminating anteriorly in a 
sharp tooth 2 

2. Costal scutes 4 pairs; dorsal scutes usually conspicuously imbricate; bony 

alveolar surface of upper jaw with a sharp-crested ridge. 

Eretmochelys (p. 16) 

Costal scutes 5 pairs or more; dorsal scutes not conspicuously imbricate; bony 

alveolar surface of upper jaw smooth or with a rounded ridge 3 

3. Four enlarged inframarginal scutes on bridge; dorsal color gray to olive-green; 

maxillaries not in contact, separated by pre vomer Lepidochelys (p. 14) 

Three enlarged inframarginal scutes on bridge; dorsal color brown or reddish 
brown; maxiUaries in contact between pre vomer and premaxillaries. 

Caretta (p. 15) 

Genus LEPIDOCHELYS Fitzinger 

Lepidochelys Fitzinger, Systema reptilium, fasc. 1, 1843, p. 30. 
Caouana Gray, Catalogue of the tortoises ... in the British Museum, 1844, p. 
52 (type, Chelonia olivacea Eschscholtz). 

Genotype. — Chelonia olivacea Eschscholtz. 
Species. — Two are generally recognized. 
Range: — Tropical oceans, world-wide. 

KEY TO SPECIES OF LEPIDOCHELYS 

1. Inframarginal scutes without pores; usually 5 pairs of costal scutes; color usually 
gray; bony alveolar surface of upper jaw with a conspicuous ridge. 

kempii (p. 15) 

Each inframarginal scute usually with a pore at its posterior border; usually 

more than 5 pairs of costal scutes; color olive; bony alveolar surface of upper 

jaw with a low (not conspicuous) ridge olivacea (p. 15) 



' Deraniyagala (Spolia Zeylanica, vol. 24, 1945, p. 98) suggests that this group may be of pleurodirous 
origin. 
' Adapted from Carr, Proc. New England Zool. Club, vol. 21, 1942, pp. 3-5. 



CHECKLIST OF REPTILES OF MEXICO 15 

LEPIDOCHELYS KEMPH (Gairoan) 

Thalassochelys {Colpochelys) kempii Garman, Bull. Mus. Comp, Zool,, vol. 6, 

1880, p. 123, 
Lepidochelys kempii, Baur, Araer. Nat., vol. 24, 1890, p. 487. — Carr, Proc. 

New England Zool. Club, vol. 21, 1942, pp. 4, 8-13, pis. 2, 4. 

Type. — Several cotypes, Mus. Comp. Zool. 

Type locality. — Gulf of Mexico, here restricted to Key West, Fla. 

Range. — Atlantic Ocean and Gulf of Alexico. In Alexico, the entire 
Atlantic coast. The only exact record known is from Quiatana 
Roo (Isla de Mujeres). 

LEPIDOCHELYS OLIVACEA (Eschscholtz) 

Chelonia olivacea Eschscholtz, Zoologischer Atlas, pt, 1, 1829, p. 2, pi. 3. 
Lepidochelys olivacea, Girard, United States Exploring Expedition . . . , vol. 

20, Herpetology, 1858, p. 435. — Deraniyagala, Tetrapod reptiles of Ceylon, 

vol. 1, 1939, pp. 123-163, figs. 43-65.— Carr, Proc. New England Zool. 

Club, vol. 21, 1942, pp. 4-5, pi. 5 (skull). 
Chelonia dussumieri Di;m:6ril and Bibron, Erp^tologie g6n4rale . . . , 1835, p. 

557 (type in Mus. Hist. Nat. Paris; type locality, Malabar). 
Careita remivaga Hay, Proc. U. S. Nat. Mus., vol. 34, 1908, pp. 194-197 (type, 

U.S.N.M. No. 9973; type locality, Ventosa Bay, Oaxaca). 

Type. — Unknown, 

Type locality. — Manila Bay, Philippine Islands. 

Range. — Indian and Pacific Oceans. In Mexico, the entire Pacific 
coast. Reported only from the states of Chiapas, Oaxaca, Guerrero, 
Colima, and Sonora (Tibur6n Island). 

Genus CARETTA Rafinesque 

Caretta Rafinesqtje, Specchio Sci. Palermo, vol. 2, 1814, p. 66. 

Thalassochelys Fitzinger, Ann. Wien Mus., vol. 1, 1835, p. 121 (type, Testudo 

caouana Daudin). 
Caouana Cocteatt, in Ram6n de la Sagra, Historia ffsica, polftica y natural de 

la Isla de Cuba, vol. 4, Rept., 1838, p. 31 (type, Testudo cephalo Schneider). 
Eremonia Gray, Hand list of shield reptiles of the British Museum, 1873, p. 91 

(type, Caouana elongata Gray). 

Genotyye. — Caretta nasuta Rafinesque=(7areffa caretta caretta (Lin- 
naeus). 

Species.' — One, with two races, is generally recognized. 
Range. — Tropical ocean, world-wide. 

KEY TO SPECIES OF CARETTA 

1. Neural bones usually 9 or more; if fewer, the costal bones usually interrupting 
the neural series by contact with each other at one or more places. 

caretta gigas (p. 16) 

Neural bones 7 or 8, the series rarely interrupted by median contact of costal 

bones caretta caretta (p. 16) 



16 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

CARETTA CARETTA CARETTA (Linnaeus) 

Testudo caretta Linnaeus, Systema naturae, ed. 10, vol. 1, 1756, p. 197. 

Caretta caretta, Stejneger, Ann. Rep. U. S. Nat. Mus., 1902 (1904), p. 715. — 

Care, Proc. New England Zool. Club, vol. 21, 1942, p. 5, pis. 1, 3. 
Caretta caretta caretta, Deraniyagala, Tetrapod reptiles of Ceylon, vol. 1, 1939, 

p. 164; Spolia Zeylanica, vol. 24, 1945, p. 95. 
Testudo cephalo Schneider, Algemeine Naturgeschichte der Schildkroten . . ., 

1783, pp. 303-308 (type unknown; type locality, Charleston, S. C, by 

present restriction) . 
Testudo caouana Daudin, Histoire naturelle des reptiles, vol. 2, 1802, p. 55, pi. 

16, fig. 2 (type in Mus. Hist. Nat. Paris; type locality, Jamaica, by present 

restriction) . 

Type. — Unknown. 

Type locality. — "About the American Islands," here restricted to the 
Bermuda Islands. 

Range. — Western Atlantic Ocean; the entire Atlantic coast of 
Mexico. Reported in Mexico only from Yucatdn. 

CARETTA CARETTA GIGAS Deraniyagala 

Caretta gigas Deraniyagala, Ceylon Journ. Sci., sect. B, vol. 28, 1933, pp. 61-62. 
Caretta caretta gigas, Deraniyagala, Tetrapod reptiles of Ceylon, vol. 1, 1939, 
pp. 164-185, figs. 66-73; Spolia Zeylanica, vol. 24, 1945, p. 95. 

Type. — Presumably in Colombo Museum, Ceylon. 

Type locality. — Ceylon. 

Range. — Indian and Pacific Oceans and possibly eastern Atlantic 
Ocean. In Mexico, the entire Pacific coast. Reported only from 
Sinaloa and Baja California. 

Genus ERETMOCHELYS Fitzinger 

Eretmochelys Fitzinger, Systema reptilium, fasc. 1, 1843, p. 30. 

Genotype. — Testudo imbricata Linnaeus. 

Species. — One is generally recognized, but two species (or sub- 
species) are distinguished by some authorities. 

ERETMOCHELYS IMBRICATA « (LinnaeuB) 

Testudo imbricata Linnaeus, Systema naturae, ed. 12, 1766, p. 350. 

Eretmochelys imbricata, Fitzinger, Systema reptilium, fasc. 1, 1843, p. 30. — 
Agassiz, Contributions to the natural history of the United States, vol. 1, 
1857, p. 381. — Deraniyagala, Tetrapod reptiles of Ceylon, vol. 1, 1939, 
pp. 187-217, figs. 74-85. 

Chelonia lachrymata Cuvier, Le rSgne animal, ed. 2, vol. 2, 1829, p. 13 (no type 
or type locality designated; Bermuda Islands by present restriction). 

Chelonia pseudo-mydas Lesson, in Belanger, Voyage aux Indes Oriental . . ., 
Zool., 1834, p. 299 (type unknown; type locality, Atlantic Ocean, here re- 
stricted to the Bermuda Islands). 

Chelonia pseudo-caretta Lesson, op. cit., p. 302 (type and type locality as in the 
preceding) . 

' The Atlantic Ocean specimens are frequently regarded as Eretmochelys imbTicata (or E. i. imbricata) , 
the Pacific Ocean ones as Eretmochelys imbricata bissa (or E. i. sguamata). We are unable to determine 
characters for such separation. 



CHECKLIST OF REPTILES OF MEXICO 17 

Caretta bissa RCppell, Neue Wirbelthiere Abyssiniens, Amphibiens, 1835, p. 4, 

pi. 2 (type unknown; type locality, Red Sea). 
Eretmochelys squamata Agassiz, Contributions to the natural history of the 

United States, vol. 1, 1857, p. 382 (cotypes, Mus. Comp. Zool. Nos. 1415, 

1416; lectoholotype, No. 1416; type locality restricted to Singapore, Straits 

Settlements) . 

Type. — Unknown. 

Type locality. — American seas, here restricted to the Bermuda 
Islands. 

Range. — Tropical oceans, world-wide; both coasts of Mexico. 
Reported from Baja California, Sonora, Oaxaca, Campeche, and 
Quintana Roo (Isla de Mujeres). 

Genus CHELONIA Brongniart 

Chelonia Brongniart, Bull. Sci. Soc. Philom., vol. 2, 1800, p. 89. 

Chelone Brongniart, Mem. Sav. fitrang, vol. 1, 1806, p. 610. 

Chelona Burmeister, Handbuch der Naturgeschichte, vol. 2, 1807, p. 731. 

Chelonias Rafinesque, Specchio Sci. Palermo, vol. 2, 1814, p. 66. 

Mydas Cocteau, in Ram6n de la Sagra, Historia Ksica, polltica y natural de la 

Isla de Cuba, vol. 4, 1838, p. 22. 
Mydasea Gervais, Diet. Hist. Nat., vol. 3, 1843, p. 457. 

Euchelonia Tschudi, Untersuchungen iiber die Fauna Peruana . . ., 1846, p. 22. 
Megemys Gistel, Naturgeschichte des Thierreichs, 1848, p. 8 (type of all above, 

Testudo mydas Linnaeus), 

Genotype. — Testudo mydas Linnaeus. 

Species. — Possibly four forms (species or subspecies) are recogniz- 
able. None of these are well enough known to us to diagnose properly. 
Range. — Tropical oceans, world-wide. 

CHELONIA MYDAS' (Linnaeus) 

Testudo mydas Linnaeus, Systema naturae, ed. 10, 1758, p. 197. 

Chelonia mydas, Brongniart, Bull. Sci. Soc. Philom., vol. 2, 1800, p. 89.— Schweig- 

ger, Konigsberg. Arch. Naturw. Math., vol. 1, 1812, p. 412.— Deranita- 

GALA, Tetrapod reptiles of Ceylon, vol. 1, 1939, pp. 218-242, figs. 86-95. 
Testudo viridis Schneider, Algemeine Naturgeschichte der Schildkroten . . ., 

1782, pp. 299-303 (type unknown; type locality, Charleston, S. C, by present 

restriction) . 
Chelonia virgata Schweigger, Prodromi monographiae cheloniorura . . ., 1814, 

p. 21 (type unknown; type locality, "Seas of Torrid Zone," here restricted to 

the Bermuda Islands). 
Chelonia maculosa Cuvier, Le r^gne animal . . ., ed. 2, vol. 2, 1829, p. 13 (type 

in Mus. Hist. Nat. Paris?; type locahty not designated, here restricted to 

Ascension Island). 
Chelonia agassizii Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, p. 122 (type in 

Mus. Hist. Nat. Paris; type locality, mouth of Rfo Nagualate, Guatemala); 

Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, 

pp. 26-28, pi. 6 (color). 



• A common arrangement restricts mydas to the Atlantic Ocean, agassizii to the Pacific, either as species 
or subspecies. We are unable to determine characters Justifying such an arrangement. 



18 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type. — UdIoiowii. 

Type locality. — Ascension Island. 

Range. — Tropical oceans, world-wide, both coasts of Mexico. 
Reported from Veracruz, Yucatan, Quintana Roo (Isla de Mujeres), 
Oaxaca, Guerrero, Sonora (Tibur6n Island), and Clarion Island. 

Suborder Trionychoidea Fitzinger 

Trionychoidea Fitzinger, Neue Classification der Reptilien . . ., 1826, p. 7. 

Families. — Two, one of which (Carettochelyidae) is restricted to 
New Guinea. 

Family TRIONYCHIDAE Gray 

Trionicidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 212. 

Genera. — Seven, only one of which occurs in the Americas. 
Range. — Asia, Africa, North America. 

Genus AMYDA Schweigger 

.i4m?/da8cHWEiGGER, in Geoflfroy, Ann. Mus. Hist. Nat. Paris, vol. 14, 1809, p. 1. — 
CoNANT and Goin, Occ. Pap. Mus. Zool. Univ. Michigan, No. 510, 1948, pp. 
11-19 (taxonomy). 

Genotype. — Amy da javanica Geoff roy [=Amyda cartilaginea (Bod- 
daert)]. 

Species. — About 21, of which 4 are American; two or three sub- 
species are recognized of one species (A. spinifera). One occurs in 
Mexico. ^° 

Range. — Asia, North America. 

AMYDA EMORYI (Agassiz) 

Aspidonecles emoryi Agassiz, Contributions to the natural history of the United 
States, vol. 1, 1857, p. 407; vol. 2, 1857, pi. 6, figs. 4, 5. 

Amyda emoryi, Stejnegbr and Barbour, Check list of North American amphib- 
ians and reptiles, 1917, p. 124. 

Type.—Mvi^. Comp. Zool. Nos. 1901-1910, cotypes; W. H.Emory 
collector. 

Type locality. — Rio Grande River, near Brownsville, Tex. 

Range. — Southern Oldahoma and Arizona southward into northern 
Mexico. Recorded from Coahuila: Hacienda Los Borregos (near 
Judrez), San Juan, Cuatro Cienegas, Hacienda La Gacha; Nuevo 
Leon: Rodriguez; Tamaulipas: Matamoros, Rio Purificaci(5n north of 
Ciudad Victoria; Baja California?. 

Suborder Cryptodira Cope 

Cryptodira Cope, Proc. Amer. Assoc. Adv. Sci., vol. 19, 1870, p. 235. 

Families. — Six, all but one of which (Platysternidae, southern Asia) 
occur in Mexico. 



•« Muller (Verb. Naturf. Ges. Basel, vol. 6, 1878, p. G41) records A. mutica also from "Mexico." The 
record requires conflrmation. 



CHECKLIST OF REPTILES OF MEXICO 19 

KEY TO MEXICAN FAMILIES OF CRYPTODIRA 

1. Abdominal scutes in contact with marginals or separated from them only by 

membrane; inguinal scute short, less than half length of bridge; 12 plastral 

scutes 4 

All plastral scutes separated from marginals by a series of inframarginals, or 
inguinal scute very long, half length of bridge or longer; 12 or fewer plastral 
scutes 2 

2. Four or 5 inframarginals Dermatemyidae (p. 19) 

Two or 3 (rarely 1) inframarginals 3 

3. Bridge very narrow, covered chiefly by the displaced abdominal scutes, which 

are widely separated from each other medially Chelydridae (p. 20) 

Bridge broad, abdominal scutes in normal position Kinostemidae (p. 21) 

4. Digits with no webs whatever; rear foot stump-shaped, plantar surface as 

broad as long Testudinidae (p. 27) 

Digits with at least some vestige of webs; rear foot more elongate, plantar 
surface longer than broad Emyidae (p. 28) 

Family DERMATEMYIDAE Gray 

Dermatemyidae Gray, Supplement to the catalogue of the shield reptiles . . ., 
1870, p. 49. 

Genera. — One. 

Range. — Central Mexico to Honduras. 

Genus DERMATEMYS Gray 

Dermatemys Gray, Proc. Zool. Soc. London, 1847, p. 55. 

Chloremys Gray, Proc. Zool. Soc. London, 1870, p. 715 (type, Dermatemys abnormis 

Cope=Z). mawii Gray). 
Limnochelone Werner, Zool. Anz., vol. 24, 1901, p. 297 (type, Limnochelone 

micrura Werner). 

Genotype. — Dermatemys mawii Gray. 

Range. — Atlantic coast from central Veracruz to Honduras. 

Species. — One recognized. 

dermatemys MAWn Gray 

Dermatemys mawii Gray, Proc. Zool. Soc. London, 1847, p. 55. — Boulenger, 

Catalogue of the chelonians . . ., 1889, pp. 28-29. 
Emys Berardii Dum^ril and Bibron, in Dum^ril and Dum^ril, Catalogue 

m^thodique de la collection des reptiles, livr. 1, 1851, p. 11 (type locality, 

environs of Veracruz, Mexico; type in Mus. Hist. Nat. Paris). 
Dermatemys abnormis Cope, Proc. Acad. Nat. Sci. Philadelphia, vol. 20, 1868, 

p. 120 (type, U.S.N.M. No. 6545; Belize River, British Honduras). 
Limnochelone micrura Werner, Zool. Anz., vol. 24, 1901, p. 298 (type locality, 

Mexico, here restricted to Alvarado, Veracruz; type unknown). 

Type. — Brit. Mus. 

Type locality. — Uriknown, here restricted to Alvarado, Veracruz. 

Range. — Rivers of the Atlantic coast from central Veracruz to 
Guatemala, probably excludmg the northern part of the Yucat§,n 
Peninsula. Recorded from the states of Veracruz, Oaxaca, Tabasco, 
Yucatan, and Campeche. 



20 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Family CHELYDRIDAE Swainson 

Chelidridae Swainson, Natural history and classification of fishes, amphibians, 
and reptiles, vol. 2, 1839, p. 116. 

Genera. — Two, one of which (Macrochelys) is restricted to the 
United States. A supposed New Guinea genus, Devisia, is an errone- 
ously labeled Chelydra serpentina. 

Range. — North America south to northern South America. 

Genus CHELYDRA Schweigger 

Chelydra Schweigger, Konigsberg. Arch. Naturw. Math., vol. 1, 1812, p. 292. 
Chelonura Fleming, Philosophy of zoology , . ., vol. 2, 1870, p. 64 (type, Testudo 

serpentina Linnaeus) . 
Rapara Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 211 (type as above). 
Saurochelys Latreille, Families naturelles du rfegne animal . . ., 1825, p. 92 

(type as above). 
Cheliurus Rafinesque, Atlantic Journ., 1832, p. 64 (type as above). 
Emysaurus Dum^kil and Bibron, Erp^tologie g^n^rale, vol. 2, 1835, p. 548 (type 

as above). 
Chelonura Holbrook, North American herpetology, vol. 1, 1836, p. 139 (type 

as above). 

Genotype. — Testudo serpentina Linnaeus. 

Range. — Southeastern Canada, United States east of the Rocky 
Mountains, south to Ecuador. 

Species. — Three species or subspecies ; one is definitely recorded from 
Mexico, another probably occurs there, and the third (acutirostris) is 
restricted to Panama and South America. 

KEY TO MEXICAN SPECIES OF CHELYDRA 

1. Bridge about one-seventh length of plastron; 4 chin barbels; second vertebral 
shield 32 to 34 percent length of carapace; height of skull at quadrate 44 to 
48 percent length of skull to condyle; width of palatine bone 28 to 30 percent 

length of skull rossignonii (p. 20) 

Bridge about one-ninth length of plastron; 2 chin barbels; second vertebral 
shield less than 30 percent length of carapace; height of skull at quadrate 
38 to 43 percent length of skull to condyle; width of palatine bone 19 to 
25 percent length of skull .serpentina (p. 21) 

CHELYDRA ROSSIGNONII (Bocourt) 

Emysaurus rossignonii Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, pp. 121-122; 

Mission scientifique au Mexique . . ., fitudes sur les reptiles, livr. 1, 1870, 

pp. 18-19, pi. 5, fig. 2. 
Chelydra rossignonii, Boulenger, Ann. Mag. Nat. Hist., ser. 7, vol. 9, 1902, pp. 

49-51. — Schmidt, Smithsonian Misc. Coll., vol. 106, No. 8, 1946, pp. 1-9. 

Type. — Mus. Hist. Nat. Paris. 

Type locality. — Panzos, Rio Polochic, Guatemala. 

Range. — Atlantic slopes of Guatemala and presumably adjacent 
Mexico southward to Costa Rica. Not yet reported definitely from 
Mexico. 



CHECKLIST OF REPTILES OF MEXICO 21 

CHELYDRA SERPENTINA (Linnaeus) 

Testudo serpentina Linnaeus, Systema naturae, ed. 10, vol. 1, 1758, p. 199. 
Chelydra serpentina, Schweigger, Konigsberg. Arch. Naturw. Math., vol. 1, 
1812, p. 293.— Cahn, Illinois Biol. Monogr., vol. 16, 1937, pp. 34-45, pis. 3-4. 

Type . — Unknown . 

Type locality. — "Warmer region," here restricted to New Orleans, 
La. 

Range. — North America east of the Rocky Mountains; in Mexico, 
Atlantic slopes south to the Yucatan Peninsula; recorded only from 
the states of Veracruz and Campeche. 

Family KINOSTERNIDAE Agassiz 

Cinosternidae Agassiz, Contributions to the natural history of the United States, 
vol. 1, 1857, p. 347. 

Genera. — Three. 

Range. — North America, south to northern South America. 

KEY TO GENERA OF KINOSTERNIDAE 

1. Plastron very small, cruciform, with 7-9 scutes; bridge narrow, its length 

(parallel to body axis) exceeded by its width; inframarginals little longer 

than broad 2 

Plastron larger, not cruciform, with 10 or 11 scutes; bridge broader, its length 
much exceeding its width; inframarginals about three times as long as broad, 
or longer Kinostemon (p. 21) 

2. A ligament between inframarginals and plastron Claudius (p. 26) 

A suture between inframarginals and plastron Staurotj^pus (p. 27) 

Genus KINOSTERNON Spix 

Kinosternon Spix, Ranae et testudinis brasiliensis species novae, 1825, p. 17. 
Swanka Gray, Catalogue of the tortoises . . ., 1844, p. 32 (type, Kinosternon 

scorpioides Gray). 
Platythyra Agassiz, Contributions to the natural history of the United States, 

vol. 1, 1857, p. 430, pi. 5, figs. 12-15 (type, Platythyra flavescens Agassiz). 
Thyrosternum Kqkbbiz, op. cit., p. 429 (type, Kinosternum integrum LeConte). 

Genotype. — Kinosternon longicaudatum Spix=i^. scorpioides Lin- 
naeus. 

Range. — United States, except the northwestern quarter, south to 
northern South America. 

Species. — About 22 species and subspecies, 12 of which are known to 
occur in Mexico. 

KEY TO MEXICAN SPECIES OF KINOSTERNON 

1. Plastron rounded behind, not incised or indented; stridulation organs 
(on concealed surfaces of shank and thigh) in males poorly developed 
or absent; plastron large, completely closing shell in adults 2 



22 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Plastron incised or indented behind; stridulation organs in males well 
developed or not; plastron at least sonaewhat smaller, not completely 
closing shell even in adults 7 

2. Carapace tricarinate; axillary and inguinal scutes (if present) widely 

separated 3 

Carapace flat or unicarinate; axillary and inguinal scutes narrowly in 
contact or narrowly separated 5 

3. Axillary scute usually absent; posterior edge of abdominals convex. 

abaxillare (p. 22) 
Axillary scute present; posterior edge of abdominals straight 4 

4. Anterior lobe of plastron longer than immovable portion of plastron. 

cruentatum cruentatum (p. 23) 
Anterior lobe of plastron as long as or shorter than immovable portion of 
plastron cruentatum censors (p. 24) 

5. Anterior lobe of plastron longer than fixed portion 6 

Anterior lobe of plastron not longer than fixed portion acutum (p. 23) 

6. Gular more than half length of anterior lobe of plastron. .creaseri (p. 23) 
Gular less than haK length of anterior lobe of plastron, .leucostomum (p. 25) 

7. Carapace tricarinate; stridulation organs absent; plastron slightly smaller 

than opening of carapace integrum (p. 25) 

Carapace flat or unicarinate; stridulation organs well developed in males; 
plastron distinctly smaller than opening of carapace 8 

8. Ninth marginal about as high as tenth, much higher than eighth 9 

Ninth marginal much lower than tenth, about height of eighth 10 

9. Gular more than half length of anterior lobe of plastron (60-64 percent); 

length of anterior lobe less than combined length of gular plus length 
of interfemoral suture plus width of nuchal (83-93 percent). 

flavescens stejnegeri (p. 24) 
Gular half or less than half length of anterior lobe of plastron (30-50 percent) ; 
length of anterior lobe greater than combined length of gular plus length 
of interfemoral suture plus width of nuchal (128-196 percent). 

fiavescens flavescens (p. 24) 

10. Anterior vertebral very narrow, widely separated from second marginal 

on each side; gular less than half length of anterior lobe; posterior marginal 

(eleventh) little lower than penultimate (tenth) herrerai (p. 24) 

Anterior vertebral broad, in contact with second marginal on each side; 
gular variable in length; posterior marginal distinctly lower than penulti- 
mate 11 

11. Carapace depressed; entire shell twice as broad as deep; head light, dark- 

spotted; sutures between scutes of carapace not distinctly black-bordered, 
those between plastral scutes not or narrowly marked with brown. 

sonoriense (p. 26) 
Carapace peaked, arched as viewed directly from front; shell less than twice 
as broad as deep; head dark, light-spotted, sutures on carapace narrowly 
but distinctly black-edged, those on plastron broadly marked with brown. 

hirtipes (p. 25) 

KINOSTERNON ABAXILLARE Baur 

Kinosternon ahaxillare Baur, in Stejneger, Journ. Washington Acad. Sci., vol. 15, 
1925, pp. 462-463. 

Tyjpe.—U.S.^M. No. 7518; C. H. Berendt collector. 

Type locality. — Tuxtla Gutierrez, Chiapas, Mexico. 

Range. — Plateau of Chiapas. Recorded only from the type locality. 



CHECKLIST OF REPTILES OF MEiXICO 23 

KINOSTERNON ACUTUM Gray 

Kinosternon scorpioides acuta Grat, Synopsis reptilium, 1831, p. 34, pi, 7, fig, 1. 
Kinosternon acutum, Stejneger, Proc. U. S. Nat. Mus., vol. 90, 1941, pp. 457-458. 
Cinosternum berendtianum Cope, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 189 

(Tabasco; U.S.N.M. No. 6517; C. H. Berendt collector). 
Swanka maculata Grat, Proc. Zool. Soc. London, 1869, p. 162 (type locality, 

Cosamaloapam, Veracruz, Mexico; cotype in Brit. Mus.), 
Cinosternon Effeldtii Peters, Monatsb. Berlin Akad. Wiss., 1873, p. 603, pi, 5, 

figs. 1-3 (type locality, "presumably" Veracruz, Mexico, here restricted to 

Cosamaloapam; type Berl. Mus.). 

Type. — Brit. Mus. 

Type locality. — Unknown, here restricted to Cosamaloapam, Vera- 
cruz. 

Range. — Atlantic coast from central Veracruz southward to British 
Honduras, excluding the northern part of the Yucatan Peninsula. 
Recorded from the states of Veracruz, Tabasco, and Campeche." 

KINOSTERNON CREASERI Hartweg 

Kinosternon creaseri Hartweg, Occ. Pap. Mus. Zool. Univ. Michigan, No. 277, 
1934, pp. 1-2. 

Type. — Univ, Mich. Mus, Zool, No. 73090, male; Edwin P. Creaser 
collector. 

Type locality. — One mile south of the Hacienda, Chichen Itzd, 
Yucatdn. 

Range. — Known only from the general region of the type locality, 
and Quintans Roo (Vivienda de Platanal). 

KINOSTERNON CRUENTATUM CRCENTATUM Dmn^ril and Bibron 

C[inosternon] cruentatum Dumeril and Bibeon, in Dum^ril and Dum^ril, Cata- 
logue m6thodique de la collection des reptiles, livr. 1, 1851, pp. 16-17. — 
GtJNTHER, Biologia Centrali- Americana, Reptilia and Batrachia, 1885, pp, 
13-14, pis. 9-11. — Siebenrock, Zool. Anz., vol. 30, 1906, pp. 98-99. 

Kinosternon cruentatum cruentatum, Wettstein, Sitzb. Akad. Wiss. Wien, math.- 
nat. Kl., Abth. 1, vol. 143, 1934, pp. 14r-l 5.— Schmidt, Publ. Field Mus. Nat. 
Hist., zool. ser., vol. 22, 1941, p. 488. 

K[inosternum] mexicanum LeConte, Proc. Acad. Nat. Sci. Philadelphia, 1854, 
pp. 182-183 (type locality, Mexico, here restricted to San Mateo del Mar, 
Oaxaca; type presumably in Acad. Nat. Sci. Philadelphia). 

Cinosternum triliratum LeConte, Proc. Acad. Nat. Sci. Philadelphia, 1859, p. 6 
(type locality, "Mexico," here restricted to San Mateo del Mar, Oaxaca; 
Pease coll.; Acad, Nat. Sci. Philadelphia?). 

Cinosternon Shawianum Bocotjrt {nee Bell), Journ. Zool., vol. 5, 1876, pp. 387, 
397-398. 

Type. — Mus. Hist. Nat. Paris. 

Type locality. — "Amer. septentr.," here restricted to San Alateo del 
Mar, Oaxaca. 

" Qadow's record (Proc, Zool. Soc. London, 1905, p. 194) for San Mateo, Oaxaca, is considered erroneous. 



24 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Range. — Atlantic and Pacific drainages in Oaxaca, Tabasco, Campe- 
che, and Chiapas. ^^ In Central America, Guatemala. 

KINOSTERNON CRUENTATUM CONSORS Stejneger 

Kinosternon cruentatum consors Stejneger, Proc. U. S. Nat. Mus., vol. 90, 1941, 
pp. 45S-459. 

Type— \J. S.N. M. No. 13912. 
Type locality. — Cozumel Island, Quintana Roo. 
Range. — The type locality and northern Yucat§,n (Progreso and 
Telchac Puerto). 

KINOSTERNON FLAVESCENS FLAVESCENS (Agassiz) 

Platythyra flavescens Agassiz, Contributions to the natural history of the United 
States, vol. 1, 1857, p. 430; vol. 2, pi. 5, figs. 12-15.— Cahn, Illinois Biol. 
Monogr., vol. 16, Nos. 1-2, 1937, pp. 56-62, pis. 6, 7. 

Kinosternon flavescens flavescens, Hartweg, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 371, 1938, pp. 2, 3. 

Type . — Unknown . 

Type locality. — Texas and Arizona, here restricted to Waco, Tex. 

Range. — Northern Mexico from Coahuila to Tamaulipas; recorded 
in Coahuila (Jaral). In the United States from Arizona and Texas 
north to Colorado and Kansas, 

KINOSTERNON FLAVESCENS STEJNEGER! Hartweg 

Kinosternon flavescens stejnegeri B.A.B.TWEa, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 371, 1938, pp. 1-5. 

Type. — Univ. Alichigan Mus. Zool. No. 72235; Morrow J. Allen 
collector. 

Type locality. — Llano, Sonora (midway between Nogales and 
Hermosillo) . 

Range. — Known only from Sonora (type locality) and Durango 
(Pedricena) ; perhaps also from southern Coahuila (said to intergrade 
with flai^escens flavescens at Jaral, Coahuila). 

KINOSTERNON HERRERAI Stejneger 

Kinosternon herrerai Stejneger, Journ. Washington Acad. Sci., vol. 15, 1925, 
p. 462. — Shannon and Smith, Trans. Kansas Acad. Sci., vol. 52, No. 4, 
1949, pp. 497-498. 

Type.—U.S.N.M. No. 61249; Alfonso Herrera, donor. 
Type locality. — Xochimilco, Valley of Mexico, Distrito Federal, 
Mexico, in errore; here restricted to La Laja, Veracruz. 
Range. — Known only from the type locality. 

1' A record from La Majada, near ApatzingSn, Michoac&n (Schmidt and Shannon, Fieldiana, vol. 31, 
1947, p. 69), is referable to K. integrum. 



CHECKLIST OF REPTILES OF MEXICO 25 

KINOSTERNON HIRTIPES Wagler 

Cinosternon hirtipes Wagler, Nattirliches System der Amphibien, . . ., 1830, 

pi. 5, figs. 29, 30. 
Cinosternum hirtipes, Gunther, Biologia Centrali-Americana, 1885, p. 15, pi. 

12-15.— SiEBENROCK, Zool. Anz., vol. 30, 1906, pp. 94-97, figs. 
Cinosternum henrici LeConte, Proc. Acad. Nat. Sci, Philadelphia, 1859, p. 4 

(type presumably in Acad. Nat. Sci. Philadelphia; type locality, New Mexico, 

T. C. Henry collector). — Ditmars, Reptile book, 1907, p. 26, pi. 11, fig. 

Type. — Munich Museum. 

Type locality. — Mexico, here restricted to Mazatl^n, Sinaloa, 
Range. — Western Texas and southern Arizona southward through 
the main Mexican Plateau, from Chihuahua to Mexico; recorded 
from the states of Chihuahua, Sinaloa, Michoacan, Colima, Guana- 
juato, San Luis PotosI, Hidalgo, Mexico, and from Distrito Federal 
and the Tres Marias Islands. 

KINOSTERNON INTEGRUM LeConte 

Kinosternum integrum LeConte, Proc. Acad. Nat. Sci. Philadelphia, 1854, p. 183. 
Cinosternum integrum, Boulenger, Catalogue of the chelonians, rhynchocepha- 

lians and crocodiles in the British Museum, 1889, p. 42. 
Cinosternum scorpioides integrum, Siebenrock, Zool. Anz., vol. 30, 1906, pp. 96-97. 
Cinosternum rostellum Bocourt, Journ. Zool., vol. 5, 1876, pp. 391-392 (type 

locality, Guanajuato, Mexico; type in Mus. Hist. Nat. Paris). 

Type. — Acad. Nat. Sci. Philadelphia; Mr. Pease collector. 
Type locality. — Mexico, here restricted to Acapulco, Guerrero. 
Range. — The plateau of Mexico from Sonora to Oaxaca east to 
Veracruz. Known from Tres Marias Islands and the states of Sonora, 
Sinaloa, Nayarit, Colima, Michoacan, Guerrero, Oaxaca, Morelos, 
Guanajuato, Aguascalientes, Jalisco, San Luis Potosi, Puebla, and 
Veracruz. 

KINOSTERNON LEUCOSTOMUM Dam6ril and Bibron 

C[inosternon] leucostomum Dum^ril and Bibron, in Dum^ril and Dum^ril, 

Catalogue m6thodique de la collection des reptiles, livr. 1, 1851, p. 17, figs. 

1-3. — GtJNTHER, Biologia Centrali-Americana, Reptilia and Batrachia, 

1885, p. 17, pis. 16, 17.— Siebenrock, Zool. Anz., vol. 30, 1906, pp. 97-98. 
Swanka lexicostoma, Gray, Catalogue of the shield reptiles in the British Museum, 

pt. 1, Testudinata, 1855, p. 69. 
Swanka maculata Gray, ibid., p. 68 (type locality, "Vera Paz," Guatemala, and 

Cosamaloapam, Veracruz, here restricted to the latter; type in Brit. Mus. 

Nat. Hist.). 
Cinosternum brevigvlare GtJNTHER, Biologia Centrali-Americana, Reptilia and 

Batrachia, 1885, pp. 17-18 (type locality, Playa Vicente, Mexico, Sall6 

collector; type in Brit. Mus. Nat. Hist.). 
Cinosternum cobanum Gxjnther, op. cit., p. 18, pi. 18, fig. B (type in Brit. Mus. 

Nat. Hist.; type locality, Cobdn and Cahabon, Guatemala, here restricted 

to Cobdn). 

Type. — ^Mus. Hist. Nat. Paris. 

Type locality. — "N. Orleans; Mexique; Rio-Sumasinta (Amer. 



26 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

centr.): M. Morelet. Am6\ septentr. cf9 ?Vallee de la Madeleine 
(N. Grenade), ad. et j. age: M. J. Goudot. Santa-Fe de Bogotd 
(N. Grenade): M. Lewy, V. V. Velins No. 38." Here restricted to 
Cosamaloapam, Veracruz. 

Range. — Atlantic slopes of southern Mexico south of central 
Veracruz; recorded from the states of Puebla, Veracruz, Tabasco, 
Campeche, and Chiapas; records from Yucatan and Cozumel Island 
presumably are referable to K. creaseri and K. cruentatum consors. 
Ranges through Central America to northern South America. 

u 

KINOSTERNON SONORIENSE LeConte 

Kinosternum sonoriense LeConte, Proc. Acad. Nat. Sci. Philadelphia, 1854, p. 184. 
Thyrosternum sonoriense, Agassiz, Contributions to the natural history of the 

United States, vol. 1, 1857, p. 428; vol. 2, pi. 5, figs. 8-11 {Cinosiernum 

sonoriense LeConte). 
Kinosternon sonoriense, Stejneger, Proc. U. S, Nat. Mus., vol. 25, 1902, p. 149. — 

Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 967-970, 

pis. 120-121. 
tCinosternon ■punctalum Or at. Catalogue of the shield reptiles in the British 

Museum, 1855, p. 45, pi. 20c, figs. 5, 6 (type in Brit. Mus.; "North America," 

here restricted to Tucson, Ariz.). 

Type. — Acad. Nat. Sci. Philadelphia. 

Type locality. — Tucson, Ariz. 

Range. — Sonora, Chihuahua, and Durango (Durango); in the 
United States: southeastern California, southern Arizona, New 
Mexico to western Texas. 

Genus CLAUDIUS Cope 

Claudius Cope, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 187. 

Genotype. — Claudius angustatus Cope. 

Range. — Restricted to the Atlantic coast from Veracruz to British 
Honduras. 
Species. — One. 

CLAUDIUS ANGUSTATUS Cope 

Claudius angustatus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1865, pp. 187, 

189. — BocouRT, Mission scientifique au Mexique . . ., fitudes sur les 

reptiles, livr. 1, 1870, pp. 20-21, pi, 4 (color). 
Claudius megalocephalus Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, p. 122 

(type locality, Mexico, here restricted to Tabasco; type in Mus. Hist. Nat. 

Paris) . 

Type.— U.S. ]<l.M. Nos. 6518, 6525. 
Type locality. — Tabasco. 

Range. — Veracruz southward to British Honduras. Recorded in 
Mexico from the states of Veracruz, Tabasco, and Campeche. 



CHECKLIST OF REPTILES OF ME^XICO 27 

Genus STAUROTYPUS Wagler 

Staurotypus Wagler, Natiirliches System der Amphibien . . ., 1830, p. 137. 
Stauremys Gray, Proc. Zool. Soc. London, 1864, p. 127 [genotype, Staurotypus 
(Stauremys) salvinii Gray]. 

Genotype. — Terrapene triporcata Wiegmann. 

Range. — Atlantic slopes of southern Mexico south to Guatemala. 

Species. — Two. 

KEY TO SPECIES OF STAUROTYPUS 

1. Suture between humerals longest; abdominals much broader than long; length 

of bridge 5}^ or more times in length of plastron salvinii (p. 27) 

Suture between pectorals longest; abdominals as broad as long; length of 
bridge less than BYi times in length of plastron triporcatus (p. 27) 

STAUKOTYPUS TRIPORCATUS (Wiegmann) 

Terrapene triporcata Wiegmann, Isis von Oken, 1828, p. 364. 

Staurotypus triporcatus, Wagler, Natiirliches System der Amphibien . . ., 1830, 

pi. 5, figs. 44-45. 
Staurotypus salvinii, BocotTRT (nee Gray), Mission scientifique au Mexique . . ., 

fitudes sur les reptiles, livr. 1, 1870, pp. 22-23, pi. 5, fig. 3. 
Claudius pictus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1872, pp. 26-27 (a 

name proposed for the preceding specimen of Bocourt; type locality. Vera Paz, 

Guatemala, here restricted to Alta Verapaz; type presumably in Acad. Nat. 

Sci. Philadelphia). 

Type. — Presumably in the Berlin Mus. ; Deppe collector. 

Type locality. — Rio Alvarado, Veracruz. 

Range. — Atlantic slopes from Veracruz to Guatemala and British 
Honduras (recorded in Mexico only from the states of Veracruz and 
Tabasco) . 

STAUROTYPUS SALVINII Gray 

Staurotypus salvinii Gray, Proc. Zool. Soc. London, 1864, p. 127. 

Staurotypus marmoratus Fischer, Archiv fiir Naturg., 1872, p. 265, pi. 10 (tj^pe 
locality, "Mexico," here restricted to Santa Efigenia, Oaxaca; St. Petersburg 
Mus). 

Claudius severus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1872, pp. 24-26 (type 
locality, Santa Efigenia, Oaxaca; type presumably in Acad. Nat. Sci. Phila- 
delphia) . 

Staurotypus biporcatiis Siebenrock, Zool. Jahrb., Suppl. 10, No. 3, 1909, p. 438 
(type locality restricted here to that of S. salvinii Gray; type here restricted, 
as lectotype, to that of S. salvinii Gray;. 

Type. — Brit. Mus. 

Type locality. — Huamuchil, Guatemala. 

Range. — Pacific slopes from the Isthmus of Tehuantepec to Guate- 
mala (in Mexico recorded only from Oaxaca). 

Family TESTUDINIDAE Gray 

Testudinidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 210. 
861316—50 3 



28 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Genera. — About four, of which one occurs in Mexico. 
Range. — World-wide in tropical and subtropical areas, except the 
East Indies and Australia. 

Genus GOPHERUS Rafinesque 

Gopherus Rafinesque, Atlantic Journ., vol. 1, 1832, p. 64. 
Genotype. — Testudo polyphemus Daudin. 
Range. — Southern United States and northern Mexico. 
Species. — Three recognized, two of which occur in Mexico. 

KEY TO MEXICAN SPECIES OF GOPHERUS " 

1. Carapace dome-shaped, rounded on top; carapace length about twice shell 

height (47-56 percent) berlandieri (p. 28) 

Carapace with flat-topped contour; carapace length more than twice shell 
height (36-48 percent) agassizii (p. 28) 

GOPHERUS BERLANDIERI (Agassiz) 

Xerobates berlandieri Agassiz, Contributions to the natural history of the United 
States, vol. 1, 1857, p. 447; vol. 2, 1857, pi. 3, figs. 17-19. 

Testudo berlandieri, Boulenger, Catalogue of the chelonians, rhynchocephalians 
and crocodiles in the British Museum, 1889, p. 156. 

Gopherus berlandieri, Stejneger, North Amer. Fauna, No. 7, 1893, p. 161. 

Type. — U.S.N.M. No. 60, two specimens. 

Type locality. — Lower Rio Grande, Tex., here restricted to Browns- 
ville. 

Range. — Nuevo Le6n, Tamaulipas, and Coahuila ^*; in United 
States: Southern Texas. 

GOPHERUS AGASSIZII (Cooper) 

Xerobates agassizii Cooper, Proc. California Acad. Sci., vol. 2, 1863, p. 125. 

Testudo agassizii, Boulenger, Catalogue of the chelonians, rhynchocephalians 
and crocodiles in the British Museum, 1889, p. 156. 

Gopherus agassizii, Stejneger, North Amer. Fauna, No. 7, 1893, p. 161. — Wood- 
bury and Hardt, Ecol. Monogr., vol. 18, 1948, pp. 145-200, figs. 1-25. 

Type. — Unknown . 

Type locality, — Mountains near Fort Mojave, Calif. 

Range. — Baja California, Sonora including Tibur6n Island; in the 
United States, southeastern California and southwestern Arizona 
north into Nevada and Utah. 

Family EMYIDAE '' Gray 

Emyidae Gray, Ann. Fhilos., ser. 2, vol. 10, 1825, p. 210. 



" From Woodbury and Hardy, Ecol. Monogr., vol. 18, 1948, p. 152. 

1* Duges's record (La Naturaleza, ser. 2, vol. 1, 1888, pp. 146-147; vol. 2, p. 479) from Bolson de Mapimf, 
Chihuahua, cannot be authoritatively allocated. It may be referable to this species. 

i» See Dermochelyidae (p. 13) for a discussion of orthography of family names. The present family has 
as its type the genus Emys, not the genus Emyda Gray, 1831, which belongs to the family Trionychidae 
(a homonym, Emyda Rafinesque, 1815, long overlooked but of earlier date, belongs to the Emyidae, but 
being a substitute for the generic name Emys, it is not available). 



CHECKLIST OF REPTILES OF MEXICO 29 

Genera. — About 25, of which 6 occur in Mexico. 
Range. — World-wide except Aiistraha and East Indies. 

KEY TO MEXICAN GENERA OF EMYIDAE 

1. Plastron hinged, movable; no inframarginals Terrapene (p. 34) 

Plastron fixed, immovable; inframarginals present 2 

2. Alveolar surface of upper jaw with a ridge either smooth or toothed, parallel to 

labial cutting edge 5 

Alveolar surface of upper jaw ridgeless, smooth 3 

3. A strong interrupted median keel on carapace, with a marked tubercle at the 

end of each vertebral; a low dorsal crest of enlarged compressed scales on 
tail; bridge much shorter than posterior lobe of plastron. 

Malaclemys (p. 36) 

Dorsal keel on carapace absent or continuous, not tuberculate; no crest 

of scales on tail 4 

4. Bridge much shorter than posterior lobe of plastron Clemmys (p. 37) 

Bridge longer than posterior lobe of plastron Geoemyda (p. 29) 

5. Rear of carapace perfectly smoothly rounded, edges of all plates even, no 

notches except a median rear one; a terminal notch flanked on either side by 
a projection on upper jaw, and cutting edge of lower jaw smooth. 

Chrysemys (p. 33) 

Rear of carapace at least somewhat serrate; no terminal notch flanked on 

either side by a projection in upper jaw, or if so cutting edge of lower jaw 

serrate Pseudemys (p. 31 ) 

Genus GEOEMYDA Gray 

Geoemyda Gray, Proc. Zool, Soc. London, 1834, p. 100.— Wettstein, Sitzb. Akad. 

Wiss. Wien, math.-naturw. Kl., Abth, 1, vol. 143, 1934, pp. 17-19 (key to 

American species). 
Rhinoclemys Gray, Ann. Mag. Nat. Hist., ser. 3, vol. 12, 1863, p. 182 (genotype, 

Geoclemmys annulata Gray). 

Genotype. — Testudo spengleri Gmehn. 

Range. — India to Japan and the Malay Archipelago; the coasts 
of central Mexico south into South America. 

Species. — Nineteen species and subspecies are known, nine in the 
Americas, four in Mexico. 

KEY TO MEXICAN SPECIES OF GEOEMYDA 

1. Bridge yellow areolata (p. 30) 

Bridge black, uniform or with a few yellow markings 2 

2. Upper jaw hooked or smooth; plastron dark with a yellow margin.rubida (p. 30) 
Upper jaw notched; plastron yellow, with a broad dark median stripe 3 

3. Carapace almost uniform brown, without distinct markings, its entire border 

strongly bowed, nowhere straight; sides of head in front of eye with usually 

2, sometimes 3, bright horizontal lines pulcherrima incisa (p. 30) 

Carapace olive-brown, with bright, yellow-red concentric or hieroglyphic 
pattern of lines on each scale; young with dark, light-centered spots on 
costals; only lateral border of carapace strongly bowed; sides of head in 
front of eye always with 3 lines pulcherrima pulcherrima (p. 30) 



30 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

GEOEMYDA AREOLATA (Diun«ril and Bibron) 

Emys areolata Dumeril and Bibhon, in Dum^ril and Dum^ril, Catalogue m6tho- 

dique de la collection des reptiles . , ., 1851, p. 10. 
Geoemyda areolata, Wettstein, Sitzb. Akad. Wiss. Wien, math.-nat. KL, Abth. 1, 

vol. 143, 1934, p. 18. 

Type. — Mus. Hist. Nat. Paris; M. Morelet collector. 

Type locality. — El Peten, Guatemala, here restricted to La Libertad. 

Range. — Coast of central Veracruz south to northern Guatemala 
and British Honduras. Recorded in Mexico from Veracruz, Tabasco, 
Campeche, Yucatan, and Quintana Roo (Coba, Cozumel Island). 

GEOEMYDA PULCHERRIMA PULCHERRIMA (Gray) 

Emys pulcherrima Gray, Catalogue of the shield reptiles in the British Museum, 
vol. 1, 1855, p. 25, pi. 25, fig. 1. — Bocourt, Mission scientifique au Mexique, 
fitudes sur les reptiles, livr. 1, 1870, pp. 15-16, pi. 7, figs. 1, la, lb (color). 

Geoemyda pulcherrima pulcherrima, Wettstein, Sitzb. Akad. Wiss. Wien, math.- 
nat. Kl., Abth. 1, vol. 143, 1934, p. 18. 

Type. — Brit. Mus. Nat. Hist. 

Type locality. — Mexico, here restricted to Presidio de Mazatlan, 
Sinaloa. 

Range. — Pacific slopes from southern Sonora to Guerrero and 
probably western Oaxaca. Recorded from the states of Sonora, 
Sinaloa, Nayarit, Colima, and Guerrero. 

GEOEMYDA PULCHERRIMA INCISA (Bocourt) 

Emys incisa Bocourt, Ann. Sci. Nat., vol. 10, 1868, p. 121; Mission scientifique 
au Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, pp. 11-13, pis. 1, 2. 

Geoemyda pulcherrima incisa, Wettstein, Sitzb. Akad. Wiss. Wien, math.-nat. 
Kl., Abth. 1, vol. 143, 1934, p. 18. 

Type. — Mus. Hist. Nat. Paris. 
Type locality. — La Union, El Salvador. 

Range. — Pacific slopes of the Isthmus of Tehuantepec southward 
to at least El Salvador. Recorded from Oaxaca and Chiapas, 

GEOEMYDA RUBIDA (Cope) 

Chelopus rubida Cope, Proc. Amer. Philos. Soc, vol. 11, 1870, p. 148. 
Geoemyda rubida, Siebenrock, Zool. Jahrb., Suppl. 10, Heft 3, 1909, p. 500. 
Rhinoclemmys mexicana Gray, Supplement to the catalogue of shield reptiles 

. . ., 1870, p. 30 (type locality, "San Juan del Rfo"; type in Brit. Mus. Nat. 

Hist.; Rebouch collector). 

Type . — Unknown. 
Type locality. — Juchitan, Oaxaca. 

Range. — The Pacific coast from Colima to the Isthmus of Tehuan- 
tepec (recorded from Colima, Michoacan, Guerrero, and Oaxaca^®). 

" The exact location of San Juan del Rio (type locality of Rhinoclemmys mexicana) is uncertain; it prob- 
ably is one of the two cities of that name in the state of Oaxaca. 



CHECKLIST OF REPTILES OF MEXICO 31 

Genus PSEUDEMYS Gray 

Pseudemys Gray, Proc. Zool. Soc. London, 1855 (1856), p. 197. 

Type. — Testudo concinna LeConte. 

Species. — About 8, and a total of about 22 species and subspecies, 
of which 8 forms occur in Mexico. 

Range. — The United States east of the Rocky Mountains, Baja 
California, both coasts of Mexico from the Rio Grande on the east 
and southern Sonora on the west southward to Argentina. 

KEY TO MEXICAN SPECIES OP PSEUDEMYS 

1. Ridge on alveolar surface of upper jaw distinctly toothed; temporal stripe 

absent or not expanded 2 

Ridge on alveolar surface of upper jaw not toothed; temporal stripe or spot 
much expanded 3 

2. A strong notch at apex of upper jaw, flanked on each side by a toothlike projec- 

tion; teeth on alveolar ridge long, numerous; nostrils terminal; snout not 

pointed, obtuse floridana texana (p. 33) 

No notch at apex of upper jaw, and no subterminal toothlike projections; teeth 
on alveolar ridge smaller, less numerous; nostrils pierced below and posterior 
to tip of snout; snout pointed, acuminate umbra (p. 33) 

3. A light, round temporal spot, completely isolated from orbital markings _ _ 4 
Supratemporal light spot elongate, reaching orbit 5 

4. Temporal light spot completely isolated, not continuous with lines either 

anteriorly or posteriorly; bridge with longitudinal black stripes. 

scripta gaigeae (p. 33) 

Temporal light spot continuous toward rear with longitudinal light stripe; 

bridge black spotted scripta nebulosa (p. 32) 

5. Markings solid, intensely black, replacing or obscuring a linear juvenile pat- 

tern, and including a broad area along middle of plastron, lengthwise along 
middle of bridge, and on posterolateral corners of the marginal scutes above 
and below; these markings evident in bony plates as well as in epidermal 

scutes scripta hiltoni (p. 32) 

Markings not as described 6 

6. Plastral markings always present, either smudged, or, if in lines, the lines 

forming ocelli; supratemporal stripe, if visible, reaching eye. 

scripta elegans (p. 32) 

Plastral markings, if visible, not smudged, but in lines that tend to be arranged 

longitudinally, not ocellate; supratemporal stripe reaching eye or not 7 

7. Dark ocellus of each costal scute nearest latter's posteroventral corner. 

scripta cataspila (p. 32) 
Dark ocelli nearest posterodorsal corners of costals scripta omata (p. 31) 

PSEUDEMYS SCRIPTA OKNATA (Gray) 

Emys omata Gray, Synopsis reptilium, 1831, p. 30. — Bocourt, Mission scientifi- 
que au Mexique . . . , fitudes sur les reptiles, livr. 1, 1870, pi. 3, fig. 1, la. 

Pseudemys scripta ornata, Carr, Herpetologica, vol. 1, 1938, p. 135, figs. 3, 4, 5. 

Emys venusta Gray, Catalogue of the shield reptiles . . . , vol. 1, 1855, p. 24, 
pi. 24a (type locality, Mexico, Honduras and America, here restricted to 
Honduras; types in Brit. Mus.). 

Type. — Brit. Mus., two cotypes. 



32 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type locality. — Mazatlan, Sinaloa. 

Range. — Pacific slopes of Mexico southward from southern Sinaloa, 
and both Atlantic and Pacific slopes (presumably) of Central America 
to Panama and possibly northern South America; Atlantic slopes of 
Mexico east of the Isthmus of Tehuantepec (recorded in Mexico from 
the states of Sinaloa, Jalisco, Oaxaca, Cliiapas, Tabasco, Campeche, 
and Yucatan, and from Cozumel Island). 

PSEUDEMYS SCRIPTA NEBULOSA (Van Denburgh) 

Chrysemys nebulosa Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 5, 

1895, p. 84, pis. 4-6. 
Pseiidemys scripta nebulosa, Carr, Amer. Mus. Nov., No. 1181, 1942, p. 1. 

Type . — D estroy ed . 

Type locality. — Los Dolores, Baja California, Mexico. 

Range. — The southern half of Baja California. 

PSEUDEMYS SCRIPTA HILTONI Carr 

Pseudemys scripta hiltoni Carr, Amer. Mus. Nov., No. 1181, 1942, pp. 1-4, fig. 1-3. 

Type. — Amer. Mus. Nat. Hist. No. 63747; John W. Hilton collector. 

Type locality. — Guirocoba, 28 miles southeast of Alamos, Sonora, 
Mexico. 

Range. — Known only from the type locality; presumably occurs in 
northern Sinaloa as well as in Sonora. 

PSEUDEMYS SCRIPTA CATASPILA (Gunther) 

Emys cataspila Gunther, Biologia Centrali-Americana, Reptilia and Batrachia, 

1885, p. 4, pi. 6, fig. B. 
Pseudemys scripta cataspila, Carr, Herpetologica, vol. 1, 1938, p. 135. — Hartweg, 

Copeia, 1939, No. 1, p. 55. 
? Emys callirostris Grat, Catalogue of the shield reptiles in the British Museum, 

vol. 1, 1855, p. 25, pi. 12B (type locality uncertain, here restricted to Alva- 

rado, Veracruz; type in Brit. Mus.). 

Type. — Brit. Mus., seven cotypes. 

Type locality. — Alexico, here restricted to Alvarado, Veracruz. 

Range. — Atlantic slopes of Mexico from the northern border to the 
Isthmus of Tehuantepec (recorded from the states of Tamaulipas 
and Veracruz). 

PSEUDEMYS SCRIPTA ELEGANS (Wied) 

Emys elegans Wied, Reise durch Nord-America, vol. 1, pt. 4, 1838, p. 213. 

Pseudemys elegans. Cope, U. S. Nat. Mus. Bull. 1, 1875, p. 53. 

Chrysemys scripta elegans, Sibbenrock, Zool. Jahrb., Suppl., vol. 10, 1909, pp. 

464-465. 
Pseudemys troostii elegans, Stejneger and Barbour, Check list North American 

amphibians and reptiles, ed. 4, 1939, p. 167. — Schmidt and Owens, Publ. 

Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, p. 101. 
Pseudemys scripta elegans, Cagle, Copeia, 1944, No. 2, p. 105. 



CHECKLIST OF REPTILES OF MEXICO 33 

Type. — Unknown. 

Type locality. — Fox River at New Harmony, Ind. 
Bange. — Northeastern Mexico: Coaliuila, Nuevo Le6n, and Tama- 
ulipas. Widely distributed in central and southern United States. 

PSEUDEMYS SCRIPTA GAIGEAE " Hartweg 

Pseudemys scripta gaigeae Hartweg, Occ. Pap. Mus, Zool. Univ. Michigan, 
No. 397, 1939, pp. 1-4. 

Type.— Vniv. Michigan Mus. Zool. No. G6472; Helen T. Gaige 
collector. 

Type locality. — Rio Grande River, Boquillas, Brewster County, 
Tex. 

Range. — Northern central Mexico (recorded from Coahuila and 
Durango). 

PSEUDEMYS UMBRA (Bocourt) 

Emys grayi Bocourt, Ann. Sci. Nat., ser. 5, vol. 10, 1868, p. 121; Mission scien- 
tifique au Mexique . . ., Etudes sur les reptiles, livr. 1, 1870, pp. 13-15, 
pi. 3, fig. 2, 2a. 

Emys umbra Bocourt, in O'Shaughnessy, Zool. Rec, vol. 13, 1876 (1878), Rapt., 
p. 6 (substitute name for Emys grayi Bocourt, preoccupied by Emys grayi 
Giinther '=Clemmys caspica caspica [Gmelin]). — Gunther, Biologia Cen- 
trali-Americana, Reptilia and Batrachia, 1885, pp. x, 5, pis. 4, 5, 6 (fig. A). 

Type. — Mus. Hist. Nat. Paris. 

Type locality. — Rio Nagualate, Guatemala. 

Bange. — Pacific slopes of the Isthmus of Tehuantepec, southeast- 
ward to Guatemala. Recorded definitely only from the state of 
Oaxaca; other records from Chiapas may refer either to this or to 
P. s. ornata. 

PSEUDEMYS FLORIDANA TEXANA Baur 

Pseudemys texana Baur, Proc. Amer. Philos. Soc, vol. 31, 1893, p. 223. 
Pseudemys floridana texana, Carr, Copeia, 1938, p. 108. 

Tyj^e.— Acad. Nat. Sci. Philadelphia No. 246. 
Type locality. — San Antonio, Tex. 

Range. — Central Texas and northeastern Mexico. Recorded only 
from Nuevo Leon. 

Genus CHRYSEMYS Gray 

Chrysemys Gray, Catalogue of the tortoises . . . , 1844, p. 27. 

Genotype. — Testudo picta Schneider. 

Range. — Eastern three-fourths of the United States, extreme north- 
ern Mexico. 

Species. — One, with four subspecies. One race occurs in Mexico. 

" Schmidt and Owens (Publ. Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, p. 101) regard this as a species 
distinct from that to which elegans belongs, but the distinguishing criterion mentioned appears to be one 
that in reality separates males from females of either form. We have not examined the specimens but re- 
gard all Pseudemys reported by Schmidt and Owens as P. s. elegans. 



34 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

CHRYSEMYS PICTA BELLII (Gray) 

Emys hellii Gray, Synopsis reptilium . . . , 1831, p. 31. 

Chrysemys picta hellii, Bishop and Schmidt, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 18, 1931, p. 136. 

Type. — Lost; originally in Brit. Mus. 

Type locality. — Unknown, here restricted to Manhattan, Kans. 

Range. — Western Illinois to Washington and British Columbia, 
south on the east of the Cascade Mountains to northern Chihuahua 
and southern Texas. Recorded only from the state of Chihuahua: 
Rio Santa Maria, near Progreso. 

Genus TERRAPENE Merrem 

Terrapene Merrem, Tentamen systematis amphibiorum, 1820, p. 27. — Ditmars, 
Zoologica, vol. 17, 1934, pp. 1-44, figs. 1-41. 

Type. — Testudo clausa Gm.e\m== Terrapene Carolina (Linnaeus). 

Range. — North America east of the Rocky Mountains and Arizona, 
south to Yucatdn and Nayarit. 

Species. — Eleven species and subspecies are recognized at present, 
six in Mexico. 

KEY TO MEXICAN SPECIES OF TERRAPENE 

1. Plastron with a conspicuous and well-defined pattern of yellow lines on a dark 

background (or dark lines on a yellow background) ornata (p. 36) 

Plastron without a lined pattern, either uniform or suffused here and there 
with dark color 2 

2. Pattern of carapace consisting of small, round or ovoid yellow dots 1.5-2 mm. 

in diameter, separated from each other by an average distance of 4 mm. 

klauberi (p. 35) 
Pattern not so 3 

3. Width of carapace 65-68 percent length of carapace, average 66 percent; height 

of shell 40-45 percent length of carapace; 4 rear claws; carapace horn-colored, 

without dark borders at sutures coahuila (p. 36) 

Width of carapace 68-79 percent length of carapace, average about 73 percent; 
height of shell 45-58 percent length of carapace, average 51 percent; rear 
claws 3 or 4; carapace either marked with yellow or horn-colored, but sutures 
always broadly margined with dark color 4 

4. No notch at apex of upper jaws; 4 rear claws nelsoni (p. 35) 

Apex of upper jaw notched; rear claws 3 or 4 5 

5. Rear claws always (so far as known) 4 mexicana yucatana (p. 35) 

Rear claws generally 3 (81 percent of '^2 specimens). 

mexicana mexicana (p. 34) 

TERRAPENE MEXICANA MEXICANA (Gray) 

Cistudo (Onychotria) mexicana Gray, Proc. Zool. Soc. London, 1848 (Feb. 1849), 

vol. 16, pp. 16-17, pi. 2. 
Cistudo mexicana, Gray, Catalogue of shield reptiles in the British Museum, pt. 

1, Testudinata, 1855, p. 40. 
Onychotria mexicana, Ducfes, La Naturaleza, ser. 2, vol. 1, 1888, pp. 107-108. 
Cistudo Carolina var. mexicana, Boulenger, Catalogue of the chelonians, rhyn- 

chocephalians and crocodiles in the British Museum, 1889, p. 118. 



CHECKLIST OF REPTILES OF MEXICO 35 

Terrapene mexicana, Baur, Amer. Nat., vol. 27, 1893, p. 677. — Ditmabs, 

Zoologica, vol. 17, 1934, pp. 18-19, fig. 28. — Muller, Zool. Anz., vol. 

113, 1936, pp. 97-114, figs. 1-4. 
Terrapene mexicana mexicana, Smith, Publ. Field IMus. Nat. Hist., zool. ser., 

vol. 24, 1939, pp. 17-18. 
Terrapene goldmani Stejneger, ^^ Proc. Biol. Soc. Washington, vol. 46, 1933, 

pp. 119-120 (type, U.S.N.M. No. 46251; Chijol or Chijoles, southeastern 

corner of the state of San Luis Potosf, Mexico; E. W. NeLson and E. A. 

Goldman collectors). — Ditmars, Zoologica, vol. 17, 1934, pp. 20-22, figs. 

34-36. 
Terrapene yucatana, Ditmars (nee Boulenger), Zoologica, vol. 17, 1934, figs. 29, 

30. 

Type. — Brit. Mus. Nat. Hist.; two female cotypes; collector un- 
known. 

Type locality. — Mexico, restricted by Muller to Tampico, Tamauli- 
pas. 

Range. — Atlantic slopes from central Tamaulipas and eastern 
Nuevo Leon to northern Veracruz. Known only from San Luis 
Potosi, Veracruz, and Tamaulipas. 

TERKAPENE MEXICANA YUCATANA (Boulenger) 

Cistudo yucatana Boulenger, Ann. Mag. Nat. Hist., ser. 6, vol. 15, 1895, p. 330. 
Terrapene yucatana, Siebenrock, Zool. Jahrb., Suppl., vol. 10, 1909, p. 492. 
Terrapene mexicana yucatana. Smith, Publ, Field Mus. Nat. Hist., zool. ser., vol. 
24, 1939, pp. 17-18. 

Type. — Brit. Mus. Nat. Hist. 

Type locality. — Northern Yucatan, here restricted to Chichen Itza. 
Range. — Northern part of the peninsula of Yucatan. Recorded 
only from Yucatan and Quintana Roo. 

TERRAPENE NELSONI Stejneger 

Terrapene nelsoni Stejneger, Journ. Washington Acad. Sci., vol. 15, 1925, p. 
463.— Ditmars, Zoologica, vol. 17, 1934, pp. 19-20, figs. 31-33. 

TVpe.— U.S.N.M. No. 46252; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — Pedro Pablo, Tepic [=Nayarit], Mexico, 2,500 feet 
altitude. 

Range. — Known only from the type locality. 

TERRAPENE KLAUBERl Bogert 

Terrapene klauheri Bogert, Amer. Mus. Nov., No. 1226, 1943, pp. 1-4, figs. 1-13. 

19 Schmidt and Owens (Publ. Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, p. 103) regard T. goldmani 
as a distinct species "of lowland San Luis Potosi," implying that Its closest relative, T. mexicana, is restricted 
to other areas, presumably highland Mexico inasmuch as lowland San Luis Potosi is scarcely distinguish- 
able faunistically from northern Veracruz, southern Tamaulipas, and eastern Nuevo Leon. The entire 
area, as a matter of fact, is small. However, in spite of a few records of occurrence of T. mexicana on the 
plateau, notably by Duges {loc. cit.), we do not regard any of them as reliable, and believe the species is 
restricted to the approximate range here stated. There can, therefore, be no geographical isolation of two 
populations to consider. 



36 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type.— Amer. Mus. Nat. Hist. No. 63751; John W. Hilton col- 
lector. 

Type locality. — Rancho Guirocoba, about 18 miles southeast of 
Alamos, Sonora. 

Range. — Known only from the type locality and Sierra de Batuc, 
8 kilometers northeast of Alatape, Sonora. 

TERRAPENE COAHUILA Schmidt and Owens 

Terrapene coahuila Schmidt and Owens, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 29, No. 6, 1944, pp. 101-103. 

Type. — Chicago Nat. Hist. Mus. No. 41234; Ernest G. Marsh 
collector. 

Type locality. — Quatro Cienegas, Coahuila, Mexico. 
Range. — Known only from the type locality. 

TERRAPENE ORNATA (Agassiz) 

Cistudo ornaia Agassiz, Contributions to the natural history of the United States, 

vol. 1, 1857, p. 445. 
Terrapene ornata, Baur, Science, vol. 17, 1891, p. 191.— Ditmars, Zoologica, vol. 

17, 1934, pp. 16-18, figs. 24-27. 

Cotypes.—U.^.'N.M. Nos. 7541, 7542, 7547(?); Mus. Comp. Zool. 
No. 1536, Burlington, Iowa; J. Ranch collector. 

Type locality. — The upper Missouri and Iowa, here restricted to 
Council Bluffs, Iowa. 

Range. — Widespread in the United States between the Rocky 
Mountains and the Mississippi River reaching east to Indiana and 
west to Arizona; extreme northern Mexico. Reported only from 
Sonora and Chihuahua. 

Genus MALACLEMYS Gray 

Malaclemys Grat, Catalogue of the tortoises . . ., 1844, p. 28. 

Genotype. — Testudo concentrica Shaw. 

Range. — Atlantic and Gulf coasts of the United States and Mexico. 
Species. — One, of five distinct races, only one of which has been 
found in Mexico. 

MALACLEMYS TERRAPIN LITTORALIS Hay 

Malaclemmys littoralis Hay, Bull. U. S. Bur. Fish., vol. 24, 1904, p. 18, pis. 8, 9, 

12, figs. 2, 3. 
M[alaclem7nys] terrapin littoralis, Carr, Copeia, 1946, p. 172. 

Type.— U.S. l<i.M. No. 33913; W. P. Hay collector. 
Type locality. — Rockport, Tex. 

Range. — Coast of Texas southward possibly to the Yucat&n Penin- 
sula. Recorded only from "Mexico" (Strauch). 



< 



CHECKLIST OF REPTILES OF MEXICO 37 

Genus CLEMMYS Ritgen 

Clemmys Ritgen, Nova Acta Acad. Leopold.-Carol., vol. 14, pt. 1, 1828, p. 272. 

Genotype. — Testudo punctata Schoepff [= Clemmys guttata (Schnei- 
der)]. 

Range. — Southern Europe, northwestern Africa, southern China 
and Japan, extreme western and the eastern third of the United States. 

Species. — Fifteen species and subspecies are recognized, five in 
North America, one of which enters Mexico. 

CLEMMYS MARMORATA PALLIDA Seeliger 

Clemmys marmorata pallida Seeliger, Copeia, 1945, pp. 158-159. 

Type. — Mus. Vert. Zool. No. 6716; J. E. Law collector. 

Type locality. — Lower Coyote Creek, near Alamitos, Orange County, 
Cahf. 

Range. — Central California near the mouth of the Sacramento River 
south to northwestern Baja California. Recorded only from northern 
Baja California. 

Subclass Lepidosauria Romer 

Lepidosauria Romer, Vertebrate paleontology, ed. 2, 1945, p. 595. 

Orders. — Two orders are represented by living members; one (Rhyn- 
chocephalia) is restricted to certain islands off North Island, New Zea- 
land, and the other is world-wide in distribution. 

Order SQUAMATA Oppel 

Squamata Oppel, Die Ordnungen, Familien und Gattungen der Reptilien . . . , 
1811, p. 14. 

Suborders. — Three living suborders are recognized: the Sauria, Ser- 
pentes, and Amphisbaenia. 

Suborder Amphisbaenia Gray 

Amphisbaenia Gray, Catalogue of the tortoises, crocodilians and amphisbaenians 
in British Museum, 1844, p. 68 (as an order). 

Families. — Two, only one of which occurs in Mexico. When the 
anatomy of the forms of amphisbaenians is better known, it is probable 
that several other families will be recognized. 

Range. — West Indies, tropical parts of South America north to 
Arizona, Florida, and Baja California; Spain, Turkey through Africa. 

Family BIPEDIDAE Stejneger 

Bipedidae Stejneger, Science, vol. 21, 1905, pp. 157-158. 

Genera. — One. For the present we prefer to refer all Mexican forms 
to the single genus Bipes despite the differences in digital structure. 



38 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Studies on the cranial anatomy may reveal differences not now 
apparent. 

Range. — The southern tip of Baja California, Arizona, the Rio 
Balsas Valley, and southern Guerrero. 

Genus BIPES Latreille 

Bipes Latrkille, Histoire naturelle des reptiles, vol. 2, 1802, p. 90. 

Bimanus Oppel, Die Ordnungen, Familien und Gattungen der Reptilien . . ., 

1811, p. 18. 
Hemichirotes DuGfes, in Cope, Amer. Nat., vol. 28, 1894, p. 436 (type, Hemichirotes 

tridactylus Duges). 
Euchirotes Cope, ibid, (type, Euchirotes biporus Cope). 

Genotype. — Bipes canaliculaius Bonnaterre. 

Range. — Baja California, Guerrero, southeastern Arizona. 

Species. — Three. 

KEY TO SPECIES OF BIPES 

1. Digits 5 2 

Digits 3, clawed tridactylus (p. 38) 

2. All digits equally developed, all generally with claws; 6 preanal pores. 

canaliculatus (p. 38) 
One digit smaller and lacking claw; 2 preanal pores biporus (p. 38) 

BIPES TRIDACTYLUS (Dugfes) 

Hemichirotes tridactylus DuGfcs, in Cope, Amer. Nat., vol. 28, 1894, pp. 436-437, 
fig. 6; La Naturaleza, ser. 2, vol. 2, 1894, pp. 411-412, pi. 20. 

Bipes tridactylus, Smith and Neckeh, Anal. Esc. Nac. Cien. Biol., vol. 3, 1943, 
pp. 196-197. 

Type. — Alfredo Duges Mus., Guanajuato, Guanajuato, Mexico. 
Type locality. — Tecpan de Galeana, Guerrero. 

Range. — Known only from type locality (only the type specimen 
known). 

BIPES BIPORUS (Cope) 

Euchirotes biporus Cope, Amer. Nat., 1894, p. 437, figs. 5-5e; Ann. Rep. U. S. Nat. 

Mus., 1898 (1900), pp. 680-682, fig. 140. 
Bipes biporus, Stejnegek and Barbour, Check list of North American amphibians 

and reptiles, 1917, p. 72. 

7V2>e.— U.S.N. M. No. 8568, cotypes (?), T. H. Street collector, and 
No. 12599, 12 specimens, L. Belding collector. 

Type locality. — Cape San Lucas, Baja California. 

Range. — The cape region of southern Baja California. Reported 
from La Paz, Cape San Lucas. 

BIPES canaliculatus Bonnaterre 

B{ipes\ canaliculatus Bonnaterre, Encyclopedic m^thodique, Erp^tologie, 1789, 

p. 68, pi. 12. 
Bipes canaliculatus, Sonnini and Latreille, Histoire naturelle des reptiles, vol. 2, 

1801, p. 90.— Smith, Journ, Washington Acad. Sci., vol. 39, 1949, pp. 42-43. 



CHECKLIST OF REPTILES OF ME'XICO 39 

Chiroles canaliculatus, Merrem, Tentamen systematis amphibiorum, 1820, p. 
161. — BocouRT, Mission scientifique au Mexique . . ., fitudes sur les reptiles, 
livr. 7, 1881, pp. 487-488, pi. 2lC, figs. 8, 8a-c.— Boulenger, Catalogue of 
the lizards in the British Museum, vol. 2, 1885, p. 432. 

Lacerta lumbricoides Shaw, Naturalists' Miscellany, vol. 6, 1795, pi. and text No. 
212 (type locality, Mexico, here restricted to Mexcala, Guerrero; type origi- 
nally in British Museum), 

Lacerta mexicana Donndorff, Zool. Beitrage zur 13. Ausg. Linneschen Natur- 
system, vol. 3, 1798, p. 135 (here restricted to Mexcala, Guerrero). 

Chalcides sulcata Suckow, Anfangsgriinde der theoret. und angewandten Natur- 
geschichte der Thiere, vol. 3, 1798, p. 147 (here restricted to Mexcala, 
Guerrero) . 

Chamaesaura propus Schneider, Historia amphibiorum naturalis et literaria, 
fasc. 2, 1801, p. 211 (here restricted to Mexcala, Guerrero). 

Type, — Not known. 

Type locality. — IMexico, here restricted to Mexcala, Guerrero. 
Range. — Kno^vn in the basin of Rio Balsas, Guerrero. Reported 
from Mexcala, Tecuaiziapan, and Balsas. ^^ 

Suborder Sauria Macartney 

Sauria Macartney, in Ross's translation Cuvier's Lecons d'anatomie compar^e, 
vol. 1, 1802, table 3. 

Families. — Nineteen, of which 10 occur in Mexico. 

KEY TO MEXICAN FAMILIES OF SAURIA 

1 . Neither forelegs nor hind legs evident externally 2 

Both forelegs and hind legs visible externally 3 

2. No eye opening; eye scarcely or not visible below lateral head scales; one 

unpaired median scale between frontal and rostral. Anelytropsidae (p. 170) 

Eye opening present, although small; a pair of scales on dorsal surface of head 

between frontal and rostral Anniellidae (p. 208) 

3. No movable eyelids; eyes permanently open 4 

Eyelids present, movable 6 

4. All scales around middle of body uniform, flat, cycloid, smooth, in about 13 

rows Teiidae 20 (p. 170) 

Not so 5 

5. Head covered with large, flat plates; ventral scales large, quadrangular, 

abruptly differentiated from granular lateral scales. .Xantusiidae (p. 151) 
Dorsal surface of head covered with granular scales; ventral scales smaller, 
rounded, not abruptly differentiated from granular lateral scales. 

Gekkonidae (p. 40) 

6. All scales around middle of body perfectly smooth (sometimes finely striated), 

cycloid, more or less equal in size (those in the middorsal row or pair or 

rows may be widened) 7 

Not so__ 8 

7. Only one scale in contact with frontal anteriorly; five supraoculars. 

Angxiidae 21 (p. 194) 

i« A record for "Morelia?," Michoacdn (DugSs, La Naturaleza, ser. 2, vol. 2, 1896, p. 480), is unacceptable 
until verified. 
"> Oymnophthaimus only. 
" Diploglossui only. 



40 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Two or three scales in contact with frontal anteriorly; three or four supra- 
oculars Scincidae (p. 155) 

8. A granular fold along sides of body, abruptly differentiated from much 

enlarged dorsal and ventral scales Anguidae (p. 194) 

Not so 9 

9. Ventral scales large, quadrangular, in 8 longitudinal series; dorsals granular. 

Teiidae (p. 170) 
Ventral scales smaller, in more numerous rows 10 

10. Ventral scales quadrangular; scales on top of head small, more or less uniform, 

tubercular; dorsal scales granular with numerous enlarged tubercles. _ 11 

Ventral scales pointed or rounded, not quadrangular; body and head scales 

as described or not 12 

11. Digits on hind leg of nearly equal length; enlarged tubercles covering most of 

dorsal surface of body; a series of about four large, paired postmentals on 

midline of chin, immediately back of mental Helodermidae (p. 192) 

Digits on hind leg varying greatly in length, fourth toe three times as long 
as fifth; enlarged tubercles on body much less numerous, not occupying as 
great an area as granules; no enlarged postmentals. _Xenosauridae (p. 207) 

12. Head and most of body, except bellj^, covered with very minute granules; no 

parietal "eye"; no keels or tubercles along ventral surfaces of digits. 

Qekkonidae (p. 40) 

Head covered with larger scales; a parietal "eye" usually visible; at least 

one tubercle, or several keels, on most or all of lamellae on ventral surfaces 

of digits Iguanidae (p. 53) 

Family GEKKONIDAE Stejneger 

Geckonidae Boulenger, Catalogue of the lizards in the British Museum, vol. 

1, 1885, p. 3. 
Gekkonidae Stejneger, U. S. Nat. Mus. Bull. 58, 1907, p. 164. 

Subfamilies. — Studies by Noble (Amer. Mus. Nov., No. 4, 1921, 
pp. 1-16) and by M. A. Smith (Rec. Indian Mus., 1933, p. 16) indi- 
cate that neither the Eublepharidae nor the Uroplatidae are worthy 
of either family or subfamily distinction. 

Genera. — About 75 genera are commonly recognized. 

Range. — World-wide, in tropical and semitropical areas. 

Remarks. — Eight genera of this family occur in Mexican territory. 
Three of these have probably been introduced accidentally from other 
parts of the world. Peropus, with a single representative, arrived on 
the west coast of Mexico probably from the Philippines. Hemidac- 
tylus, with three species, has a somewhat more complicated history. 
One species reached the west coast, probably from the Philippines 
(frenatus), a,nother came across the Atlantic to the east coast from 
Europe or Africa (turcicus), and a third (mabouia) probably came 
from somewhere in the West Indies or, not impossibly, from Mada- 
gascar or South Africa where it also occurs. Aristelliger undoubtedly 
was imported from the West Indies. A fourth genus, Gonatodes, has 



CHECKLIST OF REPTILES OF MEXICO 41 

only recently been reported in Mexico, and may possibly have been 
transported by man from Central America. 

It would appear from collections that all these genera have very 
limited or discontinuous distribution in Mexico. 

The genera Phyllodactylus, Sphaerodactylus, and Thecadactylus very 
probably reached Mexico without the aid of man. 

KEY TO MEXICAN GENERA OF GEKKONIDAE 

1. Eyelid fully developed, body covered with small cycloid somewhat imbricating 

scales, with or without somewhat larger tubercles on body or tail; a few 
preanal pores present; digits cylindrical, never distinctly widened. 

Coleonyx (p. 41) 
Eyelid rudimentary, not capable of covering eye 2 

2. Under surface of digits unwidened, covered with flat smooth scales, lacking 

leaflike lamellae; no preanal or femoral pores; diminutive size. 

Gonatodes (p. 45) 

A part of under surface of digits with widened lamellae present; femoral pores 

present or absent; size variable 3 

3. A single widened terminal lamella on digits with the claw lateral to it; no 

femoral pores; diminutive in size Sphaerodactylus (p. 52) 

Digits with one pair or several pairs of widened lamellae 4 

4. A single terminal pair of lamellae, the claw between them; size larger; preanal 

and femoral pores absent in Mexican forms Phyllodactylus (p. 46) 

Part of under surface of digits widened with a single or double series of lamellae, 
usually not present under terminal joint; pores present or absent 5 

5. Subdigital lamellae undivided, in a single row Aristelliger (p. 51) 

Subdigital lamellae in two rows, divided medially 6 

6. Tail with enlarged, sometimes spiny tubercles, forming longitudinal rows or 

transverse whorls; body with at least some enlarged tubercles, which may be 

flat or trihedral; femoral or preanal pores present Hemidactylus (p. 49) 

Tail and body lacking spines or enlarged tubercles; pores present or absent. _ 7 

7. Tail with flat ventral surface, serrated lateral edge, and a median row of en- 

larged scales on ventral surface; a long, doubly curved series of femoral 

pores; a fold of skin on posterior side of thigh and leg Peropus (p. 51) 

Tail rounded, tapering, covered with imbricate scales, a little larger on under 
surface; no femoral or preanal pores; no skin fold on hind leg. 

Thecadactylus (p. 49) 

Genus COLEONYX Gray 

Coleonyx Gray, Ann. Mag. Nat. Hist., vol. 16, 1845, p. 162. — Klauber, Trans. 

San Diego Soc. Nat. Hist., vol, 10, No. 11, 1945, pp. 133-216. 
Brachydachjliis Peters, Monatsb. Akad. Wiss. Berlin, 1863, p. 41 (type, B. 

mitratus Peters). 

Genotype. — Coleonyx elegans Gray. 

Range. — The genus extends from Southwestern United States to 
Panama. 

Species. — Five, with a total of 12 forms. Four species and 10 
forms are known or are to be expected in Mexico. 



42 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

KEY TO MEXICAN FORMS OF COLEONYX 22 

1. Dorsal surface covered with small granular scales, without larger tubercular 

scales 2 

Dorsal surface with larger tubercles intermingled with the small granular 
scales 9 

2. Two light and 3 dark bands between level of arm and leg insertions. 

fasciatus (p. 43) 

Usually 4 or parts of 4 dark bands between levels of arm and leg insertions, or 

the bands broken up into spots 3 

3. Preanal pores small, usually 4 in number (rarely more or less), the pore-bearing 

scales separated medially; maximum snout-vent measurement about 59 

mm brevis (p. 43) 

Preanal pores larger, usually 6 to 10, the series continuous, not separated 
medially; maximum snout- vent measurement 77 mm 4 

4. Usually 4 or less postmental scales touching mental .variegatus slevini (p. 45) 
Usually 5 or more postmental scales touching mental 5 

6. Preanal pores in males usually 8 or more variegatus bogerti (p. 44) 

Preanal pores in males usually 7 or less 6 

6. Dark transverse body bars in the adults considerably wider than the light 

interspaces 7 

Dark transverse body bars in adults about equal to or narrower than the 
light interspaces ; or bars obsolete and replaced by spotting 8 

7. A middorsal light longitudinal line usually splitting the dorsal body bars in 

the adults variegatus sonoriensis (p. 45) 

No middorsal light longitudinal line; adults with longitudinal edges of the 
dark body bars even, with narrow uniform interspaces. 

variegatus peninsularis (p. 44) 

8. Dark body bands in the adults unicolor; top of head unicolor; nuchal light loop 

narrow and clear variegatus abbotti (p. 44) 

Dark body bands in the adults with light centers producing a double-barred 
effect, or bars obsolete and replaced by spotting; top of head spotted; nuchal 
light loop irregular or obsolete variegatus variegatus (p. 44) 

9. Enlarged supranasals ^^ separated by a median circular scale. 

elegans nemoralis (p. 43) 
Enlarged supranasals in contact elegans elegans (p. 42) 

COLEONYX ELEGANS ELEGANS Gray 

Coleonyx elegans Gray, Ann. Mag. Nat. Hist., vol. 16, 1845, p. 162. 

Coleonyx elegans elegans, Klatjber, Trans. San Diego Soc. Nat. Hist., vol. 10, 

1945, pp. 191-195, map. 
G[ymnodactylus\ scapularis A. Dum:6ril, in Dum^ril and Dum^ril, Catalogue 

na^thodique de la collection des reptiles, 1851, pp. 45-46 (Pet6n, Guatemala, 

here restricted to La Libertad; Mus. Hist. Nat. Paris). 
Gyinnodactylus coleonyx A. Dumeril, Arch. Mus. Hist. Nat. Paris, vol. 8, 1858, 

p. 483 (same type as scapularis). 

Type. — Brit. Mus. Nat. Hist., male spec; Mr. Dysou collector. 



" Adapted from Klauber, op. cit., pp. 205-206. 

*> We believe the scale above nostril should be designated supranasal rather than prenasal; and those scales 
following the mental should be postmental rather than gular, since the latter term refers to throat or gullet, 
rather than chin. 



CHECKLIST OF REPTILES OF MEXICO 43 

Type locality. — Belize, British Honduras. 

Range. — Atlantic slopes from central Veracruz through the Yucatan 
Peninsula, including Honduras and northern Guatemala; on Pacific 
slopes in Chiapas and undoubtedly adjacent Guatemala. Reported 
from Veracruz: Jalapa, Potrero Viejo, San Lorenzo, Orizaba, Tezonapa, 
Zempoala, Presidio, San Juan Cuatotolapam ; Oaxaca: San Crist6bal, 
Cosolapa; Tabasco: Teapa, Tenosique; Campeche: Tuxpefia Camp, 
Apasote, Encarnaci6n; Yucatan: Chichen Itza, Mayapan, Puz, Gon- 
gora Caves, Ziz, Xkyc Cave; Chiapas: La Esperanza, Piedra Parada; 
Quintana Roo: Mujeres Island. 

COLEONYX ELEGANS NEMORAUS Klaaber 

Coleonyx elegans, Gadow, Proc. Zool. Soc. London (vol. 2), 1905, p. 212. 
Coleonyx elegans nemoralis Ivlauber, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 195-199. 

Type.— EB.T-HUS No. 10509; Hobart M. Smith collector. 

Type locality. — Hacienda Paso del Rio, Colima. 

Range. — Colima to southeastern Oaxaca (probably also Michoac§,n). 
Probably confined to the coastal area of these states. Reported from 
Guerrero: Cocoyul, Agua del Obispo, 4 to 5 miles north of Acapulco; 
Michoacdn: vicinity of JoruUo (?); Oaxaca: (said to be intergrades 
with elegans) Tehuantepec, Mixtequilla, Tapanatepec, etc. 

COLEONYX FASCIATUS (Boulenger) 2« 

Eublepharis fasciatus Boulenger, Catalogue of the lizards in the British Museum, 
vol. 1, 1S85, p. 423. — GtJNTHER, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1893, p. 84, pi. 31, fig. A. 

Coleonyx fasciatus, Taylor, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 203-205.— 
Klatjber, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 182-184. 

Type. — Brit. Mus. Nat. Hist.; A. Forrer collector. 

Type locality. — Ventanas, Durango. 

Range. — Known from western Durango, and Sinaloa. The only 
known specific locality other than the type locality is Sinaloa: 10 
miles south of Presidio. 

COLEONYX BREVIS Stejneger 

Coleonyx hrevis Stejxeger, North Amer. Fauna, No. 7, 1893, pp. 163-164. — 
Smith, Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 301-314, fig. IB, pi. 1, 
figs. 2-3.— Klauber, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 
184-191. 

Type.— v. S.'N.M. No. 13627; Mr. Marnock collector. 
TyjJe locality. — Helotes, Bexar County, Tex. 

Range. — The Rio Grande Valley, from northern New Mexico to 
southern Texas, and southward through eastern Chihuahua and 

" Bogert and Oliver (Bull. Amer. Mus. Nat. Hist., vol. 83, 1945, p. 330) suggest the possibility of inter- 
gradation of this form with variegalm. There is no evidence whatsoever that such is the case. See Klauber, 
loc. ett. 

861316—50 i 



44 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

western Tamaulipas to southern Coahuila and extreme eastern 
Durango. Reported from Tamaulipas: Mier; Nuevo Leon: 5 miles 
south of Sabinas Hidalgo, near China, Cienega de Flores, Mamulique 
Pass; Coahuila: Monclova, Saltillo; Durango: 6 miles northeast of 
Pedriceiia. 

COLEONYX VARIEGATUS VARIEGATUS (Baird) 

Stenodactylus variegatus Baird, Proc. Acad. Nat. Sci. Philadelphia, vol. 10, 1858, 

p. 254. 
Coleonyx variegatus, Cope, Proc. Acad. Nat. Sci. Philadelphia, vol. 18, 1866, pp. 

125, 310.— Smith, Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 301-314. 
Eublepharis variegatus, Boulenger (part), Catalogue of the lizards of the British 

Museum, vol. 1, 1885, pp. 233-234. 
Coleonyx variegatus variegatus, Klauber, Trans. San Diego Soc. Nat. Hist., vol. 

10, 1945, pp. 138-153, map. 

IVpe.— U.S.N.M. No. 3217 (now lost). 

Type locality. — Colorado Desert (fide Klauber, loc. cit.), here 
restricted to Winterhaven, Calif. 

Range. — Southern Nevada south through eastern California and 
western Arizona to northeastern Baja California and northwestern 
Arizona. Reported from Baja California: San Felipe, Colorado River 
Delta, Colorado Desert; Sonora: Paso McDougall, Sierra Blanca, 
Punta Pefiasco, etc. 

COLEONYX VARIEGATUS ABBOTTI Klauber 

Coleonyx variegatus abbotti Klauber, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 154-159. 

Type. — No. 34790, collection of L. M. Klauber; William Moore 
collector. 

Type locality. — Proctor Valley, between Jamul and Upper Otay 
Reservoir, San Diego County, Calif. 

Range. — Southwestern California and northwestern Baja California. 
Reported from Baja California: Ensenada, 65 miles southeast of 
Tecate, San Jose (lat. 31°), ? Calmalli, Isla de Cedros. 

COLEONYX VARIEGATUS BOGEBTI Klauber 

Coleonyx variegatus bogerti Klauber, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 176-182. 

Type.—L. M. Klauber No. 32486; Lee W. Arnold collector. 
Type locality. — Xavier, Pima County, Ariz. 

Range. — Southeastern Arizona and presumably adjacent Sonora 
and New Mexico. Not yet recorded from Mexico. 

COLEONYX VARIEGATUS PENINSULARIS Klauber 

Coleonyx variegatus peninsularis Klauber, Trans. San Diego Soc. Nat. Hist., 
vol. 10, 1945, pp. 160-162. 

Type. — Mus. Comp. Zool. No. 37210; Miguel L. Cornejo collector. 



CHECKLIST OF REPTILES OF MEXICO 45 

Type locality. — La Paz, Baja California, Mexico. 
Range. — Eastern side of southern Baja California. Reported from 
Baja California: San Jose del Cabo, between Loreto and Comondii. 

COLEONYX VARIEGATUS SLEVINI Klauber 

Coleonyx variegalus slevini Klauber, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 167-171. 

Type. — California Acad. Sci. No. 51697; J. R. Slevin collector. 
Type locality.— ''South Santa Inez Island" (lat. 27° W.), Gulf of 
California (Baja California coast). 

Range. — Isla Santa Inez and Isla de San Marcos, Gulf of California. 

COLEONYX VARIEGATUS SONORIENSIS Klauber 

Coleonyx variegalus sonoriensis Klauber, Trans. San Diego Soc. Nat. Hist., vol. 
10, 1945, pp. 162-166. 

Type. — Mus. Zool. Univ. Michigan No. 72140; Morrow J. Allen 
collector. 

Type locality. — Five miles southeast of Hermosillo, Sonora. 

Range. — Known only from central western Sonora. Reported 
from La Posa 10 miles northwest of Guaymas, Tepoca Bay, Sierra 
Alamo (30 miles west of Caborca), Isla de Tibur6n. 

Genus GONATODES Fitzlnger 

Gonatodes Fitzinger, Systema reptilium, 1843, pp. 91-92, 

Genotype. — Gymnodactylus albogularis Dumeril and Bibron =^Gona- 
todes albogularis (Dumeril and Bibron). 

Range. — Tropical America; West Indies. Species in South Asia, 
Africa, and Malaya formerly regarded as members of the genus are 
now placed in Cnemaspis. 

Species. — ^About 10, of which only one occm-s m Mexico. 

GONATODES FUSCUS (Hallowell) 

Stenodadylus fuscus Hallowell, Journ. Acad. Nat. Sci. Philadelphia, ser. 2, 

vol. 3, 1855, p. 33. 
Gymnodactylus fusctis, A. Dumeril, Arch. Mus., vol. 8, 1856, p. 477. — Bocourt, 

Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 2, 1873, 

pp. 48, 49, pi. X, fig. 5, 5a, 5b, 5c. 
[Gonatodes albogularis] fuscus, Boulenger, Catalogue of the lizards in the British 

Museum, ed. 2, vol. 1, 1885, pp. 59-60. 
Gonatodes fuscus, Stejneger, Pioc. U. S. Nat. Mus., vol. 53, 1917, pp. 264-265. — 

Smith, Copeia, 1944, p. 187. 
Goniodactylus hraconnieri O'Shaughnessy, Ann. Mag. Nat. Hist., ser. 4, vol. 16, 

1875, p. 265 {fide Boulenger; type locality, Baranquilla, Colombia; Brit. 

Mus. Nat. Hist.; Mr. Rippon collector). 

Type. — Acad. Nat. Sci. Philadelphia? 

Type locality. — Nicaragua, here restricted to Rama. 



46 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Range. — Huixtla, Chiapas (only definite Mexican record), south- 
ward in Central America to northwestern South America; Jamaica 
(probably introduced); Florida (probably introduced). 

Genus PHYLLODACTYLUS Gray 

Phyllodactylus Gray, Spicilegia zoologica, 1830, p. 3. 

Discodactylus Fitzinger, Systema reptilium, 1843, p. 95 (type, Phyllodactylus 

pulcher Gray). 

Genotype. — Phyllodactylus pulcher Gray. 

Range. — California to and including most of South America; 
Galapagos Islands, southern Asia, Africa, Madagascar, Australia, 
et cetera. 

Species. — ^About 48 species are known; 9 are known from Mexico.^* 

KEY TO MEXICAN SPECIES OF PHYLLODACTYLUS 

1. Dorsal squamation consisting of minute rounded scales without admixture of 

larger tubercles; no enlarged tubercles on tail; small, length 52 mm. 

unctus (p. 47) 

Dorsal squamation with an admixture of larger tubercles; larger tubercles 

present or absent on tail 2 

2. Tail lacking any trace of enlarged tubercles on dorsal surface 3 

Tail with smaller or larger tubercles intermixed with dorsal lepidosis 4 

3. Scales on head between middle of orbits 15-17 ^®; on dorsal and lateral surface 

14 to 16 rows of large, very distinct trihedral tubercles; smaller, 44 + mm. 

bordai (p. 47) 

Scales between orbits 19-24; dorsal tubercles low, small, rather indistinct, 

arranged in 8 to 12 rows; larger, 90 mm delcampi (p. 47) 

4. Two rows of tubercles on dorsal surface of tail (greatly reduced or absent in 

homolepidurus) 5 

More than two rows of enlarged tubercles, at least on basal third of tail 6 

5. Scales between middle of orbits 22-26; tubercles on back and tail large, well 

defined muralis (p. 47) 

Scales between middle of orbits 19-24; tubercles on back much smaller than in 
muralis, the caudal tubercles reduced so that they can be discerned only 
with difficulty; 67 mm homolepidurus" (p. 48) 

»» Phyllodactylus pulcher Gray, Spicilegia Zoologica, from "Tropical America," may be found to occur in 
Mexico. It is distinct from all otliers listed here by having 20 to 22 longitudinal series of dorsal tubercles 
(rather than 14 to 16), and larger ventral scales, in 22 longitudinal and 53 transverse series (rather than 25 to 
30 and 65, respectively). 

We likewise are unable to place Phyllodactylus mentalis Werner (Jahrb. Hamb. Wiss. Anstalten, vol. 27, 
No. 2, 1910, p. 6; Zool. Mus. Hamburg; L. von Poppinghausen coll.; Central America or Mexico). Mosauer 
(Copeia, 1936, p. 144) includes a few notes on the type. 

'• Not counting scales on vestigial eyelids. 

" Bogert and Oliver (Bull. Amer. Mus. Nat. Hist., vol. 83, 1945) note that at least nine species of Phyllo- 
dactylus have been described from western Mexico in recent years and that certain of these are allopatric and 
their characters strongly suggest that a more accurate "indication of the biological conditions that exist" may be 
obtained by considering them as subspecies or races of the same species. They proceed to place homolepi- 
durus as a subspecies under tuberculosus. Since the latter name cannot be fixed at the present time, we do 
not follow this proposal. Unfortunately, these authors have not examined most of the described forms and 
have overlooked the fact that at least four of the species may occur in the same general locality. They state 
that "the lizards of this genus exhibit considerable variation." These forms are probably no more variable 
than other lizards. For many years most North American species of Phyllodactylus were classified in muse- 
ums as "tuberculostis," and examination of such material might create the erroneous idea that "tuberculosus" 
was an extremely variable species. 



CHECKLIST OF REPTILES OF MEXICO 47 

6. Four distinct longitudinal rows of enlarged tubercles at least on basal fourth of 

tail, usually in rather regular transverse rows 7 

At least six distinct rows of enlarged tubercles on basal part of tail, not in- 
variably arranged in regular transverse rows 8 

7. Scales between orbits 12-16; dorsal and posterior part of thigh with numerous 

large conical tubercles; color darker above, cream color below; 80 mm. 

lanei (p. 48) 
Similar to above but tubercles on back heavier, higher; head deeper, less 
depressed, color lighter; yellow below; 66 mm. (1 specimen). 

magnatuberculatus (p. 48) 

8. Scales between orbits 25-30; 95 mm magnus (p. 48) 

Scales between orbits much fewer tuberculosus (p. 49) 

PHYLLODACTYLUS UNCTUS (Cope) 

Diplodactylus unctus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 102. 
Phyllodactylus (Diplodactylus) unctus, Bocourt, Mission scientifique au Mexique 

. . ., Etudes sur les reptiles, livr. 2, 1873, pp. 43-44. 
Phyllodactylus unctus, Taylor, Univ. Kansas Sci. Bull., vol. 28, 1942, pp. 96-99, 

fig. 2, A-D. 

Type.—U.S.lSiM. No. 5304. 

Type locality. — Cape San Lucas, Baja California. 

Range. — Southern Baja California. Keported from Baja California: 
San Jose del Cabo ; Isla Partida near Isla Espiritu Santo, Isla Gallina, 
and Isla San Francisco. 

PHYLLODACTYLUS DELCAMPI Mosaner 

Phyllodactylus delcampi Mosauer, Copeia, 1936, pp. 141-146. 

Type. — Mus. Comp. Zool. No. 41238; W. Mosauer collector. 
Type locality. — Tierra Colorada, Guerrero. 
Range. — Kno^\^^ only from the type locality. 

PHYLLODACTYLUS BORDAI Taylor 

Phyllodactylus bordai Taylor, Univ. Kansas Sci. Bull., vol. 28, pt. 1, 1942, pp. 93- 
96, fig. 1. 

?V^e.— EHT-HMS No. 27732; E. H. Taylor collector. 

Type locality. — Six miles north of Taxco, Guerrero. 

Range. — Northern Guerrero; known only from the type locality. 

PHYLLODACTYLUS MURAUS Taylor 

Phyllodactylus muralis Taylob, Univ. Kansas Sci. Bull., vol. 26, 1939 (1940), 
pp. 543-547, fig. 7. 

Type.— EB.T-HMS No. 10902; H. M. Smith collector. 
Type locality. — Totolapam, Oaxaca. 

Range. — Oaxaca. Reported from Totolapam, Tehuantepec, San 
Ger6nimo, Salina Cruz. 



48 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

PHYLLODACTYLUS HOMOLEPIDURUS Smith 

Phyllodactylus homolepidurus Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 

121-125, pi. 25, fig. 2, text fig. lA. 
Phyllodactylus tuberculosus homolepidurus, Bogert and Oliver, Bull. Amer. Mus. 

Nat. Hist., vol. 83, art. 6, 1945, pp. 344-345. 

Type.— ERT-RMS No. 10853 (originally EHT field No. 146); E. 
H. Taylor collector. 

Type locality. — Five miles southwest of Hermosillo, Sonora. 

Range. — Southern Sonora and adjacent Chihuahua. Reported from 
Sonora: La Posa, 5 miles southwest of Hermosillo, Guirocoba; Chi- 
huahua: Batopilas. 

PHYLLODACTYLUS MAGNATUBERCULATUS Taylor 

Phyllodactylus niagnatuberculatus Taylor, Univ. Kansas Sci. Bull., vol. 26, 1939 
(1940), pp. 547-549. 

Type.— ERT-UMS No. 10995; H. M. Smith collector. 

Type locality. — Acapulco, Guerrero. 

Range. — Known only from the type locality. 

PHYLLODACTYLUS MAGNUS Taylor 

Phyllodactylus magnus Taylor, Univ. Kansas Sci. Bull., vol. 28, 1942, pp. 99- 
103, fig. 3. 

Ti/^e.— EHT-HMS No. 21783; E. H. Taylor collector. 

Type locality. — Tierra Colorada, Guerrero. 

Range. — Guerrero, Oaxaca, and Chiapas. Reported from Guerrero: 
Garrapatas, El Ocotito, Tierra Colorada, Agua del Obispo; Oaxaca: 
Tehuantepec, Oaxaca, Tres Cruces, Cerro Arenal, Caj6n de Piedra, 
Escurana; Chiapas: Tonala. 

PHYLLODACTYLUS LANEI 28 Smith 

Phyllodactylus land Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 125-132, 
fig. lb; pi. 25, fig. 3.— Taylor, Univ. Kansas Sci. Bull., vol. 26, No. 15, 
1939 (1940), pp. 541-542, fig. 6. 

Type.—EWT-RM^ No. 10942 (originally EHT field No. 146); E. 
H. Taylor and H. M. Smith collectors. 

Type locality. — Near Tierra Colorada, Guerrero, Mexico. 

Range. — Sinaloa and (?) western Durango ^^ southward to central- 
southern Guerrero, probably confined to the coastal areas up to an 
elevation of about 3,500 feet. Recorded from the states of Sinaloa 
(including the Tres Marias Islands), ? Durango, Nayarit, ? Jalisco, 
Colima, Michoacan, and Guerrero. 

2' The suggestion of common identity of P. lanei and P. tuberculosus by Mosauer is, we believe, unten- 
able (Mosauer, Copeia, 1936, p. 194). 

« A specimen of Phyllodactylus collected at Durango, Mexico by Dr. T. H. Webb (U. S. N. M. No. 3208) 
and figured by Balrd (Reptiles of the Boundary, U. S. and Mexican Boundary Surveys, 1859, p. 12, pi. 
23, figs. 1-8) may or may not belong to this species, since Durango is considerably higher than this species 
is known to range. 



CHECKLIST OF REPTILES OF MEXICO 49 

PHYLLODACTYLUS TUBERCULOSUS Wiegmann 3» 

Phyllodactylus Uiberculosus Wiegmann, Nova Acta Phys.-Med. Acad. Caes. 

Leop.-Carol., vol. 17, pt. 1, 1835, p. 241, pi. 18, fig. 2. 
t Phyllodactylus Xanti Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 103 

(Cape San Lucas, Baja California). 

Type. — ? Zool. Miis. Berlin; collector unknown. 
Type locality. — "Californien." 

Range. — ? Extreme southern part of California, south through Baja 
California. 

Genus THECADACTYLUS Gray 

Thecadactylus Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 198. 

Genotype. — Gecko rapicauda Houttuyn. 

Range. — Southern Mexico to northern South America (Peru); 
Antilles; islands north of Australia in Torres Strait. 
Species. — Two. 

THECADACTYLUS RAPICAUDUS (Houttuyn) 

Gekko rapicauda Houttuyn, Verh. Genootsch. wet. Vlissing., vol. 9, 1782, p' 

323, pi. 3, fig. 1. 
Thecadactylus rapicauda, Gray, Catalogue of the lizards in the collection of the 

British Museum, 1845, p. 146. — Boulenger, Catalogue of the lizards in the 

British Museum, vol. 1, 1885, pp. 111-112, 

TyjJe . — Unknown . 

Type locality. — American Islands, here restricted to Chichen Itzd, 
Yucatan. 

Range. — Southward from Yucatan through Central America to 
northwestern South America; Lesser Antilles. Reported from 
Yucatan: Chichen Itzd; Puz Cave, Oxkutzcab; Chalcxix Cave, Tekax. 
The Guadalajara reference of Cope (U. S. Nat. Mus. Bull. 32, 1887, 
p. 28) is very doubtfid. 

Genus HEMIDACTYLUS Oken 

Hemidactylus Oken, Isis, 1817, p. 1183 (based on Cuvier's Hemidactyle, R6gne 

animal, vol. 2, 1817, p. 47). 
Tachybates Fitzinger, Systema reptihum, 1843, p. 105 (type, "Hemidactylus 

mabuya Cuvier"). 
Pnoepus Fitzinger, Systema reptilium, 1843, p. 105 (type, " Heviidactylus javanicus 

Cuvier"). 

M We have here followed the idea that Baja California or California is the type locality. However, since 
there are two forms (one undescribed) in Baja California, one with a spiny tail, the other without spines 
on the tail, it is not certain to which the name must apply, inasmuch as the type has a reproduced tail. 
Direct comparison of the two species with the type of tubercutosus will be necessary to determine the proper 
application of the name on the basis of characters not now discernible from the description or figure. 

There are many reports of this form in the literature, and it is impossible to determine to what species 
many of them refer. Although for that reason many localities are thus not here cited, we have included 
every state from which "tuherculosus" has been recorded in the records for at least one species, basing our 
allocation either upon actual examination of specimens or a reasonably accurate geographic indication. 
There is one exception: Izucar de Matamoros, Puebla. 



50 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Genotype. — Oekko tuherculosus Daudin [= Hemidactylus mabouia 
Moreau de Jonnes]. 

Range. — Widespread in Africa, southern Eurasia, tropical America, 
Polynesia. 

Species. — About 60 known. Three are found in Mexico. 

KEY TO MEXICAN SPECIES OF HEMIDACTYLUS 

1. Preanal pores 4 to 10, forming a short angular series; no femoral pores; dorsal 

scales minute granules intermixed with large tubercles usually larger than 

interspaces between them; inner digits well developed turcicus (p. 50) 

Preanal and femoral pores forming a continuous series 2 

2. Small granules of the dorsal surface with only a few relatively small, convex 

tubercles, these usually on posterior part of body; pores 15-18 on each side, 

not separated mesially frenatus (p. 50) 

Small granules of the dorsal surface mixed with very numerous trihedral 
tubercles, irregularly scattered; pores from 15 to 30 on each side. 

raabouia (p. 50) 

HEMIDACTYLUS MABOUIA (Moreau de Jonnfes) 

Gecko mabouia Moreau de Jonnes, Bull. Soc. Philom., 1818, p. 138. 
Hemidactylus mabouia, Boulenger, Catalogue of the lizards of the British 
Museum, vol. 1, 1885, pp. 122-123. — Loveridge, Copeia, 1941, p. 248. 

Type.— Mns. Hist. Nat. Paris. 

Type locality. — "Antilles," here restricted to St. Vincent, Lesser 
Antilles. 

Range. — In Mexico known only from "Veracruz" (probably city). 
Widespread in the West Indies, and the northern and eastern coast of 
South America; Madagascar and Africa. The American localities are 
probably due to relatively recent introductions from Africa, or the 
West Indies. 

HEMIDACTYLUS FRENATUS Schlegel 

Hemidactylus frenatus Schlegel, in Dum6ril and Bibron, Erp^tologie generale 
. . ., vol. 3, 1836, pp. 366-368.— Taylor, Univ. Kansas Sci. Bull., vol. 26, 
1939, pp. 444. — Burt and Myers, Stanford Univ. Publ., biol. sci., vol. 8, 
1942, pp. 285-286.— Smith, Anal. Inst. Biol., vol. 14, 1943, p. 342.— Edgren, 
Nat. Hist. Misc., No. 55, 1950, p. 1. 

Type. — Leyden Mus. 

Type locality. — Java. 

Range. — Guerrero, in the region about Acapulco. Widespread in 
southern Asia, Pacific Islands, etc. Reported from Guerrero: Aca- 
pulco, Tierra Colorada. 

HEMIDACTYLUS TURCICUS TURCICUS (Linnaeus) 

Lacerta turcica Linnaeus, Systema naturae, ed. 10, 1758, p. 202, 
Hemidactylus turcicus, Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 132-133, 
pi. 23, fig. 2. — Grant and Deuel, Herpetologica, vol. 2, 1944, pp. 130-135. 
Hemidactylus turcicus turcicus, Loveridge, Copeia, 1941, p. 247. 



CHECKLIST OF REPTILES OF MEXICO 51 

Hemidactylus exsul Barbour and Cole, Bull. Mus. Comp. Zool., vol. 50, 1906, 
p. 148 (type locality, Progreso, Yucatdn; Mus. Comp. Zool. No. 7039). — 
Stuart, Copeia, 1934, No. 4, p. 185. 

Type . — Unknown . 

Type locality — "Habitat in Oriente," here restricted to Cairo, 
Egypt. 

Range. — Tamaulipas south to Yucatdn, distribution very probably 

discontinuous. Found in the West Indies, Florida, North Africa, 

and southwestern Asia. Reported from Tamaulipas: 12 miles east 

of Llera; Veracruz: no specific locality; Tabasco: Alvaro Obreg6n, 

Emiliano Zapata; Campeche: Ciudad del Carmen; Yucatan: Progreso. 

The Mexican records are due to modern introductions, probably from 

northern Africa. 

Genus ARISTELLIGER Cope 

Aristelliger Cope, Proc. Acad. Nat. Sci. Philadelphia, 1861, p. 496. 
Idiodactylus Bocourt, Mission scientifique au Mexique . . ., Etudes sur les 
reptiles, livr. 2, 1873, p. 41 (type, Idiodactylus georgeensis Bocourt). 

Genotype. — Aristelliger lar Cope. 

Species. — Six, aU West Indian except the one Yucatan form. 

Range. — West Indies and the peninsula of Yucatdn. 

ARISTELLIGER GEORGEENSIS (Bocourt) 

Idiodactylus georgeensis Bocourt, Mission scientifique au Mexique . . ., Etudes 

sur les reptiles, livr. 2, 1873, pp. 41-42, pi. 10, fig. 1. 
Aristelliger georgeensis, Schmidt, Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, 

1941, p. 490. 
Aristelliger praesignis, Boulenger, Catalogue of the lizards in the British 

Museum, vol. 1, 1885, pp. 146-147 (part). 
Aristelliger irregularis Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, p. 387 

(Cozumel Island; Ridgway coll.; U.S.N.M. No. 13903). 

Type. — Mus. Hist. Nat. Paris. 

Type locality. — St. George Island, near Belize, British Honduras. 
Range. — The eastern edge of the Yucatan Peninsula. Recorded in 
Mexico only from Cozumel Island (Quintana Roo). 

Genus PEROPUS Wiegmann'i 

Peropus WiEGMANN, Nova Acta Acad. Leop. -Carol., vol. 17, pt. 1, 1835, p. 238. 
Dactyloperus Fitzinger, Systema reptilium, 1843, p. 103 (type, Peropus peronii 

Dumdril and Bibron= Peropws mutilatus Wiegmann). 
Peripia Gray, Catalogue of the specimens of lizards in the British Museum, 

1845, p. 158 (type, Peripia peronii Dum6ril and Bihron^ Peropus mutilatus 

Wiegmann). 
Chalinocnemis Duoiis, La Naturaleza, vol. 6, 1883, p. 312 (type, Hemidactylus 

navarri Duges= Peropus mutilatus Wiegmann). 
Spasmocnemis Duofcs, loc. cit. (type as for preceding). 

Genotype. — Hemidactylus mutilatus Wiegmann. 

" We are inclined to maintain Peropus and Oehyra based on the different character of the subdigital 
lamellae. 



52 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Range. — Widespread in southern Asia from Ceylon to the Philip- 
pines; south through the Malay Archipelago to New Guinea; Sey- 
chelles; western Mexico. 

Species. — One form occurs in Mexico, probably introduced from the 
Philippines by man; about 10 others are known. 

PEROPUS MUTILATUS (Wiegmann) 

Hemidaclyhis (Peropus) jnutilatus Wiegmann, Nova Acta Acad. Leop. -Carol., 
vol. 17, 1835, p. 238. 

Peropus mutilatus, Smith and Necker, Anal. Esc. Nac. Cienc. Biol., vol. 3, 1943, 
pp. 197-199, pi. 3, fig. 3. 

Hemidaclylus Navarri Duces, La Naturaleza, vol. 6, 1883, pp. 309-312, pi. 7a 
(type locality, San Bias, Nayarit; 2 types in Alfredo Duges Museum, Guana- 
juato, Guanajuato.) 

Type. — Zool. Mus. Berlin?. 

Type locality. — Manila, Philippine Islands. 

Range. — Seaports of Nayarit and Sinaloa. Widespread in south- 
eastern Asia, Philippine Islands, Dutch Indies, to New Guinea and 
certain Pacific Islands. Reported from Nayarit: San Bias, Tepic; 
Sinaloa: Presidio, MazatMn. 

Genus SPHAERODACTYLUS Wagler 

Sphaerodadylus (part) Wagler, Naturliches System der Amphibien, 1830, p. 
143.— Barbour, Mem. Mus. Comp. Zool., vol. 47, 1921, pp. 215-279, pis. 
1-26. 

Genotype. — Sphaerodactylus sputator Cuvier. 
Range. — West Indies, southern Mexico, Central America. 
Species. — About 38 forms. Three are known or expected from 
Mexico .^^ 

KEY TO MEXICAN SPECIES OF SPHAERODACTYLUS 

1. Dorsal scales granular, keeled lineolatus (p. 52) 

Dorsal scales imbricate, smooth 2 

2. A single nuchal band; two distinct spots at base of tail, light spots at elbow 

and knee usually discernible glaucus glaucus (p. 53) 

A cream-colored nuchal band bordered bj black bands of equal width anteriorly 
and posteriorly; two narrow longitudinal light lines from collar to eye; in- 
distinct narrow light bands on back; spots on tail base absent or obsolescent; 
no light spots at knee and elbow in young or adult. 

glaucus torquatus (p. 53) 

SPKAERODACTYLUS lineolatus Lichtenstein 

Sphaerodadylus Imeolatus Lichtenstein, Nomenclator reptilium et amphibiorum 
musei zoologici berolinensis, 1856, p. 46. — Barbour, Mem. Mus. Comp. 
Zool., vol. 47, 1921, pp. 233-240, pi. 4, figs 1, 2, pi. 14, figs. 1-4. 

Type. — Zool. Mus. Berlin, tliree cotypes. 

'" Until new material is discovered in Mexico, the Mexican reports of Sphaerodactylus fantasfkus, an- 
thracinus, punctatissimus (cinereus), and sputator, are to be regarded as uncertain. The type locality of 
anthracinus is "Mexico," but possibly in error (see p. 213) . It would not be impossible for a part of these forms 
to occur, as recent introductions in Mexico. Banana fruits serve as a ready means of transportation. 



CHECKLIST OF REPTILES OF MEXICO 53 

Type locality. — Veraguas, Panamd. 

Range. — The eastern edge of the Yucatan Peninsula southeastward 
to Colombia. No definite records from Mexico. ^^ 

SPHAERODACTYLUS GLAUCUS GLAUCUS Cope 

Sphaerodactylus glaucus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 192. — 

BouLENGER, Catalogue of the lizards in the British Museum, vol. 1, 1885, 

p. 221, pi. 18, fig. 3.— Barbour, Mem. Mus. Comp. Zool., vol. 47, 1921. 

pp. 240-241, pi. 14, figs. 5-8.— Taylor, Univ. Kansas Sci. Bull., vol. 31, 

pt. 2, 1947, pp. 305-309, fig. 2. 
Sphaerodactylus glaucus glaucus, Smith, Journ. Washington Acad. Sci., vol. 39, 

1949, pp. 34-35. 
Sphaerodactylus inornatus Peters, Monatsb. Akad. Wiss. Berlin, 1873, p. 738 

(Zool. Mus. Berlin No. 4589; Uhde coll.; Mexico ^^ here restricted to Te- 

huantepec, Oaxaca. 

Type.— U.S.N.M. No. 6572, thi-ee cotypes (now one, No. 13570, 
in Mus. Comp. Zool.) ; Arthur Schott collector. 

Type locality. — Near Merida, Yucatan. 

Range. — Known from southern Veracruz to Guatemala and British 
Honduras, and possibly to Costa Rica. Reported from Oaxaca: Te- 
huantepec, Salina Cruz, Guengola Mountain; Tabasco: Teapa; 
Campeche: Tuxpena Camp, Ciudad del Carmen, Balchacaj, Apazote; 
Veracruz: Perez, Coatzacoalcos River; Yucatan: Merida. 

SPHAERODACTYLUS GLAUCUS TORQUATUS Strauch 

Sphaerodactylus torquatus Strauch, Mem. Acad. Sci. St. Petersburg, ser. 7, vol. 

35, 1887, p. 35.— Taylor, Univ. Kansas Sci. Bull., vol. 31, 1947, pp. 302-305, 

figs. 1-2. 
Sphaerodactylus glaucus torquatus, Smith, Journ. Washington Acad. Sci., vol. 

39, 1949, p. 35. 

Type. — Alus. Petrograd No. 3268, tlii-ee specimens; Hr. Salrain 
collector. 

Type locality. — "Mazatian, Mexico" (= Mazatlan, Oaxaca?). 

Range. — Uncertain. Known from the type locality and Cajon de 
Piedra, Oaxaca.^® 

Family IGUANIDAE Gray 

Iguanidae Gray, Philos. Mag., ser. 2, vol. 2, 1827, p. 56. 

Genera. — About 65 genera are commonly recognized; 21 occur in 
Mexico. 



" The species is included here on the strength of its occxirrence in British Honduras (Schmidt, Publ. 
Field Mus. Nat. Hist., zool. ser., vol. 22, 1941, p. 489), for although it has been cited from "Mexico" by- 
numerous authors, no definite record of its occurrence in that country Is known to us. Its existence in at 
least Quintana Roo is virtually a certainty. 

" It now seems most probable that this name was applied to a unlcolor intergrade between S. g. glaucus 
and S. g.torguahis such as those known to occur in the vicinity of Tchuantepec, Oaxaca. The entire descrip- 
tion, of both scutellation and coloration, fits glaucus, not lineolatus, although Barbour (op. cit., p. 238) places 
Peters's name in the synonymy of the latter species. See Smith, loc. cit., and Taylor, loc. cit., for 
discussion.— H. M. S. 

*' The subspecies does not occur in the vicinity of the city of Tehuan tepee, but only, so far as we are aware, 
near the coast southwest of Salina Cruz, where a village by the name of Mazatlan does exist. 



54 BULLETIN 199, XHSTITED STATES NATIONAL MUSEUM 

Range. — Extreme southwestern Canada and extreme southeastern 
New York, southward throughout the Americas; Bermuda Islands; 
West Indies; Revillagigedos Islands; Galapagos Islands; Madagascar 
(two genera); Fiji and Tonga Islands (one genus). 

KEY TO MEXICAN GENERA OF IGUANIDAE 

1. No femoral pores 2 

Femoral pores present, conspicuous in males, less well defined in females. _ 5 

2. Ventral lamellae on next to last phalanx of digits expanded laterally, forming 

a padlike structure; posterior part of head not produced Anolls (p. 55) 

Ventral lamellae on digits not expanded to produce a pad; posterior part of 
head more or less produced 3 

3. Infradigital lamellae with a single, tubercular keel Laemanctus (p. 69) 

Infradigital lamellae with several sharp keels 4 

4. Toes with a lateral fringe Basiliscus (p. 71) 

No fringe on toes Corythophanes (p. 68) 

5. A single, median, dorsal, longitudinal series of enlarged scales 6 

Scales in vertebral row not conspicuously if any larger than adjacent scales. 9 

6. Tail with whorls of much-enlarged, spiny scales, usually separated from each 

other by whorls of small scales 7 

Scales on dorsal and lateral surfaces of tail (except the vertebral row) subequal 
in size, small 8 

7. A group of much enlarged, spinose, protuberant scales on shank and some- 

times on thigh; basal caudal region (or entire tail in short- tailed species) 
highly modified, sometimes flattened and widened, with whorls of greatly 
enlarged scales separated from each other by no more than 1 row of small 

scales; total length less than 450 mm Enyaliosaurus (p. 75) 

No conspicuously enlarged scales on shank or thigh; basal caudal area much 
like rest of tail (which is always long), not flattened but cylindrical, with 
whorls of enlarged scales separated from each other by two or more rows 
of small scales throughout most of tail; total length often more than 450 
mm., reaching at least 1,200 mm Ctenosaura (p. 73) 

8. A large, circular shield below tympanum; a well-developed gular append- 

age Iguana (p. 72) 

No shield below tympanum; no gular appendage Dipsosaurus (p. 77) 

9. Head bearing bony spines posteriorly, or elevated somewhat in a projecting 

ridge Phrynosoma (p. 94) 

Head normal, not i^roduced posteriorly in a bony ridge or in spines 10 

10. Superciliary scales not imbricate, but juxtaposed, like other scales on head; 

rostral subdivided, with a median suture meeting lip Sauromalus (p. 79) 

Superciliary scales strongly imbricate, large; rostral not divided, no median 
suture reaching lip 11 

11. Sutures between supralabials diagonal, not vertical; mental very small, 

bordered posteriorly by a median postmental 12 

Sutures between supralabials vertical; mental larger, not bordered posteriorly 
by median postmental 14 

12. No ear opening Holbrookia (p. 81) 

A distinct ear opening 13 

13. Interparietal much smaller than ear opening, not more than twice as large 

as any other dorsal head scale; several enlarged auricular lobules; a patch 
of enlarged scales with projecting points on rear of femur Uma (p. 89) 



CHECKLIST OF REPTILES OF ]MEXICO 55 

Interparietal nearly as large as ear opening, or larger, much larger than 
any other dorsal head scale; no enlarged auricular lobules; no patch of 
enlarged scales on rear of femur Callisaurus (p. 85) 

14. No complete transverse gular fold, or if so body and tail compressed and 

dorsal scales nearly imiform, large, no more than 75 from interparietal to 

rear of thighs 15 

A complete transverse gular fold, marked by granular or reduced scales, 
much smaller than those preceding or following fold; dorsal scales very 
small, or at least not uniformly large and as few as 75 from interparietal 
to rear of thighs (except Uta squamata); body and tail flattened, never 
compressed even slightly 16 

15. A gular fold usually present; lateral scales granular; body and tail somewhat 

compressed Sator (p. 139) 

Gular fold never present; lateral scales not granular in those species with 
compressed tail Sceloporus (p. 105) 

16. Interparietal very large, about as large as or larger than ear opening 18 

Interparietal very small, a great deal smaller than ear opening 17 

17. Two scales bordering mental posteriorly between infralabials; head very broad, 

neck narrow Crotaphytus (p. 91) 

Three or more scales bordering mental posteriorly between infralabials; head 
elongate, neck not so sharply defined Qambelia (p. 93) 

18. Dorsal scales all small, granular, smooth, very little larger than lateral scales; 

enlarged supraoculars in more than one row; one or more blackish bars 

across shoulders; size frequently over 75 mm. snout to vent 19 

Dorsal scales, at least toward middle of back, larger than lateral scales, keeled; 
enlarged supraoculars in one row; no blackish bars across shoulders; size 
less than 75 mm. snout to vent 20 

19. Caudal scales smaller than ventrals on body, not mucronate; three or four 

black bands across body Petrosaurus (p. 90) 

Caudal scales large, mucronate, much larger than ventrals on body; a single 
narrow black band across shoulders Streptosaurus (p. 91) 

20. A longitudinal dorsolateral line or dermal fold, usually with some enlarged 

scales on it; usually a second, lateral fold; no small, rounded blue blotch 

behind axilla; frontal divided or not Urosaurus (p. 140) 

No longitudinal dorsolateral dermal fold; often a smaU, rounded, dark blue 
blotch behind axilla; frontal always divided Uta (p. 147) 

Genus ANOLIS Daudin 

Anolis Daudin, Histoire naturelle gen^rale et particuliftre des reptiles, year X 

(1802), pp. 50-51.— Barbour, Bull. Mus. Comp. Zool., vol. 78, 1934, pp. 

121-155. 
Dactyloa Wagler, Natvirliches System der Amphibien, 1830, p. 148 (type, Anolis 

viridis Wied). 
Tropidopilus Fitzinger, Systema reptilium, 1843, p. 66 (type, Anolis fusco- 

auraius d'Orbigny, monotype). 
Trachypilus Fitzinger, op. cit., p. 67 (type, Anolis sagraez Dumdril and Bibron). 
Pristicercus Fitzinger, op. cit., pp. 07-68 (type, Dactyloa biporcata Wiegmann). 
Ctenocercus Fitzinger, op. cit., pp. 17, 68 (type, Dactyloa buUaris Wagler — Anolis 

carolinensis Linnaeus). 
Gastrotropis Fitzinger, loc. cit. (type, Dactyloa nebulosa Wiegmann). 
Coccoessus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1862, p. 178 (type, Anolis 

pentaprion Cope). 



56 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Genotype. — Anolis bullaris Daudin {=Anolis carolinensis Voigt). 

Species. — Perhaps 350. Thirty-five forms are Usted here, but two 
are not known from any definite locality, four are known only from 
"Mexico," and the Mexican range of another is unknown. 

Range. — Southeastern United States to Brazil; West Indies. 

KEY TO MEXICAN SPECIES OF ANOLIS 

1. Tail strongly compressed 2 

Tail subcylindrical, no more than slightly compressed 4 

2. Two continuous rows of scales along middorsal line on tail, .barker! (p. 58) 
One row of scales along middorsal line of tail 3 

3. Ventrals keeled sagrei mayensis (p. 59) 

Ventrals smooth cozumelae (p. 59) 

4. Dorsal scales as large as or larger than ventrals, and both strongly keeled. 5 
Dorsal scales smaller than ventrals, or one or both sets of scales smooth; or, 

if body scales as above, occipital much larger than ear opening 8 

5. Supraorbital semicircles in contact or separated by a single series of scales. 

megapholidotus (p. 59) 
Supraorbital semicircles separated by 2 or 3 series of scales 6 

6. Dorsal and ventral scales subequal in size metallicus (p. 59) 

Dorsal scales distinctly larger than ventrals 7 

7. Tibia 75 percent to 85 percent length of head (to anterior border of ear) ; adult 

size smaller, about 36 mm.; dewlap brilliant red with a large purple spot 

at base in life humilis uniformis (p. 60) 

Tibia 90 percent length of head, or longer; adult size larger, about 50 mm.; 
dewlap red, without a purple spot tropidonotus (p. 60) 

8. Ventrals smooth 3' 9 

Ventrals keeled 25 

9. Supraorbital semicircles in contact and a single row of only 4 or fewer large 

supraoculars 10 

Supraorbital semicircles separated or, if in contact, supraoculars numerous (8 
or more enlarged) 14 

10. Two rows of loreals; female with a large gular "pouch"; all scales on body 

flat and granular, small and equal on back and sides, larger on belly. 

impetigosus (p. 60) 

Four to six rows of loreals; females (not known in schmidti) lacking a large 

gular pouch; dorsal scales keeled, at least faintly larger than lateral scales. 

11 

11. Ventrals and dorsals subequal in size; enlarged supraoculars three-fourths 

width of supraocular area schmidti (p. 60) 

Ventrals distinctly larger than (at least 1% times as large as) dorsals; enlarged 
supraoculars narrower, half width of supraocular area or less 12 

12. Two gulars in contact with mental between chin shields; 9 or fewer scales 

across snout between second canthals (counting as the first canthal that 

which borders the superciliaries) 13 

Three or (generally) 4 gulars in contact with mental between chin shields; 
10 to 12 scales across snout between second canthals gadovii (p. 61) 

13. Six scales between nasals; 5 scales in contact with rostral between supralabials; 

ventrals larger dunni (p. 61) 

'« In six species (cymbops, laeriventris, limifrons rodriguezii, milleri, schiedii, and schmidti) the ventrals may 
appear either smooth or keeled; these are keyed out under both categories. All others are uniformly and 
plainly either smooth or keeled. 



CHECKLIST OF REPTILES OF ME'XICO 57 

Seven or 8 scales between nasals; 6 scales in contact with rostral between 
supralabials; ventrals smaller taylcri (p. 61) 

14. All dorsals and laterals equal in size, minute, granular; tibia 85 percent length 

of head (to anterior border of ear); scales of supraorbital semicircles en- 
larged; head scales feebly keeled damulus (p. 61) 

At least 2 or more vertebral rows of slightly enlarged scales, or, if all dorsal 
scales uniform, tibia less than 70 percent length of head, and either scales of 
supraorbital semicircles not enlarged, or head scales perfectly smooth-. 15 

15. Scales of supraorbital semicircles, at least in the frontal and frontoparietal 

area, not or scarcely larger than scales between; tail compressed, sub- 
triangular in section with a serrate crest; tibia much shorter than head 
(70 percent or less) all dorsal and lateral scales equal in size. 

pentaprion (p. 61) 

Scales of supraorbital semicircles distinctly larger than scales between; tail not 

or onh' shghtly compressed, with httle or no crest 16 

16. Supraorbital semicircles either in contact or separated by a single series of 

scales; postanals enlarged in males (males not known in utoivanae, cymho-ps, 

beckeri, and baccatus) 17 

Supraorbital semicircles separated by 2 or 3 rows of scales; postanals not 
enlarged in males (not known in guntherii) 22 

17. Interparietal smaller than ear opening 18 

Interparietal larger than ear opening 19 

18. Three loreal rows utowanae (p. 62) 

Seven loreal rows cymbops (p. 62) 

19. All dorsal and lateral scales equal in size, minute, granular, smooth; tibia only 

60 to 65 percent head length; all dorsal head scales smooth. 

beckeri (p. 62) 
At least a few vertebral scales slightly enlarged, keeled; tibia longer in propor- 
tion to head length; dorsal head scales smooth or not 20 

20. Upper head scales tricarinate baccatus (p. 62) 

Upper head scales smooth, shghtly rugose, or feebly unicarinate; postanals 

enlarged in males 21 

21. One series of 3 enlarged supraoculars; male lacking first digit on all four feet*^; 

dewlap red liogaster (p. 62) 

Two series of 7 to 9 enlarged supraoculars; males with 5 digits on each foot; 
dewlap yellow laeviventris (p. 62) 

22. Tibia distinctly longer than head (15.5 mm. and 13.5 mm. respectively in the 

type and only known specimen) schiedii (p. 63) 

Head (snout to anterior border of ear opening) at least a little longer than 
tibia 23 

23. Interparietal plate twice as long as ear opening, much larger than adjacent, 

very small head scales guntherii (p. 63) 

Interparietal plate not more than !){ times as long as ear opening, not more 
than 4 times as large as adjacent scales 24 

24. Ventral scales only slightly larger than dorsals; one large scale in anterolateral 

corner of supraocular area, in contact with superciliaries and separated from 

enlarged supraoculars by one row of small scales milleri (p. 64) 

Ventral scales much larger than dorsals limifrons rodriguezii (p. 64) 

25. Head shorter than tibia 26 

Head longer than tibia 27 



" Materia Ireeently examined demonstrates that this species like others of Anolis possesses five digits on 
both limbs in both sexes. 



58 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

26. Ventrals very strongly keeled; dorsal scales smooth, no larger than laterals; 

head broad capito (p. 65) 

Ventrals very weakly keeled or smooth; dorsal scales keeled, somewhat larger 
than laterals; head longer schiedii (p. 63) 

27. Lamellae under second and third phalanges of fourth toe, about 27; inter- 

parietal scarcely enlarged, much smaller than ear opening petersii (p. 65) 

Lamellae under second and third phalanges of fourth toe, 23 or less; inter- 
parietal variable 28 

28. Two or 3 rows of scales between supraorbital semicircles 29 

Supraorbital semicircles in contact or separated from each other by no more 

than 1 row of scales 30 

29. Ventrals very strongly keeled; 22 or 23 lamellae under phalanges 2 and 3 of 

fourth toe biporcatus (p. 65) 

Ventrals very weakly keeled; not over 16 or 17 lamellae 24 

30. A single row of not more than 5 large supraoculars 31 

Two or more rows of 7 or more enlarged supraoculars 35 

31. Ventrals very weakly keeled; postanals enlarged and dewlap yellow in males; 

supraorbital semicircles broadly in contact schmidti (p. 60) 

Ventrals very strongly keeled 32 

32. Males with enlarged postanals and dewlap more or less unicolor 33 

Males without enlarged postanals and dewlap with a conspicuous central 

dark blue spot 34 

33. Dorsal scales very slightly smaller than ventrals nebulosus (p. 65) 

Dorsal scales markedly smaller than ventrals nebuloides (p. 66) 

34. Supraorbital semicircles in contact with each other and with interparietal. 

kidderi (p. 66) 
Supraorbital semicircles separated from each other (generally) and from inter- 
parietal (invariably) ustus (p. 66) 

35. Interparietal little if any larger than ear opening; dewlap in males without a 

central blue spot 36 

Interparietal nearly or fully twice as long as ear opening; dewlap in males with 
a central blue spot or not 38 

36. Ventral scales indistinctly keeled; dorsal scales granular except two vertebral 

rows of abruptly enlarged scales cymbops (p. 62) 

Ventral scales strongly keeled; dorsal scales uniform in size in both vertebral 
and paravertebral areas 37 

37. Dorsals very small, much smaller than ventrals; 17-19 lamellae under pha- 

langes 2 and 3 of fourth toe; 6-8 rows of loreals- -lemurinus bourgeaei (p. 66) 

Dorsals rather large, distinctly smaller than ventrals; 15 or 16 lamellae; 4-6 

rows of loreals cumingii (p. 67) 

38. Dorsals very slightly smaller than ventrals, in about 14 rows; a purple spot in 

center of gular fan in males; postanals not enlarged heliactin (p. 67) 

Dorsals smaller; a purple spot or not; postanals enlarged or not "' 39 

39. Both dorsals and ventrals smaller; no purple spot in center of dewlap; post- 

anals enlarged in males laeviventris (p. 62) 

Dorsals and ventrals larger; a purple spot in center of dewlap; postanals not 
enlarged in males sericeus (p. 67) 

ANOLIS BARKERI Schmidt 

Anolis barkeri Schmidt, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, 
pp. 7-9, fig. 2. 

Ttjpe.— Brit. Mus. Nat. Hist. No. 36.6.6.12; R. Wright Barker 
collector. 



CHECKLIST OF REPTILES OF MEXICO 59 

Type locality. — Cascajal, upper Uzpanapa River, Veracruz. 
Range. — Known only from the type locality. 

ANGUS SAGREI MAYENSIS Smith and Burger 

Anolis sagrei mayensis Smith and Burger, Anal. Inst. Biol., vol. 20, 1949, p. 407. 

Type.—E. H. Taylor No. 11523; H. M. Smith collector. 

Type locality. — Panlao, Campeche (a small island at mouth of Rio 
Mamantel, Laguna de Terminos). 

Range. — The Atlantic coast from central Mexico to northern South 
America. Recorded in Mexico from the states of Yucatan: Progreso, 
Merida, Chichen Itzd; Campeche: Balchacaj, Ciudad del Carmen, 
Panlao; Tabasco: San Juan Bautista; Quintana Rao: Cozumel Island. 

ANGUS CGZUMELAE Smith 

Anolis cozumelae Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, 
pp. 19-20, fig. 4 (at left). 

!r?/pe.— Chicago Nat. Hist. Mus. No. 751; male; C. F. Mdlspaugh 
collector. 

Type locality. — Cozumel Island, Yucatan Peninsula. 
Range. — Cozumel Island, Quintana Roo. 

ANGUS METALUCUS Bocourt » 

Anolis metallicus Bocourt, Ann. Sci. Nat., ser. 5, vol. 17, art. 2, 1873, p. 1; 

Mission scientifique au Mexique . . ., fitudes sur les reptiles, livr. 2, 1873, 

pi. 17 bis, fig. 1. — BouLENGER, Catalogue of the lizards in the British Museum, 

vol. 2, 1885, p. 84. 
Anolis uniformis, Barbour (part), Bull. Mus. Comp. Zool., vol. 77, No. 4, 1934, 

p. 153. 
Anolis tropidonotus, Stuart (part), Occ. Pap. Mus. Zool. Univ. Michigan, No. 464, 

1942, p. 2. 

Type. — Mus. Hist. Nat. Paris. 
Type locality. — Mexico, 
Range . — Unknown . 

ANGUS MEGAPHGUDGTUS Smith 

Anolis megapholidotus Smith, Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 318-320. 

r^/pe.— EHT-HMS No. 11149 (originally EHT-HMS field No. 
1509); E. H. Taylor and H. M. Smith collectors. 

Type locality. — Between Rinc6n and Cajones (about 40-45 km. 
south of Chilpancingo), Guerrero. 

Range. — Known only from the type locality; presumably ranges 
throughout the Sierra Madre del Sur in Guerrero. 

•9 Barbour (loc. cit.) synonymizes this with A. h. uniformis, and Stuart {loc. cit.) suggests synonymy with 
A. tropidonotus. Inasmuch as both the original description and the subsequent illustration indicate the 
possession by the type of relatively small dorsal scales equal in size to the ventrals, while in both tropidonotus 
and uniformis the dorsals are much larger, we regard metallicus as distinct from either. 

861316—50 5 



60 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

ANOLIS HUMIUS UNIFORMIS Cope 

Anolis unijormis Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, p. 392. — 
BouLENOER, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
p. 84,— Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 464, 1942, pp. 1-2. 

Anolis humilis unijormis, Stuart, Misc. Publ. Mus. Zool. Univ. Michigan, No. 
69, 1948, p. 48. 

Anolis ruthveni Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 310, 1935, 
pp. 1-4 (type, Univ. Michigan Mus. Zool. No. 76622, 2 miles north of Santa 
Teresa, El Pet6n, Guatemala). 

Type.—U.S.'N.M., Nos. 6774, 24734-48, 24750 (Guatemala), 24859 
(Yucatan), cotypes; Mus. Comp. Zool. No. 10933 (Guatemala). 

Type locality. — Yucatan and Guatemala, here restricted to 2 miles 
north of Santa Teresa, Guatemala. 

Range. — The southern portion of the Yucatan Peninsula, from 
Chiapas and probably Tabasco to El Peten and British Honduras. 
Known in Mexico from Chiapas: Usumacinta River (across from 
Piedras Negras, El Peten, Guatemala), Tlacotalpa, Mineral de Santa 
Fe; Tabasco: Teapa. 

ANGUS TROPIDONOTUS Peters 

Anolis tropidonotus Peters, Monatsb. Akad. Wiss. Berlin, 1863, p. 135. — Bocourt, 
Mission scientifique au Mexique . . ., fitudes sur les reptiles, livr. 2, 1873, 
pp. 103-105, pi. 16, fig. 30. — BouLENGER, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, pp. 83-84. 

Norops tropidonotus O'Shaughnessy, Ann. Mag. Nat. Hist., ser. 4, vol. 4, 1869, 
p. 273. 

Norops yucatanicus Barbour and Cole, Bull. Mus. Comp. Zool., vol. 50, 1906, 
p. 149 (type, Mus. Comp. Zool. No. 7036, Chichen Itzd, Yucatdn). 

Type. — Zool. Mus. Berlin, 6 specimens; Dr. Hille collector. 

Type locality. — Huanuco {sic) = probably, Huatusco, Veracruz. 

Range. — Atlantic slopes from Veracruz to Nicaragua, including the 
Yucatan Peninsula. Recorded from the states of Veracruz, Tabasco, 
Oaxaca, Campeche, Yucatan, and Quintana Roo (Cobd). 

ANGUS IMPETIGOSUS Cope 

Anolis impetigosus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 174. — 
Boulenger, Catalogue of the lizards in the British Museum . . .. ed. 2, 
1885, p. 55, pi. 2, fig. 3. 

Type.— Brit. Mus. Nat. Hist. 
Type locality. — Unknown. 
Range . — Unknown . 

ANGUS SCHMIDTI Smith 

Anolis schmidti Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, No. 4, 
1939, pp. 21-23, fig. 4 (at right). 

Type.— Chicago Nat. Hist. Mus. No. 1667; C. H. T. Townsend 
collector. 

Type locality. — Manzanillo, Colima. 

Range. — Kjiown only from the type locality. 



CHECKLIST OF REPTILES OF MEXICO 61 

ANGUS GADOVn Boulenger 

Anolis gadovii Boulenger, Proc. Zool. Soc. London, 1905, vol. 2, p. 245, pL 6, 
fig. 1. — MosAUER, Herpetologica, vol. 1, no. 2, 1936, pp. 61-63. — Smith and 
Spieler, Copeia, 1945, pp. 165-168. 

Type. — Brit. Mus. Nat. Hist.; Hans Gadow collector. 
Tyye locality. — Tierra Colorada, Guerrero. 
Range. — Known only from the type locality. 

ANOLIS DUNNI Smith 

Anolis gadovii, Smith (part), Trans. Kansas Acad. Sci., vol. 36, 1933, pp. 316-318. 
Anolis dunni Smith, Copeia, 1936, p. 9. — Smith and Spieler, Copeia, 1945, pp. 
165-168. 

Ty^e.— EHT-HMS No. 1506; E. H. Taylor coUector. 
Type locality. — Between Rinc6n and Cajones, Guerrero (a place 
now called Agua del Obispo). 

Range. — Known only from the type locality. 

ANGUS TAYLGRI Smith and Spieler 

Anolis laylori Smith and Spieler, Copeia, 1945, pp. 165-168. 

ri/2)e.— U.S.N.M. (H. M. Smith field No. 10085); E. H. Taylor 
coUector. 

Type locality. — One mile north of Acapulco, Guerrero. 
Range. — Known only from the type locality. 

ANGUS DAMULUS Cope 

Anolis damulus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 169. — Bou- 
lenger, Catalogue of the lizards in the British Museum . . ., ed. 2, vol. 2, 
1885, pp. 47-48, pi. 2, fig. 2. 

Type. — Brit. Mus. Nat. Hist. 
Type locality. — Unknown. 
Range. — Unknown. 

ANGUS PENTAPRION Cope 

Anolis (Coccoessus) pentaprion Cope, Proc. Acad. Nat. Sci. Philadelphia, 1862, 

p. 178. 
Anolis pentaprion, Boulenger, Catalogue of the lizards in the British Museum, 

ed. 2, vol. 2, 1885, p. 64. 
Anolis sulcifrons Cope, Sci. Bull. Philadelphia Mus., No. 1, 1899, p. 6, pi. 2, fig. 1 

(type, Amer. Mus. Nat. Hist. No. 38750; Baranquilla, Colombia). 
Anolis panamensis Boulenger, Proc. Zool. Soc. London, 1890, p. 81, pi. 13, fig. 2 

(type, Brit. Mus. Nat. Hist. No. 89-17-2-31, 2 cotypes; Panama). 

Type. — Formerly in U.S.N.M., now lost; A. Schott coUector. 
Type locality. — Truando River, Colombia. 

Range. — Atlantic slopes from the Isthmus of Tehuantepec to 
Colombia. KJiown in Mexico only from Chiapas: Palenque. 



62 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

ANOLIS UTOWANAE Barbour 

AnoUs utowanae Barbour, Copeia, 1932, p. 11. 

Type. — Mus. Comp. Zool. No. 31035; Thomas Barbour collector 
(purchased from Indian) . 

Type locality. — Ten miles north of Mazatlan, Sinaloa. 
Range. — Coastal Sinaloa, known only from the type locality. 

ANOLIS CYMBOPS Cope 

Anolis cymhops Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 173-174. — 
BouLENGER, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
pp. 73-75. 

Type.— Brit. Mus. Nat. Hist. 

Type locality. — ^"Veracruz," Mexico. 

Range. — Known only from the type locality. 

ANOLIS BECEERI Boulenger 

Anolis beckeri Boulenger, Proc. Zool. Soc. London, 1881, pp. 921-922; Cata- 
logue of the lizards in the British Museum, vol. 2, 18S5, pp. 46-47. 

Type. — Royal Belg. Mus., two co types; A. Boucard, collector. 
Type locality. — Yucatan. 

Range.— Yuc&tkn. Peninsula, including British Honduras. Re- 
corded in Mexico only from Yucatan: Chichen Itza. 

ANOUS BACCATUS Bocourt 

Anolis baccatus Bocourt, Mission scientifique au Mexique . . ., Etudes sur les 
reptiles, livr. 2, 1873, pp. 59-61, pi. 14, fig. 14. — Boulenger, Catalogue of 
the lizards in the British Museum, vol. 2, 1885, p. 54. 

Type. — Mus. Hist. Nat. Paris; Auguste Salle collector. 

Type locality. — Mexico. 

Range. — Uncertain. The specimen reported from Sepaquite, Alta 
Verapaz, Guatemala (Barbour, Bull. Mus. Comp. Zool., vol. 77, 1934, 
p. 124), is actually A. sericeus {fide L. C. Stuart, in litt.). No definite 
Mexican records are known. 

ANOUS LIOGASTER Boulenger 

Anolis liogaster Boulenger, Proc. Zool. Soc. London, 1905, vol. 2, pp. 245-246, 
pi. 6, fig. 2. 

Type. — Brit. Mus. Nat. Hist., two cotypes; Hans Gadow collector. 
Type locality. — Omilteme, Guerrero (7,600 feet). 
Range. — Known only from the type locality.^® 

ANOLIS LAEVIVENTRIS (Wiegmann) 

D[actyloa] {A[nolis]) laeviventris Wiegmann, Herpetologia Mexicana, 1834, p. 47. 



" Gadow (Proc. Zool. Soc. London, 1905, pp. 195, 216) suggests also the locality Nevado de Collma, Jalisco. 
The record is undoubtedly unacceptable. 



CHECKLIST OF REPTILES OF MEXICO 63 

Anolis laeviventris, Bocotjrt, Mission scientifique au Mexique . . ., fitudes sur 
les reptiles, livr. 2, 1873, pp. 87-88; livr. 3, 1874, pi. 16, fig. 18, 18a.— Bou- 
IjEnger, Catalogue of the lizards in the British Museum, vol. 2, 1885, p. 76. — 
Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, pp. 20-21. 

Anolis wiegmanni Fitzinger, Systema reptilium, 1843, p. 67 (substitute name for 
Dactyloa laeviventris Wiegmann). 

Anolis nannodes Cope (part only; a lectoparatype) , Proc. Acad. Nat. Sci. Phila- 
delphia, vol. 16, 1864, p. 173 (lectotype, U.S.N.M. No. 12200, Arriba, Costa 
Rica)." 

Anolis intermedins, Boulenger, Catalogue of the lizards in the British Museum, 
ed. 2, vol. 2, 1885, pp. 78-79 (part). 

Type. — Zool. Mus. BerL; F. Deppe collector. 

Type locality. — Mexico, here restricted to Jalapa, Veracruz. 

Range. — Atlantic slopes from Central Veracruz to the Isthmus of 
Tehuantepec. Recorded from the states of Veracruz: Jalapa, 
Orizaba, Mirador, Actopam, Jicaltepec, Xico; and Tabasco. '^^ 

ANOLIS SCHIEDII (Wiegmann) 

D[actyloa] schiedii Wiegmann, Herpetologia Mexicana, 1834, p. 48. 

Anolis schiedii, Lichtenstein and Martens, Nomenclator reptilium et amphib- 
iorum Musei Zoologici Berolinensis, 1856, p. 8. — Bocourt, Mission scienti- 
fique au Mexique . . ., Etudes sur les reptiles, livr. 2, 1873, pp. 64-65 
(part)^^ pi. 15, fig. 19 (of type). — ?Boulenger, Catalogue of the lizards in 
the British Museum, vol. 2, 1885, p. 52. 

Type. — Zool. Mus. BerL; Schiede collector. 
Type locality. — Mexico (by inference). 
Range . — Unknown .*^ 

ANGUS GUNTHERII Bocourt 

Anolis guntherii Bocourt, Mission scientifique au Mexique . . ., fitudes sur les 
reptiles, livr. 2, 1873, pp. 61-62, pi. 15, fig. 15. 

" The original type series included two, a male and a female, in the Brit. Mus. Nat. Hist., from CobSn, 
Guatemala; one, U.S.N.M. No. 12206, from Arriba, Costa Rica, and one, U.S.N.M. No. 6117, now lost, from 
Jalapa, Veracruz. Dunn (Proc. New England Zool. Club, vol. 12, 1930, p. 18), in discussing A. irUermedius 
of Central America, mentions "the type of nannodes, U.S.N.M. No. 12206. Costa Rica," stating that it is the 
same as intermedins. We regard this action as constituting a restriction of the name to mtermedius and 
rendering U.S.N.M. No. 12206 the lectotype of nannodes. The Mexican specimen obviously represents 
laeviventris, while the Guatemalan species represented by the Coban specimens remains uimamed. Stuart 
(Misc. Publ. Mus. Zool. Univ. Michigan, No. 69, 1948, p. 50) restricts the name nannodes to the Guatemalan 
species, inasmuch as a name is available for each of the other two species (now laeviventris of Mexico, inter- 
medins of Costa Rica) represented in the cotypes. This procedure certainly is the most desirable, but in 
view of the fact that Dunn previously restricted the name to the Costa Rican species, Stuart's course can- 
not, apparently, be maintained. The name Anolis sluarti Smith , new name, is proposed for the Guatemalan 
species represented by the two cotypes of nannodes in the British Museum of Natural History, from Cobta, 
Alta Verapaz. For a description of the types, see Boulenger, Cat. Liz. Brit. Mus., vol. 2, 1885, p. 78. 

The Anolis nannodes (auct.) from numerous localities in Mexico is the species laeviventris. — H. M. S. 

<' In various papers (first in Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 123) Cope records this species 
from Yucatan and from "Tehuantepec"; Duges (La Naturaleza, ser. 2, vol. 2, 1896, p. 479) records it from 
Taugancicuaro (Michoacan), Guadalajara (Jalisco), and Cerro de los Amoles (state?); Mocquard (Bull. 
Soc. Philom. Paris, ser. 9, vol. 1, 1899, pp. 155-156) records it from Guadalajara, Jalisco; and Peters (Monatsb. 
Berl. Acad., 1869, p. 874) records it from Matamoros, Puebla. We believe all these records refer to other 
species with smooth ventrals. We are unable at this time to place them accurately. 

" Bocourt possessed for his description the type of schiedii and a series of specimens from CobSn, Guate- 
mala; all the latter are, however, apparently referable to A. cobanensis Stuart. 

<' Cope's record (U. S. Nat. Mus. Bull. 32, 1887, p. 31) for Jalapa, Veracruz, is highly questionable. 



64 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Anolis guentherii, Boulenger, Catalogue of the lizards in the British Museum, 
vol. 2, 1885, pp. 54-55. 

Type. — Mus. Milano. 
Type locality. — Mexico. 
Range. — Unknown . 

ANGUS MILLERI Smith, new species « 

Anolis schiedii, Smith and Laufe (nee Wiegmann), Herpetologica, vol. 3, 1945, 
pp. 2-4. 

Type.— v. S.'N.M. No. 120957; Walter S. Miller collector. 
Type locality. — Quetzaltepec, Oaxaca. 
Range. — Known only from the type locality. 

ANOLIS UMIFRONS RODRIGUEZn Bocourt 

Anolis rodriguezii Bocourt, Mission scientifique au Mexique . . . , Etudes sur 

les reptiles, livr. 2, 1873, pp. 62-63, pi. 13, fig. 1. 
Anolis limifrons rodriguezii,*^ Stuart, Misc. Publ. Mus. Zool. Univ. Michigan, 

No. 69, 1948, pp. 49-50. 
Anolis rubigenosus Bocourt, Ann. Sci. Nat., ser. 5, art. 2, 1873, p. 1; Mission 

scientifique au Mexique . . . , Etudes sur les reptiles, livr. 3, 1874, pi. 17 bis, 

fig. 2 (type, Mus. Hist. Nat. Paris; Oaxaca [state])." 
Anolis aureolus Cope, Proc. Amer. Philos. Soc, 1885, pp. 390-391 (Guatemala, 

2 cotypes, U.S.N.M. Nos. 24850-1; Yucatdn, 3 cotypes U.S.N.M. Nos. 25854- 

5, 25857, one cotype Mus. Comp. Zool. No. 10929, type locality here restricted 

to Chichen Itzd, Yucatdn). 
Anolis acutirostris Ives, Proc. Acad. Nat. Sci. Philadelphia, 1891, pp. 459-460 

(type, Acad. Nat. Sci. Philadelphia No. 7889; Citilpech, Yucatdn). 

Type. — Mus. Hist. Nat. Paris; collected by Commission Scientifique 
du Mexique. 

Type locality. — Panz6s, Alta Verapaz, Guatemala. 

Range. — Atlantic slopes from the Isthmus of Tehuantepec to 
Honduras, including the peninsula of Yucatan except its semiarid 
northwestern coast. Recorded in Mexico from Oaxaca (no locality); 
Chiapas: Palenque; Campeche: Balchacaj, Champot6n, Tres Brazos, 
Encarnacion; Yucatan: Citilpech, Chichen Itza, Merida; and Quintana 
Roo: Cozumel Island. 

** Diagnosis.— As in the accompanying key (p. 57). Differs from schiedii in two important respects: The 
head length from snout to ear (15.8 mm.) is distinctly greater than tibia length (13.6 mm.), and the inter- 
parietal is much larger, its length equaling the diameter of ear opening (the tibia, 15.5 mm., is longer than 
head, 13.5 mm., and length of the interparietal is little more than half the greatest diameter of the ear open- 
ing, in schiedii). A. schiedii is unique, in its group of the genus, in its very long tibia. 

Description.— A lengthy description of the type was published in Smith and Laufe (lac. cit.) and needs no 
repetition.— H. M. S. 

*» Spelled ridriguezii by error. 

*• The type, according to Bocourt's figure and description, does not differ from the form most authors in the 
past have called aureolus, except perhaps in size of the interparietal. It appears probable to us that the 
deceptive marking on the type of rodriguezii (a broad vertebral light stripe) , which Bocourt apparently 
thought was constant, may have been responsible for his failure to consider rodriguezii and the type of his 
rubigenosus as conspeciflc. 

Although no other record indicates the occurrence of rodriguezii in Oaxaca, the known distribution else- 
where strongly implies occurrence on Atlantic slopes of that state near or at the Isthmus of Tehuantepec. 



CHECKLIST OF REPTILES OF MEXICO 65 

ANGUS CAPITO Peters 

Anolis (Draconura) capita Peters, Monatsb. Acad. Wiss. Berlin, 1863, p. 142. 
Anolis capita, BocouRT, Mission scientifique au Mexique . . . , Etudes sur les 

reptiles, livr. 2, 1873, pi. 16, fig. 27. — Boulenger, Catalogue of the lizards 

in the British Museum, vol. 2, 1885, pp. 94-95. 
Anolis carneus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 169 (Brit. Mus. 

Nat. Hist., two cotypes; lower Verapaz forest). 

Type. — Zool. Alus. Ber. No. 4086, two cotypes; C. Hoffman collector. 
Type locality. — Costa Rica, here restricted to Palmar. 
Range. — Tabasco to Panama. Recorded in Mexico only from 
"Tabasco." 

ANOUS PETERSn Boconrt 

Anolis petersii Bocourt, Mission scientifique au Mexique . . . , fitudes sur les 
reptiles, livr. 2, 1873, pp. 79-80, pi. 13, fig. 2, and pi. 15, figs. 11, 11a.— 
Boulenger, Catalogue of the lizards in the British Museum, vol. 2, 1885, pp. 
66-67. 

Anolis petersii bivittatus Werner, Verh. zool.-bot. Ges. Wien, vol. 46, 1898, p. 351 
(Munich Mus., two cotypes; Guatemala, here restricted to Cobdn). 

Type. — Mus. Hist. Nat. Paris No. 1641 /3, two specimens; collected 
by Commission Scientifique du Mexique. 

T7jpe locality. — Alta Verapaz, Guatemala, here restricted to Finca 
Samac. 

Range. — Atlantic slopes from central Veracruz to Alta Verapaz in 
Guatemala. Recorded in Mexico only from Veracruz: Cuautlapan, 
Mirador, Orizaba; San Luis Potosi: Xilitla. 

ANOLIS BIPORCATUS (Wiegmann) 

D.[actyloa] biporcata Wiegmann, Herpetologia Mexicana, 1834, p. 47. 

Anolis biporcatus, Schmidt, Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, 1941, 
p. 491.— Smith, Rev. Soc. Mex. Hist. Nat., vol. 7, 1946, pp. 66-67.— Stuart, 
Misc. Publ. Mus. Zool. Univ. Michigan, No. 69, 1948, pp. 46-47. 

Anolis copei Bocourt, Mission scientifique au Mexique . . . , Etudes sur les 
reptiles, livr. 2, 1873, pp. 77-78, pi. 15, fig. 10 (Panzos, Alta Verapaz, Guate- 
mala; Mus. Hist. Nat. Paris). — Boulenger, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, pp. 65-66. 

Type. — Zool. Mus. Berlin; F. Deppe collector. 
Type locality. — Mexico, here restricted to Piedra Parada, Chiapas. 
Range. — Chiapas to Panama. Known in Mexico only from Chiapas 
(Piedra Parada). 

ANOLIS NEBULOSUS (Wiegmann) 

D[actyloa] nehulosa Wiegmann, Herpetologia Mexicana, 1834, p. 47. 

Anolis nebulosus, Bocourt, Mission scientifique au Mexique . . . , Etudes sur les 

reptiles, livr. 2, 1873, pp. 68-69. — Boulenger, Catalogue of the lizards in 

the British Museum, vol. 2, 1885, pp. 76-77. 
Anolis boulengerianus Thominot, Bull. Soc. Philom., ser. 7, vol. 11, 1887 (Mus. 

Hist. Nat. Paris, three cotypes, Isthmus of Tehuantepec, F. Sumichrast 

collector). 

Type. — Zool. Mus. Berlin, two cotypes; F. Deppe collector. 



66 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type locality. — Mexico (by inference), here restricted to Mazatlan, 
Sinaloa. 

Range. — Western Mexico, from Sinaloa to the Isthmus of Te- 
huantepec. Known from the states of Morelos, Puebla, Nayarit, 
CoUma, Oaxaca, Guerrero, Michoacan, Sinaloa, Jalisco, Durango, 
and the Tres Marias Islands.*^ 

ANGUS NEBULOIDES Bocoort 

Anolis nebuloides Bocourt, Mission scientifique au Mexique . . ., fitudes sur les 
reptiles, livr. 2, 1873, pp. 74-75, pi. 13, fig. 10. — Botjlenger, Catalogue of 
the lizards of the British Museum, vol. 2, 1885, p. 77. 

Type. — Mus. Hist. Nat. Paris, four co types; collected by (?) Com- 
mission Scientifique du Mexique. 

Type locality. — Putla, Oaxaca. 

Range. — Western Mexico from Chihuahua to Oaxaca, Recorded 
from the states of Chihuahua, Sonora, Colima, Guerrero, Puebla, 
and Oaxaca .^^ 

ANGUS KIDDERI Ruthven 

Anolis kidderi Ruthven, Occ. Pap. Mus. Zool. Univ. Michigan, No. 257, 1933, 
pp. 1-2. 

Ty^e. — Mus. Zool. Univ. Michigan No. 72851; Edwin P. Creaser 
collector. 

Type locality. — Merida, Yucatan. 

Range. — The western part of the Yucatan Peninsula. Recorded 
from Yucatan and Campeche. 

ANGUS USTUS Cope 

Anolis ustus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 172. — Boulengbr, 
Catalogue of the lizards in the British Museum, vol. 2, 1885, p. 73. — Schmidt, 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, No. 8, 1941, p. 493. — Stuart, 
Misc. Publ. Mus. Zool. Univ. Michigan, No. 69, 1948, pp. 49, 51. 

Type. — Brit. Mus. Nat. Hist., two cotypes. 
Type locality. — Belize, British Honduras. 

Range. — The Yucatan Peninsula and the area at its base. Recorded 
in Mexico from Yucatan and Campeche. 

ANGUS LEMURINUS BGURGEAEI Bocourt 

Anolis bourgeaei Bocourt, Mission scientifique au Mexique . . ., fitudessurles 
reptiles, livr. 2, 1873, pp. 76-77, pi. 15, fig. 9. — Boulenger, Catalogue of the 
lizards in the British Museum, vol. 2, 1885, pp. 74-75. 

Anolis limifrons bourgeaei, Stuart, Misc. Publ. Univ. Michigan Mus. Zool., 
No. 69, 1948, p. 49. 

*'' Records from the states of Veracruz and Yucat&n are probably erroneous, and that from Chihuahua is 
said to be referable to A. nebuloides. Virtually all records for both species are, for that matter, open to 
question. The two forms have never been adequately characterized, and very likely more than two species 
are involved. 

*' Undoubtedly some records of nebulosm belong here, and perhaps vice versa, A record for Veracruz is 
probably in error. 



CHECKLIST OF REPTILES OF MEXICO 67 

Anolis biporcatus, Bocourt" (nee Wiegiiiann), Mission scientifique au Mexique . . ., 

fitudes sur les reptiles, livr. 2, 1873, pp. 98-99, pi. 15, fig. 8.— Boulenger, 

op. cil., pp. 88-89. 
Anolis usius veraepacis Barbour, Proc. New England Zool. Club, vol. 12, 1932, 

p. 98 (type, Mus. Comp. Zool. Mo. 32324; A. W. Anthony coll.; Hacienda 

Chimoxan, Alta Verapaz, Guatemala).^" 

Type. — Mus. Hist. Nat. Paris, one female (Orizaba, collected by 
Bourgeau); Zool. Mus. Berlin, one male (Huatusco, collector?). 

Type locality. — Huatusco and Orizaba, Veracruz, here restricted to 
Huatusco. 

Range. — Atlantic slopes from Veracruz to Guatemala and British 
Honduras. Recorded in Mexico from the states of Veracruz, Tabasco, 
Campeche, and Yucatan. 

ANGUS CUMINGn Peters 

Anolis cumingii Peters, Monatsb. Akad. Wiss. Berlin, 1863, p. 140. — Bocourt, 
Mission scientifique au Mexique . . . , Etudes sur les reptiles, livr. 2, 1873, 
p. 89, pi. 16, fig. 20. — BotJLENGER, Catalogue of the lizards in the British 
Museum, vol. 2, 1885, p. 80. — Burt and Myers, Stanford Univ. Publ., 
biol. sci., vol. 8, 1942, p. 291.— Smith, Anal. Inst. Biol. Mexico, vol. 14, 1942, 
p. 342. 

Type. — Zool. Mus. Berlin; Herr Cuming collector. 
Type locality. — Mexico. 

Range. — Uncertain; possibly the upper Balsas Basin. Recorded 
only from La Paz (Puebla?). 

ANGUS HEUACTIN Cope 

Anolis heliadin Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 172-173. — 
Bocourt, Mission scientifique au Mexique . . . , Etudes sur les reptiles, 
livr, 2, 1873, pp. 106-108, pi. 13, figs. 4, 4a-c (in color), pi. 16, fig. 32.— 
Boulenger, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
p. 72. 

Type.— Ac&d. Nat. Sci. Philadelphia No. 7914. 

Type locality. — Mexico, 

Range. — Recorded only from "Oaxaca." 

ANGUS SERICEUS Hallowell 

Anolis sericeus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 8, 1856, 
pp. 227-228.— Barbour, BuU. Mus. Comp. Zool., vol. 77, 1934, pp. 149-150.— 
Schmidt, Publ. Field Mus. Nat. Hist., zool. ser., vol. 22, 1941, p. 492. 

Anolis sallaei GtJNTHER, Proc. Zool. Soc. London, 1859, p. 421 (Brit. Mus. Nat. 
Hist.; Mexico; Salle coll.). — Bocourt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 2, 1873, p. 90, pi. 13, fig. 3, pi. 16, fig. 21.— 
Boulenger, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
pp. 79-80. 

" And of other authors prior to Schmidt, 1941; see synonymy of A. biporcatus. 

»» According to Stuart (op. cit., 1948, p. 51) the type and several of the paratypes oiA. u. veraepacis Barbour 
are referable to A. bourgeaei while the remaining paratypes are A. sericeus. 



68 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Anolis jacobi Bocotirt, op. cit., p. 74, pi. 13, fig. 8 (Mus. Hist. Nat. Paris; Vera- 
cruz, here restricted to Veracruz). 

Type. — Acad. Nat. Sci. Philadelphia, now lost; Mr. Pease donor. 

Type locality. — El Encero de Jalapa, Veracruz. 

Range, — Atlantic slopes from Tamaulipas and Pacific slopes from 
the Isthmus of Tehuantepec south to Nicaragua. Recorded in Mexico 
from the states of Tamaulipas, San Luis Potosi, Veracruz, Oaxaca. 
Tabasco, and Chiapas. 

Genus CORYTHOPHANES Boie 

Coryto-phanes [sic] BoiE, in Schlegel, Isis von Oken, vol. 20, pt. 3, 1826, p. 290. 
Chamaeleopsis Wiegmann, in Gray, in Griffith, The animal kingdom ... by 

the Baron Cuvier . . . , vol. 9, 1831, Synopsis, p. 45 (type, Chamaeleopsis 

hernandezii Wiegmann) .^i 

Genotype. — Agama cristata Merrem. 
Species. — Three. 

Range. — Central Veracruz and the Pacific coast of Chiapas south- 
eastward to Costa Rica. 

KEY TO SPECIES OF CORYTHOPHANES 

1. Nuchal crest interrupted, not continuous with dorsal crest. hernandezii (p. 68) 
Nuchal crest continuous with dorsal, although sometimes low where the two 

meet 2 

2. Parietal crests joining very near extremity of helmet; upper head scales keeled 

or rugose percarinatus (p. 69) 

Parietal crests joining about halfway from orbits to extremity of helmet; upper 
head scales smooth cristatus (p. 69) 

CORYTHOPHANES HERNANDEZU (Wiegmann) 

Chamaeleopsis hernandezii Wiegmann, in Gray, in Griffith, The animal kingdom 
. . ., by the Baron Cuvier . . ., vol. 9, 1831, Synopsis, p. 45; Isis von Oken, 
1831, p. 298; Herpetologia Mexicana, pt. 1, 1834, pp. 15, 37, pi. 6. 

Corythophanes hernandezii, Lichtenstein and Martens, Nomenclator reptilium 
et amphibiorum Musei Zoologici Berolinensis, 1856, p. 8. — Boulenger, 
Catalogue of the lizards in the British Museum, vol. 2, 1885, pp. 103-104. 

Corythophanes chamaeleopsis Dumeril and Bibron, Erpetologie generale, vol. 4, 
1837, p. 175-177 (substitute name for Chamaeleopsis hernandezii Gray). 

Corythophanes mexicanus Bocourt, Mission scientifique au Mexique . . ., Etudes 
sur les reptiles, livr. 3, 1874, pp. 122-123, pi. 17, fig. 1 (substitute name for 
Chamaeleopsis hernandezii Wiegmann, adopting Hernandez's pre-Linnaean 
name Chamaeleo Mexicanus) .^^ 

" The author of this generic name, and of the specific name as well, has been accepted as Gray by some 
authors, as Wiegmann by others. Wiegmann pubhshed his own description in 1831, in great detail; Gray 
published but a few words, also in 1831, but attributed both names to "Wiedemann, MSS" (in error; = 
Wiegmann). Even though Gray's description may have preceded Wiegmann's, it seems obvious that 
Wiegmann was ethically the author, having provided the only reasonable description, and his manuscript 
having been acknowledged by Gray as the source of the latter's information. It is true that Gray stated 
specimens were in the British Museum (Boulenger cites one purchased by Gray), so that Gray could well 
have drawn up his description from them and not from Wiegmann's manuscript. The correct authorship 
thus remains debatable. 

M Wiegmann cited Chamaeleo Mexicanus of HernSndez in both of his papers (1831 and 1834) but did not 
adopt the name. 



CHECKLIST OF REPTILES OF MEXICO 69 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, here restricted to Jalapa, Veracruz. 

Range. — Atlantic slopes from central Veracruz south and east to 
Guatemala. Recorded in Mexico from various localities in the states 
of Veracruz, San Luis Potosi (Xilitla), Oaxaca, Chiapas, Yucatan, 
and Quintana Roo (Coba). 

CORYTHOPHANES CRISTATUS (Merrem) 

Agama crislata Merrem, Tentamen systematis amphibiorura, 1821, p. 50. 

Corythophanes cristatus,^^ Gravenhorst, Acta Acad. Leop. -Carol., vol. 16, 1833, 
p. 938, pi. 65, figs. 6-10.— Dum]6ril, Arch. Mus. Hist. Nat. Paris, vol. 8, 
1856, p. 517, pi. 20, fig. 1.— BouLENGER, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, pp. 101-102. 

Type . — Unknown . 

Type locality. — Ceylon (in error), here restricted to Orizaba, Veracruz. 

Range. — Atlantic slopes from central Veracruz to Costa Rica. 
Reported in Mexico from the states of Veracruz, Campeche, Yucatan, 
and Chiapas (Usumacinta River across from Piedras Negras, Peten, 
Guatemala) . 

CORYTHOPHANES PERCARINATUS Dum£ril 

Corythophanes percarinatus Dum^ril, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, 
p. 518, pi. 20, fig. 3. — BocouRT, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 3, 1874, pp. 120-122, pi. 17, fig. 2.— Smith, Journ. 
Washington Acad. Sci., vol. 39, 1949, p. 34. 

Type. — Mus. Hist. Nat. Paris. 

Type locality. — Escuintla, Guatemala. 

Range. — Pacific slopes of Guatemala and Chiapas, and Atlantic 
slopes in Alta Verapaz, Guatemala. Known in Mexico only from Col- 
onia Hidalgo, 8 kilometers north of La Esperanza, Chiapas. 

Genus LAEMANCTUS Wiegmann 

Laemandus (part) Wiegmann, Herpetologia Mexicana, pt. 1, 1834, pp. 45-46. 

Genotype. — Laemanctus longipes Wiegmann. 

Range. — Lowlands from San Luis Potosi south to and including 
Guatemala, and British Honduras; ? Colima. 
Species. — Four, all occurring in Mexico. 

KEY TO MEXICAN SPECIES OF LAEMANCTUS 

1. Scales on upper surface of snout much larger than those on occipital region. _ 2 

Scales on upper surface of snout not distinctly larger than those on occipital 

region; no dorsal crest; part of scales on body bicarinate or tricarinate; head 

yellowish above, the posterior contour black; body above dark purple or 



•• While Boie designated Merrem's species as type of his genus Corytophanes, he did not actually print the 
combination. That it was his intent so to spell the name is indicated in his article in vol. 21 (1828) of Isis 
von Oken, p. 363, where he (Boie) spells the name Corytophanes. Kaup (Isis von Oken, vol. 21, 1828, p. 
1147) does likewise. 



70 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

dark green, with blackish cross bands; a light lateral streak; lilac below; 690 
mm deborrei (p. 7 1) 

2. A series of projecting triangular scales on the posterior contour of the head; a 

feeble dorsal crest 3 

No distinct serration on the posterior contour of head; no dorsal crest present; 
reddish or olive above with darker bands across back; a broad dark band 
from the eye to foreleg, passing through tympanum, bordered by a yellowish 
streak inferiorly; a yellowish streak from axilla to groin sometimes broken; 
a yellow spot on each side of base of tail; lower surfaces pink; 730 mm. 

longipes (p. 71) 

3. Scales around middle of body, 57-61 ; reddish or purplish above with dark brown 

bands across body, most distinct in the vertebral region; a dark brown band 
from eye to tympanum, sometimes continued along side of back; a white 
streak from below the eye to forelimb, and from axilla to groin; a white spot 

on each side of base of tail serratus (p. 70) 

Scales round middle of body, 45-51; gulars elongate; a less distinct vertebral 
serration; a white spot in front of and one behind thigh. 

alticoronatus " (p. 70) 

LAEMANCTUS ALTICORONATUS Cope 

Laemanctus alticoronatus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1865, p. 192; 
Proc. Amer. Philos. Soc, 1869, pi. 11. — Boulenger, Catalogue of the 
lizards of the British Museum, vol. 2, 1885, p. 105. 

Type.—U.S.'NM. No. 308; Arthur Schott collector. 

Type locality. — Merida, Yucatan. 

Range. — Yucatan. Reported from Chichen Itza and Merida. 

LAEMANCTUS SERRATUS Cope 

Laemanctus longipes, Dumeril, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 512, 
pi. 21, fig. 4. — BocouRT, Mission scientifique au Mexique . . . , Etudes sur 
les reptiles, livr. 3, 1874, pp. 114-116, pi. 17, fig. 4 (non Wiegmann, fide 
Cope loc. cit.). 

Laemanctus serratus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 176. — 
Boulenger, Bull. Soc. Zool. France, vol. 2, 1877, p. 463, pi. 7, fig. 3.— 
Smith and Laufe, Trans. Kansas Acad. Sci., vol. 48, 1945, pp. 229-231. — 
Peters, Chicago Acad. Sci. Nat. Hist. Misc., No. 27, 1948, pp. 1-3, fig. 1. 

Type. — Mus. Leyden. 

Type locality. — "Orizaba Valley, Mexico." 

Range. — Lowlands from extreme eastern Guanajuato to Yucatdn. 
Reported from Yucatan: Chichen Itza; Oaxaca: "Oaxaca" (no specific 
locality), Tlacolula; Veracruz: Boca del Rio, Jicaltepec, Misantla, 
Orizaba; Guanajuato: Huasteca Potosina; Hidalgo: Zacualtipan; Cam- 
peche: Champot6n, Oxpemul; San Luis Potosi: Xilitla; Tamaulipas: 
near G6mez Farias. 



M Mrs. Helen T. Oaige suggests the possibility that serratus and alticoronatus may be identical since speci- 
mens from Yucat&n and Campeche seem to show the characters of both (Carnegie Inst. Washington Publ. 
No. 457, 1936, p. 296). 



CHECKLIST OF REPTILES OF MEXICO 71 

LAEMANCTUS DEBORREI Boulenger 

Laemandus deborrei Boulenger, Bull. Soc. Zool. France, vol. 2, 1877, pp. 462- 
463, 465-466, pi. 7, fig. 1; Catalogue of the lizards in the British Museum, 
vol. 2, 1885, p. 106. — Smith, Journ. Washington Acad. Sci., vol. 34, 1944, pp. 
155-156.— Weber, Nat. Geogr. Mag., vol. 87, No. 2, 1945, pp. 209, 210, pi. 
14 (col.). 

Type. — Museum de Bruxelles, two adults; M. Linden collector (?). 

Type locality. — Tabasco. 

Range. — Atlantic slopes from Tabasco to British Honduras. 
Reported from Tabasco: La Venta; Oaxaca: Mountains near Santo 
Domingo (U. S. Nat. Mus.). 

LAEMANCTUS LONGIPES Wiegmann 

Laemandus longipes Wiegmann, Herpetologia Mexicana, pt. 1, 1834, p. 46, pi. 
4. — Boulenger, Bull. Soc. Zool. France, vol. 2, 1877, pp. 464-465, pi. 7, 
fig. 2; Catalogue of the lizards in the British Museum, vol. 2, 1885, pp. 105- 
106. 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Jalapa, Veracruz. 

Range. — Eastern slopes of the plateau from Veracruz to Oaxaca, and 
(?) western slopes in Colima. Reported from Veracruz: Actopam, 
Jalapa, "near Veracruz," Mu-ador, Potrero Viejo; ? Colima: Paso del 
Rio; Oaxaca: Oaxaca. 

Genus BASILISCUS Laurenti 

Basiliscus Laurenti, Specimen medicum exhibens synopsin reptilium, 1768, p. 50. 
Corythaeolus Kaup, Isis von Oken, vol. 21, 1828, p. 1147 (type, Basiliscus vittatus 

Wiegmann) . 
Oedicoryphus Wagler, Natiirliches System der Amphibien, 1830, p. 148 (type, 

Basiliscus vittatus Wiegmann). 
Cristasaura Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 439 (type, 

Cristasaura mitrella Gray^ Basiliscus vittatus Wiegmann). 
Paraloma Cope, Proc. Acad. Nat. Sci. Philadelphia, 1867, p. 181 (type, Dacon- 

ura hivittata Hallo well =Bosi7iscus vittatus Wiegmann). 

Genotype. — Basiliscus americanus ljQMre,ni\= {Basiliscus basiliscus 
(Linnaeus)). 

Range. — Lowlands of Mexico south through Jalisco and Tamaulipas, 
through Central America to northwestern South America. 

Species. — Five; one occurs in Mexico. 

BASIUSCUS VITTATUS Wiegmann 

Basiliscus vittatus Wiegmann, Isis von Oken, vol. 21, 1828, p. 373. — Dum£ril, 
Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, pp. 522-524, pi. 21, figs. 2, 3.— 
BocouRT, Mission scientifique au Mexique . . . , fitudes sur les reptiles, livr. 
3, 1874, pp. 129-131, pi. 17, fig. 3. — Boulenger, Catalogue of the lizards of 
the British Museum, vol. 2, 1885, pp. 109-110. 

Cristasaura mitrella Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 439 
(Brit. Mus. Nat. Hist.; Dyson coll.; Honduras). 



72 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Daconura bivittata Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 482 
(Acad. Nat. Sci. Philadelphia; Nicaragua, here restricted to Greytown). 

Basiliscus (Cristasaura) nuchalis Cope, Proc. Acad. Nat. Sci., Philadelphia, 1862, 
p. 181 (U.S.N.M. No. 5845; Caldwell collector; Greytown, Nicaragua). 

Type. — Zool. Mus. Berlin; F. Deppe collector. 
Type locality. — Mexico, here restricted to Veracruz, Veracruz, 
Range. — The coasts from Jalisco and southern Tamaulipas, south 
to Nicaragua. Reported from numerous localities in the states of 
Tamaulipas, Veracruz, Tabasco, Campeche, Yucatan, Quintana Roo, 
Chiapas, Oaxaca, Guerrero, Michoacan, Colima, and Jalisco. 

Genus IGUANA Laurent! 

Iguana Latjrenti, Specimen medicum exhibens synopsin reptilium, 1768, p. 47. 
Hypsilophus Wagler, Natiirliches System der Amphibien, 1830, p. 147 (type, 
Lacerta iguana Linnaeus). 

Genotype. — Lacerta iguana Linnaeus. 

Range. — Mexico on the Pacific coast from Sinaloa to Chiapas, on 
the eastern coast north to central Veracruz. Absent on the higher 
parts of the plateau. Lowlands of Central America, and central and 
northern South America. 

Species. — Two, one with two subspecies. Extralimital are Iguana 
iguana iguana and Iguana delicatissima of South America. 

IGUANA IGUANA RHINOLOPHA Wlegmann 

Ilguana] H[ypsilophu8] rhinolophus Wiegmann, Herpetologia Mexicana, pt. 1, 

1834, pp. 44-45. 
Iguana tuberculata var. rhinolopha, Boulenger, Catalogue of the lizards in the 

British Museum, vol. 2, 1885, pp. 190-191. 
Iguana iguana rhinolopha, Van Denburgh, Proc. Acad. Nat. Sci. Philadelphia, 

1897, p. 461.— Dunn, Copeia, 1934, p. 1. 
Ilguana Hernandesii Jan, Indice sistematico dei rettili ed anfibi esposti nel 

medesino, 1857, p. 58 (nomen nudum). 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, by inference here restricted to C6rdoba, 
Veracruz. 

Range. — Veracruz and Sinaloa southward along both coasts to 
Panama. Reported from Sinaloa: Mazatlan, Presidio; Nayarit: 
San Bias; Colima: Manzanillo, Tecoman, Colima; Jalisco: ?San Bias 
(possibly Nayarit); Michoacan: Hacienda California, Chacan River 
Sierra Madre, Jorullo; Guerrero: Pie de la Cuesta, Acapulco; Puehla: 
Chiautla; Oaxaca: Tehuantepec, Rio Coatzacoalcos, ?Tetela, Agua 
Fria, Tequesixtlan, Totolapan; Veracruz: Cordoba, Vega de Alatorre, 
vicinity of Cuatotolapam, Matacabestro, Puente Nacional; Tabasco: 
Tlacotalpan, Tenosique; Campeche: Ciudad del Carmen; Quintana 
Roo: Cozumel Island; Chiapas: La Esperanza. 



CHECKLIST OF REPTILES OF MEXICO 73 

Genus CTENOSAURA Wiegmann 

Ctenosaura Wiegmann, Isis von Oken, 1828, p. 371. — Bailey, Proc. U. S. Nat. 
Mus., vol. 73, art. 12, 1928, pp. 1-58, pis. 1-30 (part)." 

Genotype. — Ctenosaura cycluroides Wiegmann [= C. acanthura 
(Shaw)]. 

Species. — Five, all but one of which (C. bakeri of Utilla Island, 
Honduras) occur in Mexico. 

Range. — Baja California, Sonora, and perhaps southern Arizona 
on the west, and Tamaulipas on the east, south along both coasts 
to Panama. 

KEY TO MEXICAN SPECIES OF CTENOSAURA 

1. Small scales between whorls of enlarged scales reduced to 1 row (and no more 

than 1 row) between at least a few whorls (generally near middle of tail), 

often on entire distal half or more of tail 2 

Small scales in at least one complete and another incomplete row between all 
whorls of enlarged scales on tail 3 

2. Fifth whorl of enlarged scales preceded by more than a single row of small 

scales acanthura (p. 74) 

Fifth whorl of enlarged scales preceded by one row and one row only (no in- 
complete row) of small scales hemilopha (p. 75) 

3. Three rows of small scales, complete or incomplete, preceding each of the 

proximal 5 whorls of enlarged caudal scales pectinata (p. 75) 

Small scales reduced to only two rows preceding one or more of the proximal 
five caudal whorls similis (p. 73) 

CTENOSAURA SIMIUS SIMIUS (Gray) 

Iguana {Ctenosaura) similis Gray, in Griffith, Animal Kingdom, by the Baron 

Cuvier, vol. 9, Synopsis . . . , 1831, p. 38. 
Cyclura {Ctenosaura) similis, Wiegmann, Herpetologia Mexicana, 1834, p. 42. 
Ctenosaura similis, Bailey, Proc. U. S. Nat. Mus., vol. 73, 1928, pp. 32-37, pis. 

16-20.— Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 139-140; Journ. 

Washington Acad. Sci., vol. 39, 1949, pp. 35-36. 
Ctenosaura similis [similis], Barbour and Shreve, Occ. Pap. Boston Soc. Nat. 

Hist., vol. 8, 1934, p. 197. — Smith, Ann. Carnegie Mus., vol. 30, 1944, p. 89. 
Ctenosaura completa Bocourt, Mission scientifique au Mexique . . ., Etudes 

6ur les reptiles, livr. 3, 1874, pp. 145-146 (Mus. Hist. Nat. Paris Nos. 2251- 

2252, La Uni6n, El Salvador; Mus. Comp. Zool., 1 spec, "Mexico"; Mus 

Hist. Nat. Paris, 2 cotypes, "Guatemala").^' 

Type. — Lost. 

Type locality. — Restricted (by Bailey) to Tela, Honduras. 

Range. — Coastal areas from the Isthmus of Tehuantepec on the 



•' We are not totally in agreement with Bailey's treatment of the species in this genus, nor with some of 
the more recent records no doubt influenced by his work. The four forms we recognize are more or less 
completely allopatric; at least such suggestions as of C. acanthura in Chihuahua and C. similis in Guerrero 
appear to us wholly unwarranted . The only extensive overlapping of ranges of which we are aware occurs 
on the Atlantic coast and involves C. acanthura and C. similis. 

88 Bocourt (loc. cit.) states that the types are from "Guatemala" and "La Union," EI Salvador. Barbour 
and Loveridge (Bull. Mus. Comp. Zool., vol. 69, 1929, p. 247), however, state that a cotype in the Mus. 
Comp. Zool. is from "Mexico." The type locality is here restricted to La Union. 



74 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Pacific and central Veracruz on the Atlantic south to Panama. 
Recorded in Mexico from the states of Veracruz, Tabasco, Campeche, 
Yucatan, Quintana Roo (including Mujeres and Cozumel Islands), 
Oaxaca, and Chiapas (Tonala, Palenque)." 

CTENOSAURA ACANTHURA (Shaw) 

Lacerta acanthura Shaw, General zoology, vol. 3, 1802, pp. 216-217. 

Ctenosaura acanthura, Gray, Catalogue of the lizards in the collection of the 

British Museum, 1845, p. 191.— Bailey, Proc. U. S. Nat. Mus., vol. 73, 1928, 

pp. 9-16, pis. 1-4. — Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 137-139; 

Journ. Washington Acad. Sci., vol. 39, 1949, p. 36. 
Cyclura teres Harlan, Journ. Acad. Nat. Sci. Philadelphia, vol. 4, 1825, pp. 242- 

251, pi. 26 (type lost; Tampico, Tamaulipas). 
Ctenosaura cycluriodes Wiegmann, Isis von Oken, 1828, vol. 21, p. 371 (Zool. Mus. 

Berl. Nos. 576, 578, Mus. Comp. Zool. No. 2253, cotypes; Mexico, here 

restricted to Veracruz, Veracruz). 
Iguana (Ctenosaura) arrnata Gray, in Griffith, Animal Kingdom, by the Baron 

Cuvier, vol. 9, Synopsis . . . 1831, p. 38 (type lost; no type locality cited, 

here restricted to Tampico, Tamaulipas). 
Cyclura shawii Gray, loc. cil. (synonymic name for Iguana (Ctenosaura) acanthura 

Gray). 
Iguana (Ctenosaura) lanceolata Gray, loc. cit. (type lost; no type locality cited, 

here restricted to Tampico, Tamaulipas). 
Iguana (Ctenosaura) hellii Gray, loc. cit. (type lost; no type locality cited, here 

restricted to Tampico, Tamaulipas). 
Cyclura articulata Wiegmann, Herpetologia Mexicana, 1834, p. 43 (types unknown; 

type locality, Mexico). 
Cyclura denticulata Wiegmann, ibid., pp. 43-44 (substitute name for Ctenosaura 

cycluroides Wiegmann). 
Cyclura semicristata Fitzinger, Systema reptilium, 1843, p. 56 (synonymic name 

for Cyclura denticulata Wiegmann) .^^ 
Ctenosaura multispinis Cope, Proc. Amer. Philos. Soc, vol. 23, 1886, p. 197 

(U.S.N. M. No. 72737; Captain Dallas collector; Dondominguillo, Oaxaca). 

Type.— Brit. Mus. Nat. Hist. No. XXII, 20-a (^(/e Bailey) ; collector 
unknown. 

Type locality. — Not given by Shaw. Boulenger (Catalogue of the 
lizards in the British Museum, vol. 2, 1885, p. 196) gives "California," 
which probably means Baja California. Since the species does not 
occur there we accept Bailey's restriction to Tampico, Tamaulipas. 

Range. — Atlantic slopes from central Tamaulipas south to the 
Isthmus of Tehuantepec. Reliably reported only from the states of 
Tamaulipas, San Luis Potosi (Pujal), Veracruz, and Oaxaca.^^ 

" Burt and Myers (Stanford Univ. Publ., biol. ser., vol. 8, 1942, p. 299) record the species, in error, from 
Acapulco, Guerrero. 

»» Fitzinger {loc. cit.) attributes this name to Wiegmann, as do also Lichtenstein and Martens (Nomen- 
clator . . . Berolinensis, 1856, p. 8) ; we are, however, unable to locate Wiegmann's use of the name and 
attribute it to Fitzinger, as did Bailey, Proc. U. S. Nat. Mus., vol. 73, 1928, p. 10). 

" Reexamination of specimens in the U. S. Nat. Mus. referred by Bailey (loc. cit.) to C. acanthura reveals 
that two other species were confused with C. acanthura. One specimen from Mirador, Veracruz, is a C. 
similis; and all from Uruapan, Michoacdn; Tehuantepec, Oaxaca (except Nos. 26341, 58498); Isabel Island; 
Tres Marias Islands; Cuicatlfin, Oaxaca; Tlapa and Balsas, Guerrero, are C. pectinata. That all other west- 
coast specimens in his list, from other museums, are also C. pectinata we believe is virtually a certainty. 



CHECKLIST OF REPTILES OF MEXICO 75 

CTENOSAURA PECTINATA (Wiegmann) 

Cychira pedinata Wiegmann, Herpetologia Mexicana, 1834, p. 42, pi. 2. 

Ctenosaura pedinata, Gray. Catalogue of the specimens of lizards in the collection 
of the British Museum, 1845, p. 191. — Bailey, Proc. U.S. Nat. Mus., vol. 73, 
1928, pp. 24-27, pis. 7-11.— Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, 
pp. 134-137; Journ. Washington Acad. Sci., vol. 39, 1949, pp. 36, 37. 

Ctenosaura brevirostris Cope, Proc. Amer. Philos. Soc, vol. 23, 1886, pp. 266-268 
(type, U.S.N. M. No. 24709; John Xantus collector; Colima, Colima). — 
Bailey, op. cit., pp. 27-29, pis. 12, 13, 15. 

Ctenosaura teres brachylopha Cope, Proc. Amer. Philos. Soc, vol. 23, 1886, p. 269 
(type, U.S.N. M. Nos. 7180-3; Bischoff collector; Mazatldn, Sinaloa).— 
Bailey, Proc. U. S. Nat. Mus., vol. 73, art. 12, pp. 22-24, pi. 6. 

Ctenosaura parkeri Bailey, Proc. U. S. Nat. Mus., vol. 73, art. 12, pp. 29-32, 
pis. 14, 15 (type, U.S.N. J\I. No. 18967; P. L. Jouy collector; Barranca 
Ibarra, Jalisco). 

Type. — Mus. Zool. Berlin, No. 574; F. Deppe collector. 

Type locality. — Restricted to Colima, Colima. 

Range. — Pacific coast from southern Sinaloa southward to the 
Isthmus of Tehuantepec. Recorded from Isabel and the Tres Marias 
Islands and the states of Sinaloa, Durango (Ciudad, Ventanas), 
Nayarit, Jalisco, Colima, Alichoacan, Morelos, Guerrero, Puebla, and 
Oaxaca. A record for the state of Guanajuato is unacceptable. 

ctenosaura HEMILOPHA (Cope) 

Cyclura (Ctenosaura) hemilopha Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, 
pp. 105-106. 

Ctenosaura hemilopha, Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 312. — 
Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 39, 
66-71.— Bailey, Proc. U. S. Nat. Mus., vol. 73, 1928, pp. 17-22, pi. 5.— 
Smith, Univ. Kansas Sci. BuU., vol. 22, 1935, pp. 140-142, pi. 23, fig. 1. 

Ctenosaura interrupta Bocourt, Le Naturaliste, vol. 2, 1882, p. 47 (Mus. Hist 
Nat. Paris Nos. 2243, 2245, 2843, Brit. Mus. Nat. Hist. 1 spec, cotypes 
M. Botta collector; "Baja California", here restricted to Cape San Lucas) 

Ctenosaura conspicuosc Dickerson, Bull. Amer. Nat. Hist., vol. 41, 1919, p 
461 (U.S.N.M. No. 64440; C. H. Townsend collector; San Esteban Island 
Sonora) . 

Ctenosaura insulana Dickerson, op. cit., pp. 462-463 (U.S.N.M. No. 64439 
C. H. Townsend collector; Cerralvo Islands, Baja California). 

Type. — U.S.N.M. No. 529, four cotypes; John Xantus collector. 

Type locality. — Cape San Lucas, Baja California. 

Range. — The southern two-thirds of Baja California, central 

Sonora (including San Pedro Nolasco Island) south to northern 

Sinaloa, and southwestern Chihuahua. Recorded only from the 

states cited. It is possible that the species extends as far north as 

southern Arizona, 

Genus ENYALIOSAURUS Gray 

Enyaliosaurus Gray, Catalogue of the specimens of lizards in the collection of 
the British Museum, 1845, p. 192. 
861316—50 6 



76 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Cachryx Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 124 (genotype, Cteno- 
saura erythromelas Boulenger). 

Genotype. — Cyclura quinguecarinata Gray. 

Range. — Michoacan to the Isthmus of Tehuantepec, the Peninsula 
of Yucatan, and northern Guatemala. 

Species. — Five, all but one of which (C. palearis of Guatemala) 
occur in Mexico. 

KEY TO MEXICAN SPECIES OF ENYALIOSAURUS 

1. Whorls of enlarged caudal scales not interspersed with small scales. 

defensor (p. 77) 
Whorls of enlarged caudal scales interspersed with smaller scales 2 

2. Dorsal and lateral caudal scales of nearly uniform character in each whorl. 

erythromelas (p. 77) 

Dorsal and lateral caudal scales not of nearly uniform character but median 

longitudinal series and outer lateral series differentiated 3 

3. Median row of dorsal scales extending to or almost to sacrum, noticeably 

raised, of medium height; tail twice length of body.quinquecarinata (p. 76) 

Median row of dorsal scales extending only two-thirds distance to sacrum 

and not noticeably raised; tail about lyi times body length. .clarki (p. 76) 

ENYALIOSAURUS QUINQUECARINATUS (Gray) 

Cyclura quinquecarinata Gray, Zoological miscellany, 1842, p. 59. 
Enyaliosaurus quinquecarinatus, Gray, Catalogue of the specimens of lizards in 

the collections of the British Museum, 1845, p. 192. 
Cyclura (Ctenosaura) quinquecarinata, Cope, Proc. Amer. Philos. Soc, vol. 11, 

p. 161. 
Ctenosaura quinquecarinata, Sumichrast, Bull. Soc. ZooL, vol. 10, 1880, p. 175. — 

Bailey, Proc. U. S. Nat. Mus., vol. 73, art. 12, 1928, pp. 42-44, pis. 24, 25, 26. 

Type. — Brit. Mus. Nat. Hist. No. 61; collector unknown. 

Type locality. — Unknown (restricted to Tehuantepec, Oaxaca, by- 
Bailey, loc. cit.). 

Range. — Lowland regions of southern Oaxaca. Reported from 
Oaxaca: Tehuantepec, "Hugma, Temiinos," San Pedro, and Mixte- 
quilla Mountains, between Salina Cruz and Tequesixtlan. 

ENYALIOSAURUS CLARKI (Bailey) 

Enyaliosaurus quinquecarinatus, DuGfes, La Naturaleza, ser. 2, vol. 2, 1897, pp. 

523-524, pi. 34 (color). 
Ctenosaura clarki Bailey, Proc. U. S. Nat. Mus., vol. 73, art. 12, 1928, pp. 44-46, 

pi. 27. 

Type. — Mus. Comp. Zool. No. 22454; Hans Gadow collector. 

Type locality. — Ovopeo, Michoacan, elevation 1,000 feet. 

Range. — Known definitely only from the type locality; a record for 
Cuautla, Morelos, probably belongs here (Duges, La Naturaleza, ser. 
2, vol. 2, 1896, p. 480). 



CHECKLIST OF REPTILES OF MEXICO 77 

ENYAUOSAURUS ERYTHROMELAS (Bonlenger) 

Ctenosaura erythromelas Boulenger, Proc. Zool. Soc. London, 1886, p. 241, pi. 

23 (color). — Bailey, Proc. U. S. Nat. Mus., vol. 73, art. 12, 1928, pp. 46-48, 

pis. 28, 29. — Smith, Occ. Pap. Univ. Michigan Mus. Zool., No. 388, 1938, p. 15. 
Cachryx erythromelas, Cope, U. S. Nat. Mus. Bull. 32, 1887, p. 43. 
Ctenosaura (Cachryx) annectens Werner, Jahrb. Hamburg Wiss. Anst., pt. 2, 

1911, p. 25 (locality unknown; P. Phol collector, Hamburg Museum; 

"Mexico"). 

Type.— Brit. Mus. Nat. Hist. No. 1 (Jide Bailey, loc. cit., p. 48) ; 
purchased alive. 

Type locality. — Unknown; here restricted to Balchacaj, Campeche. 
Range. — Known only from Balchacaj, Campeche. 

ENYAUOSAURUS DEFENSOR Cope 

Cachryx defensor Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 124; Proc. 

Amer. Philos. Soc, 1869, pi. 10. — Bocourt, Mission scientifique au Mexique 

. . ., Etudes sur les reptiles, livr. 3, 1874, pp. 148-149, pi. 17 bis, fig. 12, 12a. 
Ctenosaura defensor, GOnther, Biologia Centrali-Americana, Reptilia and Ba- 

trachia, 1890, pp. 58-59. — Bailey, Proc. U. S. Nat. Mus., vol. 73, art. 12, 

1928, pp. 48-50, pi. 30. 

Type.—U. S. N. M. No. 12282; male, adult, and half grown, three 
CO types; Arthur C. V. Schott collector. 

Type locality. — Yucatan, here restricted to Chichen Itza. 

Range. — Northern part of the Yucatan Peninsula. Reported 
only from Chichen Itza, Yucatan. 

Genus DIPSOSAURUS Hallowell 

Dipso-saurus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 1854, p. 92. 

Genotype. — Crotaphytus dorsalis Baird and Girard. 

Range. — Sonora, Baja California, and some coastal islands of the 
Gulf. In United States: southern and central Arizona, Colorado and 
Mojave Deserts east to the Colorado River, southern Nevada, and 
Utah. 

Species. — Three, with five forms recognized. 

KEY TO MEXICAN SPECIES OF DIPSOSAURUS 

1. Two or more scale rows between nostril and rostral (62-83 percent) ; frequently 

with longitudinal dark lines on the body dorsalis dorsalis (p. 78) 

One scale row between nostril and rostral (75 to 80 percent) ; dark lateral lines 
usually lacking 2 

2. Dorsal pattern consisting of large, round, white spots, and no longitudinal lines; 

brown reticulation on throat enclosing round or oval cream spots. 

dorsalis sonoriensis (p. 79) 

Dorsal pattern consisting of short longitudinal streaks, and few if any light 

spots, which are not dark-edged; throat either striped with dark longitudinal 

streaks or without streaks, in which case large rounded spots are present. 3 



78 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

3. Gular region without longitudinal dark streaks; large rounded spots on lower 

jaw and side of neck; central gular region suffused with dark brown; femoral 

pores average 19.1 catalinensis (p. 78) 

Gular region with longitudinal dark streaks, without definite rounded light 
spots laterally, sometimes suffused with dark centrally , 4 

4. Femoral pores fewer, average 18.46 dorsalis lucasensis (p. 78) 

Femoral pores more numerous, average 21.8 carmenensis (p. 78) 

DIPSOSAURUS CARMENENSIS Van Denbnrgh 

Dipsosaurus carmenensis Van Denburgh, Occ. Pap. California Acad. Sci., No. 
10, vol. 1, 1922, pp. 81-82. 

Type. — California Acad. Sci. No. 50504; Joseph R. Sleviii collector. 

Type locality. — Carmen Island, Baja California. 

Range. — Carmen Island and Los Coronados Islands, Baja California. 

DIPSOSAURUS CATAUNENSIS Van Denburgh 

Dipsosaurus catalinensis Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 
vol. 1, 1922, pp. 83-84. 

Type. — California Acad. Sci. No. 50505; Joseph R. Slevin collector. 
Type locality. — Santa Catalina Island, Baja California. 
Range. — Known only from the type locality. 

DIPSOSAURUS DORSALIS LUCASENSIS Van Denburgh 

Dipsosaurus dorsalis lucasensis Van Denburgh, Proc. California Acad. Sci., ser. 4, 
vol. 10, 1920, p. 33; Occ. Pap. California Acad. Sci., No. 10, 1922, vol. 1, 
pp. 78-81. 

Type.— California Acad. Sci. No. 46090. 

Type locality. — "San Jose del Cabo, Baja California, Mexico." 
Range. — Southern tip of Baja California, southward from Santa 
Margarita Island; Cerralbo Island. Reported from Miraflores, San 
Jose del Cabo, Santa Anita, Agua Caliente, Eureka, etc. 

DIPSOSAURUS DORSALIS DORSALIS (Baird and Girard) 

Crotaphytus dorsalis Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 

vol. 6, 1852, p. 126. 
Dipsosaurus dorsalis, Hallowell, ibid., vol. 7, 1854, p. 92. 
Dipsosaurus dorsalis dorsalis. Van Denburgh, Proc. California Acad. Sci., ser. 4, 

vol. 10, No. 4, 1920, pp. 33-34; Occ. Pap. California Acad. Sci., No. 10, vol. 1, 

1922, pp. 73-78, pi. 2. 

Type.— U.S.'N.M. No. ?; John LeConte collector. 

Type locality. — "Desert of Colorado, Cal.," here restricted to Winter- 
haven (=Fort Yuma), Imperial County. 

Range. — Extreme southwestern Utah and eastern California south- 
ward into northwestern Sonora, and eastern Baja California south to 
Magdalena Island; recorded on the latter island and on those of San 
Luis, Angel de la Guarda, San Marcos, Monserrate, and San Jose. 
Reported from Baja California: Volcano Lake, San Felipe Bay, 13 
miles northwest of El Mejor, El Caj6n Canon, San Ignacio, 600 feet, 
etc. ; Sonora: Punta Penasco. 



CHECKLIST OF REPTILES OF MEXICO 79 

DIPSOSAURUS DORSAUS SONORIENSIS Allen 

Dipso-saurus dorsalis sonoriensis Allen, Occ. Pap. Mus. Zool. Univ. Michigan, 
No. 259, 1933, pp. 4-6. 

Type. — Mus. Zool. Univ. Michigan No. 72121; John Piatt, John 
Scofield, and Morrow J. Allen collectors. 

Type locality. — Hermosillo, Sonora. 

Range. — Gulf slope of Sonora, except extreme northwestern part, 
and southward into northern Sinaloa. Reported from Sonora: 5 miles 
southwest of Hermosillo, Miramar, La Posa, Guaymas, Rancho Costa 
Rica, Bocoit; Sinaloa: Ahome. 

Genus SAUROMALUS Dumeril 

Sauromalus Dumeril, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 536. — Shaw, 
Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 269-306, map. 

Euphryne Baird, Proc. Acad. Nat. Sol. Philadelphia, vol. 10, 1858, p. 253 (type, 
Euphryne obesus Baird). 

Genotype. — Sauromalus ater Dumeril. 

Species. — Seven, one with three subspecies. 

Range. — Southern Utah and southern Nevada south through 
southern CaUfornia, western Sonora and the Gulf slopes of Baja 
California. 

KEY TO SPECIES OF SAUROMALUS «o 

1. One or more transverse bands dorsally, across body or rump 4 

No transverse body bands present 2 

2. Largest nuchal scales equally as large as or larger than frontal plates. 

hispidus (adult) (p. 80) 
Largest nuchal scales smaller than frontal plates 3 

3. Dorsal pattern of large, irregular, dark brown or black blotches on a yellowish 

ground color varius (p. 81) 

Dorsal pattern of small, dark brown or black spots on a gray ground color. 

klauberi (p. 80) 

4. Ventral scale rows usually less than 130 5 

Ventral scale rows usually more than 130 6 

5. Dorsal scales in a head length usually less than 20-hispidus (juvenile) (p. 80) 
Dorsal scales in a head length usually more than 20 slevini (p. 80) 

6. Transverse body bands with light centers and dark brown or black borders 

giving a double-banded effect 9 

Transverse bands unicolor 7 

7. Scales around middle of upper foreleg (humeral scales) usually fewer than 

50 8 

Humeral scales more than 50 obesus obesus (p. 81) 

8. No reddish suffusion on dorsal and ventral areas in adult males; maximum 

adult length averaging somewhat less than in o. tumidus; average scale 

counts lesser obesus townsendi (p. 81) 

More or less brilliant reddish suffusion on dorsal and ventral areas in adult 
males; maximum adult length averaging more than in o. townsendi; average 
scale counts greater obesus tumidus (p. 81) 

9. Ventral scale rows 151 or more australis (p. 80) 

Ventral scale rows usually less than 151 ater (p. 80) 

M From Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, p. 303. 



80 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

SAUROMALUS HISPIDUS Stejneger 

Sauromalus hispidus Stejneger, Proc. U. S. Nat. Mus., vol. 14, 1891, p. 409. — 
Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), p. 264, fig. 22.— Van Den- 
burgh, Occ. Pap. California Acad. Sci., No. 10, vol. 1, 1922, pp. 99-101, pis. 
5-6.— Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 270-280. 

Type.—U.S.^M. No. 8563; Thomas H. Streets collector. 

Type locality. —Angel de la Guarda Island, Baja California. 

Range.— Angel de la Guarda, Smith, Pond, Nuevo Amor, Granite, 
Mejia, and South San Lorenzo Islands, Gulf of California, Baja 
California. 

SAUROMALUS ELAUBERI Shaw 

Sauromalus klauberi Shaw, Trans. San Diego Soc. Nat. Hist., vol. 9, 1941, p. 285; 
vol. 10, 1945, pp. 282-283. 

Type.—h. M. KJauber No. 6859. 

Type locality. — Santa Catalina Island, Baja California. 

Range. — The type locality. 

SAUROMALUS SLEVINI Van Denbargh 

Sauromalus slevini Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 
vol. 1, 1922, pp. 97-99.— Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, pp. 280-282. 

TVpg.— California Acad. Sci. No. 50503, adult male; Joseph R. 
Slevin collector. 

Type locality. — South end of Monserrate Island, Baja California. 

Range. — Monserrate, Carmen, and Coronados Islands, Baja Cali- 
fornia. 

SAUROMALUS AUSTRALIS Shaw 

Sauromalus australis Shaw, Trans. San Diego Soc. Nat. Hist,, vol. 10, 1945, 
pp. 286-288. 

Type.—L. M. Klauber No. 30170; Robert S. Hoard collector. 
Type locality. — San Francisquito Bay, Baja California. 
Range. — The mainland of central and southern Baja California on 
the Gulf coast. 

SAUROMALUS ATER Dnm^ril 

Sauromalus ater Dumeril, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 536^ 
pi. 23, figs. 3, 3a.— Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, 
pp. 284-286. 

Type. — Mus. Hist. Nat. Paris; M. Jaures collector? ("qui a ete 
donne . . . parM. Jaures, lieutenant a bordde la fregateZa Z>ari<ztc?e")- 

Type locality. — Unknown; presumably one of the islands in the 
southern part of the Gulf of California, Baja California, here restricted 
to Espiritu Santo Island. 

Range. — Espiritu Santo, Isla Partida, San Francisco, San Diego, 
Santa Cruz, and San Marcos Islands, Gulf of California, Baja Cali- 
fornia. 



CHECKLIST OF REPTILES OF MEXICO 81 

SAUROMALUS VARIUS Dickerson 

Sauromalus varius Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 
464-465.— Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, pp. 288-290. 

TV^e.— U.S.N.M. No. 64441; C. H. Townsend collector. 
Type locality. — San Esteban Island, Gulf of California. 
Range. — The type locality. 

SAUROMALUS OBESUS TOWNSENDI Dickerson 

Sauromalus townsendi Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 

p. 464. 
Sauromalus obesus townsendi, Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 

1945, pp. 290-292. 

IVpe.— U.S.N.M. No. 64442; C. H. Townsend collector. 
Type locality. — Tiburdn Island, Gulf of California. 
Range. — Tiburdn Island, Sonora, and adjacent coast of Sonora at 
least as far south as Guaymas and east to the vicinity of Hermosillo. 

SAUROMALUS OBESUS TUMIDUS Shaw 

Sauromalus obesus tumidus Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 
1945, No. 15, pp. 269-306. 

Type.—L. M. Klauber No. 27323; L. M. Klauber collector. 

Type locality. — Telegraph Pass, Gila Mountains, Yuma County, 
Ariz. 

Range. — Extreme northwestern Sonora, southwestern and south 
central Arizona. Intergrades only, with S. o. townsendi, are recorded 
from Mexico; typical specimens undoubtedly occur in extreme north- 
western Sonora and extreme northeastern Baja California, along the 
Colorado River. 

SAUROMALUS OBESUS OBESUS (Baird) 

Euphryne obesits Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253; United 
States and Mexican boundary survey, vol. 2, Rept., 1859, p. 6, pi. 27. 

Sauromalus obesus obesus, Shaw, Trans. San Diego Soc. Nat. Hist., vol. 10, 1945, 
pp. 295-301. 

Type.—V.S.IS.M. No. 4172; G. H. Thomas collector (?). 

Type locality. — Fort Yuma, Calif. 

Range. — Southeastern California, southern Nevada, southern Utah, 
western Arizona, and extreme northern Baja California. Although 
known from numerous localities within a few hundred feet of the 
Baja California line, none have actually been taken in that state. 

Genus HOLBROOKIA Girard 

Holbrookia Girard, Proc. Amer. Assoc. Adv. Sci., vol. 4, 1851, p. 201. 
Cophosaurus Troschel, Arch, fiir Naturg., 1850 (1852), p. 389 (type, Cophosaurus 
texana Troschel). 



82 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Genotype. — Holbrookia maculata Girard. 

Range. — Northern states of Mexico south to San Luis Potosf and 
Durango. In the United States: Texas, New Mexico, and Arizona. 

Species. — Four, with twelve forms, of which ten are known to 
occur or are to be expected in Mexico. 

KEY TO MEXICAN SPECIES OF HOLBROOKIA 

1. Tail flat, with broad, black, ventral bands; lateroventral black marks placed 

far posteriorly, continued dorsally above lateral fold, and slanting ante- 

roventrally on belly tezana (p. 85) 

Tail rounded; no ventral subcaudal marks save small spots in one subspecies; 
lateroventral marks placed farther anteriorly, not extending upon dorsal 
surface, and slanting posteroventrally on belly 2 

2. Several black spots under tail; dorsal pattern of large, well-defined spots 

equally sharp-edged on all sides; no light dots in pattern. 

maculata lacerata (p. 83) 
No black spots under tail; pattern not as described 3 

3. Dorsal body scales distinctly keeled, rather small 11 

Dorsal body scales not keeled (although sometimes pointed and somewhat 

convex) , larger, flat 4 

4. Tail generally shorter than the snout-vent measurement, usually not over 

51 percent of total length (the higher proportions are of males) 5 

Tail generally as long as or longer than snout-vent measurement, 60-58 
percent of total length (lower proportions are of females) 7 

5. Enlarged supraoculars in contact with frontals maculata elegans (p. 84) 

Enlarged supraoculars separated from frontals by a series of small scales. 6 

6. Size greater, maximum 71 mm., average 59 mm. snout to vent; in males a 

large blue patch surrounding lateroventral black spots on belly. 

raaculata dickersonae (p. 83) 

Maximum size 61 mm., average 50 mm. snout to vent; in males a smaller 

blue patch, or none, partially surrounding lateroventral black spots on 

belly maculata approximans (p. 83) 

7. Femoral pores usually less than 12 8 

Femoral pores usually 12 or more 10 

8. Upper labial region strongly flared, almost flat bunkeri (p. 84) 

Upper labial region less flared, obtuse 9 

9. Scales on limbs distinctly keeled; upper labials strongly imbricate. 

maculata pulchra (p. 84) 

Scales on limbs smooth; upper labials less strongly imbricate. 

maculata dickersonae (p. 83) 

10. Femoral pores about two-thirds as wide as pore scales are long in adult males, 

about one-half in adult females maculata elegans (p. 84) 

Femoral pores about one-half as wide as the pore scales are long, in adult 
males, about one-fourth in adult females maculata thermophila (p. 84) 

11. Anterior gulars a transverse row of 4 subequal scales, the inner pair not 

separated by linear or rounded scales propinqua piperata (p. 85) 

Not so propinqua propinqua (p. 85) 



CHECKLIST OF REPTILES OF MEXICO 83 

HOLBROOKIA MACULATA APPROXIMANS Baird 

Holbrookia approximans Baib.!),^^ Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. 
Holbrookia maculata approximans, Stejneger, North Amer. Fauna, No. 3, 1890, 

p. 109.— Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 721-723, 

pi. 60.— Smith, Handbook of lizards, 1946, pp. 119-122, pi. 17. 
Holbrookia maculata flavilenta Cope, Proc. Acad. Nat. Sci. Philadelphia, 1883, 

p. 10 (Lake Valley, Dona Ana County, N. Mex.; Acad, Nat. Sci. Philadelphia, 

2 cotypes; E. D. Cope collector). 

Type. — Lost. 

Type locality. — Lower Rio Grande of Tamaulipas (Jide Baird, 
United States-Mexican boundary survey, Reptiles, 1859, p. 8). 

Fange. — Western Texas to eastern Arizona, and from southeastern 
Utah southward through Chihuahua and central Coahuila, and 
southward in the west to Jalisco. Reported from Chihuahua: 35 and 
40 miles south of Moctezuma, between Churo and Chiricahui, Ojos 
del Diablo, Rio Santa Maria, Ramos, Casas Grandes; Coahuila: ? 
Concordia, Sierra de Santa Rosa, Buena Vista, Hda. La Rosita; San 
Luis Potosi: 5 miles northeast of Candido Navarro ; Sonora: headwaters 
of San Pedro; Durango: El Oro; Jalisco: Colotlan.®^ 

HOLBROOKIA MACULATA DICKERSONAE Schmidt 

Holbrookia dickersonae Schmidt, Amer. Mus. Nov., No. 22, 1921, p. 2; Bull. 

Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 724-725.— Smith, Univ. Kansas 

Sci. Bull., vol. 22, 1935, pp. 188-189. 
Holbrookia maculata dickersonae, Smith and Mittleman, Trans. Kansas Acad. 

Sci., vol. 46, 1943, pp. 243-244. 

Ty^pe.— U.S.N.M. No. 2668A; Lt. D. N. Couch collector. 

Type locality. — Castanuelas, Coahuila. 

Range. — The northern portion of the central plateau, as far south 
as Guanajuato, and adjacent areas immediately northward in southern 
Coahuila and Durango. Reported from Coahuila: Castanuelas, 
Hipolito, Jaral, Agua Nueva, Jimulco, Saltillo, Alamo de Parras, 10 
miles east of Torredn; Durango: near Avilco, 5 miles north of Conejos, 
25 miles north of Bermejillo, Gomez Palacio; San Luis Potosi: Jesus 
Maria; Zacatecas: La Colorada, Berriozabal; Guanajuato: San Felipe. 

HCLBRCOKIA MACULATA LACERATA Cope 

Holbrookia lacerata Cope, U. S. Nat. Mus. Bull. 17, 1880, p. 15. — Schmidt, Bull. 

Amer. Mus. Nat. Hist., vol. 46, 1922, p. 718. 
Holbrookia maculata lacerata, Stejneger, North Amer. Fauna, No. 3, 1890, 

p. 109.— Smith, Handbook of lizards, 1946, pp. 122-124, pi. 18. 

91 The type locality for this name strongly indicates that the species involved is //. propinqua, since that is 
the only one of this group whose range reasonably may include the state of Tamaulipas. Nevertheless, 
since the description better fits H. maculata, and since Baird presumably well knew H. propinqua (having 
described it also), we retain the previous allocation of the name H. approximans pending further investiga- 
tions. 

•' A number of records for "Tamaulipas,' without specific locality, are regarded erroneous either by 
misidentification or incorrect locality data. 



84 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type.— V.S.N. M. No. 10160. 

Type locality. — Erath County, Tex., west of the Upper Brazos; on 
the Guadalupe River in Kendall or Comal County. Here restricted 
to Erath County, Tex. 

Range. — Central Texas plains region south to northern Coahuila. 
Recorded in Mexico only from Coahuila: Sabinas. 

HOLBROOKIA MACULATA PULCHRA Schmidt 

Holbrookia pulchra Schmidt, Amer. Mus. Nov., No. 22, 1921, p. 1. 

Holbrookia maculata pulchra, Smith, Handbook of lizards, 1946, pp. 124-126, pi. 19. 

Type. — Amer. Mus. Nat. Hist. No. 14777; R. D. Camp collector. 
Type locality. — Carr Canyon, Huachuca Mountains, Ariz., 5,200 feet. 
Range. — Known from the general region of the type locality; to be 
expected in adjacent Chihuahua and Sonora. 

HOLBROOKIA MACULATA THERMOPHILA Barbour 

Holbrookia thermophila Barbour, Proc. New England Zool. Club, vol. 7, 1921, 

p. 79. 
Holbrookia elegans thermophila, Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, 

pp. 194-195, pis. 27, fig. 1, pi. 28, fig. 4. 
Holbrookia maculata thermophila, Smith, Handbook of lizards, 1946, pp. 129-132, 

pi. 21. 

Type.— Mus. Comp. Zool. No. 14281; W. W. Brown and J. E. 
Thayer collectors. 

Type locality. — San Jose de Guaymas, Sonora. 

Range. — Sonora; possibly in northern Sinaloa. Reported from 
Sonora: 8 miles south of Nogales, 53 miles south of Nogales, 8 miles 
south of Magdalena, Canoa, near Noria, 30 miles south of Noria, 5 
miles southwest of Hermosillo, 54 miles south of Hermosillo, 10 miles 
north of Guaymas, Batamotal, Alamos, Guirocoba. ISinaloa: ?Bacu- 
birito. 

HOLBROOKIA MACULATA ELEGANS Bocourt 

Holbrookia Bischoffi Cope, Proc. Acad. Nat. Sci. Philadelphia, 1868, p. 310 

(nomen nudum). 
Holbrookia elegans Bocourt, Mission scientifique au Mexique . . . , Etudes sur 

les reptiles, livr. 3, 1874, pp. 164-165, pi. 17 bis, figs. 8, 8a. 
Holbrookia elegans elegans, Smith, Univ. Kansas Sci. Bull., vol. 22, No. 8, 1935, 

pp. 191-194. 

Type.— Mus. Hist. Nat. Paris. 
Type locality. — Mazatlan, Sinaloa. 

Range. — Sinaloa. Reported from Presidio de Mazatlan, Mazatlan, 
Escuinapa, ?Bacubirito, Rosario. 

HOLBROOKIA BUNKERI Smith 

Holbrookia bunkeri Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 185-188, 
pi. 28, fig. 3. 



CHECKLIST OF REPTILES OF MEXICO 85 

Type.— Univ. Kansas Mus. Nat. Hist. No. 19236; David H. Dunkle 
and Hobart M. Smith collectors. 

Type locality. — 15 miles south of Juarez, Chihuahua. 
Range. — Known only from the type locality. 

HOLBROOKIA PROPINQUA PROPINQUA Baird and Girard 

Holbrookia propinqua Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 
vol. 6, 1852, p. 126.— Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, 
p. 714.— Smith, Handbook of lizards, 1946, pp. 132-134, pi. 22. 

Holbrookia propinqua propinqua Smith and Burger, Trans. Kansas Acad. Sci., 
vol. 53, 1950, p. 169. 

Type . — Unknown . 

Type locality. — Between Indianola and San Antonio, Tex., here 
restricted to 9 miles southwest of Somerset, Atascosa County. 

Range. — Central and southern Texas, southward presumably into 
Tamaulipas. No definite record for Mexico is known. 

HOLBROOKIA PROPINQUA PIPERATA Smith and Burger 

Holbrookia propinqua piperata Smith and Burger, Trans. Kansas Acad. Sci., 
vol. 53, 1950, pp. 167-169. 

Type. — Univ. Illinois Mus. Nat. Hist. No. 4048; R. W. Reese and 
P. W. Smith collectors. 

Type locality. — Beach at Etiopa, 2 miles south of Tecolutla, Veracruz. 

Range. — Presumably southern Tamaulipas and northern Veracruz; 
recorded only from the type locality. 

HOLBROOKIA TEXANA (Troschel) 

Cophosaurus texanus Troschel, Wiegmann's Arch, fiir Naturg., Jahrg. 16, vol. 

1, 1850 (1852), p. 289, pi. 6. 
Holbrookia texana, Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, vol. 

6, 1852, p. 124.— Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 286- 

289, fig. 30.— Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 190-191 

(very numerous localities listed). 
Holbrookia affinis Baird and Girard, Proc. Acad. Nat. Sci. Pliiladelphia, vol. 

6, 1852 (types unknown; San Pedro, Tex.). 

Type. — Not known. 

Type locality. — New Braunfels, on the Guadalupe River, Tex.; 
latitude 28° N. (=New Braunfels, Comal County, Tex.). 

Range. — Southeastern Arizona to central Texas and southward 
through the northern parts of all the border states in Mexico except 
Baja California. Reported from Tamaulipas, San Luis Potosf, 
Nuevo Leon, Coahuila, Chihuahua, Durango, and Sonora. 

Genus CALLISAURUS Blainville 

Callisaurus Blainville, Nouv. Ann. Mus. Hist. Nat. Paris, vol. 4, 1835, p. 286. 
Megadactylus Fitzinger, Systema reptilium, 1843, p. 59 (type, Callisaurus dra- 
conoidcs Blainville). 



86 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Homalosaurus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 179 
(type, Homalosaurus ventralis Hallowell). 

Genotype. — Callisaurus draconoides Blainville. 

Range. — Southwestern United States, southward from Nevada to 
Baja Cahfornia, Sonora, and adjacent islands. 

Species. — One species with 10 forms; only one (C. d. myurus Rich- 
ardson of Nevada) is extralimital, 

KEY TO MEXICAN SPECIES OF CALLISAURUS 

1. A fringe of small spinose scales along one side of toes somewhat as in Uma. 

draconoides crinitus (p. 87) 
No such fringe 2 

2. Males with two nearly vertical lateral black blotches usually followed by a 

third black spot; dorsal caudal bars never black, usually with strongly 

undulate or pointed posterior margins draconoides draconoides (p. 86) 

Males with two large oblique lateral black blotches usually not followed by a 
third black spot; dorsal bars black and undulate or not 3 

3. Dark dorsal bars on tail with strongly undulate or pointed posterior margins 

from base to beyond middle of tail; dorsal tail bars never black. 

draconoides carmenensis (p. 87) 

Dark dorsal bars on tail with nearly straight or slightly undulate posterior 

margins at least from middle to end of tail; dorsal tail bars often black. _ 4 

4. Dorsal tail bars black in both sexes; usually more or less red in dorsal coloration. 

draconoides splendidus (p. 87) 
Dorsal tail bars black in males only ; usually little or no red in dorsal coloration. 5 

5. Lateroventral black bars in males ill defined, blurred or obsolete, usually united 

below 6 

Lateroventral black bars in males well defined, intense, separate below 8 

6. Lamellae under fourth toe 25-28 draconoides bogerti (p. 89) 

Lamellae under fourth toe 30 or more 7 

7. Lamellae under fourth toe 30 to 33 draconoides brevipes (p. 88) 

Lamellae under fourth toe 35-39 draconoides inusitatus (p. 88) 

8. Femoral pores generally 16 or fewer; hind leg generally 92 percent of body 

length, or more; tail generally 58 percent of total length, or more; inter- 
parietal and supraorbital semicircles separated completely. 

draconoides gabbii (p. 87) 
Femoral pores generally 17 or more; hind leg generally 91 percent of body 
length, or less; tail generally 56 percent of total length, or less; interparietal 
and supraorbital semicircles usually not separated completely. 

draconoides ventralis (p. 88) 

CALLISAURUS DRACONOIDES DRACONOIDES Blainville 

Callisaurus draconoides Blainville, Nouv. Ann. Mus. Hist. Nat. Paris, vol. 4, 

1835, p. 286, pi. 24, fig. 2, 2a.— Schmidt, Bull. Amer. Mus. Nat. Hist., 

vol. 46, 1922, pp. 646-647, fig. 5, pi. 51. figs. 1-3. 
Callisaurus draconoides draconoides, Cope, Ann. Rept. U. S. Nat. Mus., 1898 

(1900), p. 272, fig. 24.— Van Denbuegh, Occ. Pap. California Acad. Sci., 

No. 10, 1922, pp. 142-145. 

Type. — Mus. Hist. Nat. Paris; M. Botta collector. 
Type locality. — "California," here restricted to Cape San Lucas, 
Baja California. 



CHECKLIST OF REPTILES OF MEXICO 87 

Range. — Cape region of Baja California; Isla Espiritu Santo. 

CALLISAURUS DRACONOIDES CARMENENSIS Dickerson 

Callisaurus carmenensis Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 

p. 465. 
Callisaurus draconoides carmenensis, Van Denburgh, Occ. Pap. California 

Acad. Sci., No. 10 (vol. 1), 1922, pp. 145-148.— Linsdale, Univ. California 

Publ. Zool., vol. 38, 1932, p. 358. 
Callisaurus plasticus Dickerson, ibid., p. 466 (Agua Verde Bay, Baja California, 

Amer. Mus. Nat. Hist. No. 5349; C. H. Townsend collector). 

Type. — Amer. Mus. Nat. Hist. No. 5388, male; C. H. Townsend 
collector. 

Type locality. — Carmen Island, Gulf of California, Baja California. 

Range. — The southern two-thirds of Baja California, except the 
Pacific coast in the center of that area, and the Cape region, between 
lat. 29°30' N. and 24° N.; the islands of San Jose, Carmen, San Mar- 
cos, South Santa Ines, Coronados, and Santa Margarita. 

CALLISAURUS DRACONOIDES SPLENDIDUS Dickerson 

Callisaurus splendidus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
p. 467.— Schmidt, ibid., vol. 46, 1922, pp. 647-648, pi. 51, fig. 1, and text fig. 
6. — Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 148- 
150. 

Type.—kmeT. Mus. Nat. Hist. No. 5372, male; C. H. Townsend 
collector. 

Type locality. — Isla Angel de la Guarda, Gulf of California, Baja 
California. 

Range. — Island of Angel de la Guarda, Baja California. 

CALLISAURUS DRACONOIDES CRINITUS Cope 

Callisaurus crinilus Cope, Amer. Nat., vol. 30, 1896, p. 1049. — Van Denburgh, 
Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 140-142.— Schmidt, Bull. 
Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 645-646, pis. 51-52, text fig. 4. 

Callisaurus draconoides crinilus, Tevis, Copeia, 1944, pp. 7-12. 

Callisaurus rhodostictus Cope, ibid., vol. 30, 1896, pp. 1049-1050 ("Lower Cali- 
fornia," here restricted to Santo Domingo; A. W. Anthony coll.; U. S. Nat. 
Mus.). 

Type.—V. S. N. M. No. 14895. 

Type locality. — "Lower California" [=Ballenas Bay, Baja Cali- 
fornia]. 

Range. — Along the Pacific shore of central Baja California from 
Millers Landing (lat. 28°30' N.) to Santo Domingo (lat. 25°30' N.). 

CALLISAURUS DRACONOIDES GABBII Cope 

Callisaurus ventralis gabbii Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), p. 272. — 
Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 648-651, pi. 51, 
fig. 1 (part). 



88 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Callisaurun draconoides gabbii, Linsdale, Univ. California Publ. Zool., vol. 38, 
1932, pp. 358-359; Proc. Amer. Acad. Arts and Sci., vol. 73, 1940, pp. 220- 
221.- — Tevis, Copeia, 1944, pp. 7-12, map fig. 2. — Smith, Handbook of 
lizards, 1946, pp. 138-145, pi. 24. 

Type. — Apparently none designated. 

Type locality. — "Northern Lower California," here restricted to 
Gardners Laguna, Salton River. 

Range. — Southern Nevada, western Arizona, and southeastern 
California southward into extreme northwestern Sonora and north- 
eastern Baja California. Reported from several localities in Baja 
California, and from Sonora: Colorado River, Shoal Point. 

CALLISAURUS DRACONOIDES VENTRAUS (Hallowell) 

Homalosaurus ventralis Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 

1852, p. 179. 
Callisaurus ventralis, Baird, United States and Mexican Boundary Survey, vol. 2, 

pt. 2, 1859, Rept., p. 8. 
Callisaurus ventralis ventralis, Camp, Univ. California Pubis. Zool., vol. 12, 1916, 

pp. 508, 519. — Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, 

pp. 152-163, pi. 12 (part). — Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 

1922, p. 651. 
Callisaurus draconoides ventralis, Linsdale, Univ. California Publ. Zool., vol. 38, 

1932, p. 359. — Smith, Handbook of lizards, 1946, pp. 146-147, pi. 26. 

Tijpe.—JJ. S. N. M. No. 2670; S. W. Woodhouse collector. 

Type locality. — New Mexico, here restricted to Tucson, Ariz. 

Range. — Southern Arizona except the western and eastern edges, 
southward into northern Sonora. Recorded in Mexico only from the 
latter state, from between Sonoyta and Puerto Penasco. 

CALLISAURUS DRACONOIDES INUSITATUS Dickerson 

Callisaurus inusitatus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 
pp. 465-466. — Van Denbtjrgh, Occ. Pap. California Acad. Sci., No. 10, 
1922, pp. 150-152. 

Callisaurus ventralis inusitatus, Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 
1922, p. 648, pi. 51, fig. 1, pi. 52, fig. 2. 

Type. — Amer. Mus. Nat. Hist. No. 5324, male; C. H. Townsend 
collector. 

Type locality. — Tibur6n Island, Sonora. 

Range. — Tiburon Island, and adjacent Sonoran coast to Guaymas. 

CALLISAURUS DRACONOIDES BREVIPES Bogert and Dorson 

Callisaurus draconoides brevipes Bogert and Dorson, Copeia, 1942, No. 3, 
pp. 173-175. — Bogert and Oliver, Bull. Amer. Mus. Nat. Hist., vol. 83, 
1945, pp. 314, 346-347, 401, 412, 418, 419. 

Type.—Amer. Mus. Nat. Hist. No. 63668; J. W. Hilton collector. 
Type locality. — Guirocoba, 18 miles southeast of Alamos, Sonora, 
Mexico, elevation approximately 1,485 feet. 



CHECKLIST OF REPTILES OF MEXICO 89 

Range. — Extreme southern Sonora (Alamos and Guirocoba) and 
northern Sinaloa (Rio de Choix). 

CALLISAURUS DRACONOIDES BOGERTI Martin del Campo 

Callisaurus draconoides bogerti Martin del Campo, Anal. Inst. Biol. (Mexico), 
vol. 15, 1943, pp. 619-621. 

Type. — Inst. Biol. Mexico; Victoriano Rodriguez collector. 

Type locality. — Isla de los Chivos, en el puerto de Mazatlan, 

Sinaloa. 

Range. — Known only from the type locality and the vicinity of 

Mazatlan. 

Genus UMA Baird 

Uma Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. — Heifetz, Copeia, 
1941, pp. 99-111, figs. 1-7. 

Genotype. — Uma notata Baird. 

Species. — Four, one of which includes two subspecies. Three forms 
occur in Mexico. 

Range. — Southeastern Cahfornia and adjacent Arizona, Sonora 
and Baja California, and southwestern Coahuila. 

KEY TO MEXICAN FORMS OF UMA 

1. Two strongly enlarged auricular lobules; a black bar on rear of femur; dorsal 

pattern with 3 median rows of spots and black chevrons with reticulum out- 
lining whitish dots or blotches exsul (p. 89) 

Four strongly enlarged auricular lobules; no black bar on femur 2 

2. Ventrolateral blotch smaller; femoral pores 20-31, usually fewer than 26. 

notata notata (p. 89) 
Ventrolateral blotch larger; femoral pores 23-32, usually more than 26. 

notata cowlesi (p. 90) 
UMA EXSUL Schmidt and Bogert 

Uma exsul Schmidt and Bogert, Amer. Mus. Nov., No. 1339, 1947, pp. 1-9, 
figs. 1-4. 

Type.— Amer. Mus. Nat. Hist. No. 67404; Karl P. Schmidt and 
C. M. Bogert collectors. 

Type locality. — Dunes 12 miles north of San Pedro de Las Colonias, 
CoahuUa. 

Range. — Presumably restricted to the vicinity of the type locality, 
the northern edge of the dry Laguna de Mayran. 

UMA NOTATA NOTATA Baird 

Uma notata Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. 
Uma notata notata, Heifetz, Copeia, 1941, pp. 101-104, figs. Ic, 2c, 3. 
Uma rufopunctata Cope, Amer. Nat., vol. 29, 1895, p. 939 (U.S.N.M. Nos. 21846-52, 
7 cotypes; "Yuma Desert," Ariz.). 

Type. — U.S.N.M. No. 4124; WiUiamson and Heerman collectors. 
Type locality. — "Mojave Desert," in error; restricted by Heifetz 



90 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

to the "Colorado Desert" and here restricted to the vicinity of Yuma, 
Ariz. 

Range. — Extreme southeastern Baja CaHfornia, southwestern 
Arizona, and northeastern Baja Cahfornia; possibly occurs in Sonora. 
Reported in Mexico only from Baja California. 

UMA NOTATA COWLESI Heifetz 

Uma notata cowlesi Heifetz, Copeia, 1941, pp. 104-106, fig. 5 (distribution). 

Type. — California Acad. Sci. No. 53370; J. E,. Slevin collector. 
Type locality. — Shores of Tepoca Bay, Sonora. 
Range. — The coast of northern Sonora, from Cerro Prieto to 
Tepoca Bay. 

Genus PETROSAURUS Boulenger 

Petrosaurus Boulengek, Catalogue of the lizards in the British Museum, ed. 2, 
1885, vol. 2, p. 205.— MiTTLEMAN, BuU. Mus. Comp. Zool., vol. 91, 1942, 
pp. 110-111. 

Genotype. — Uta thalassina Cope. 

Range. — Southern Baja California and certain adjacent islands in 
the Gulf of California. 
Species. — Two. 

KEY TO SPECIES OF PETROSAURUS 

1. Two series of scales between subocular and labials; snout longer, less truncate; 
three distinct anterior dorsal cross bars; throat yellow, orange, or pale blue 

centrally thalassina (p. 90) 

One series of scales between subocular and labials; snout shorter and more 
truncate; four distinct anterior dark bands; throat brownish marked with 
blackish centrally repens (p. 90) 

PETROSAURUS REPENS (Van Denburgh) 

Uta repens Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 5, 1895, p. 
102, pis. 7, 8, figs, a-e; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 189- 
191.— LiNSDALE, Univ. California Publ. ZooL, vol. 38, 1932, pp. 360-361. 

Petrosaurus repens, Mittleman, BuU. Mus. Comp. Zool., vol. 91, 1942, p. 111. 

Type.— California Acad. Sci. No. 633; W. E. Bryant collector. 
Type locality. — Comondu, Baja California. 

Range. — The southern half of Baja California, exclusive of the 
Cape region; Isla Danzante. 

PETROSAURUS THALASSINUS (Cope) 

Uta thalassina Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 104. — Van 
Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 186-189. 

Petrosaurus thalassinus, Boulenger, Catalogue of the lizards in the British 
Museum, ed. 2, vol. 2, 1885, p. 205. 

T?/pe.— U.S.N.M. No. 5302; John Xantus collector. 
Type locality. — Cape San Lucas, Baja California. 



CHECKLIST OF REPTILES OF MEXICO 91 

Range. — Cape region of Baja California and certain adjacent 
islands. Reported on the mainland from Cape San Lucas, La Paz, 
San Jose del Cabo, Miraflores, etc.; also Isla Partida, Isla Espiritu 
Santo. 

Genus STREPTOSAURUS Mittleman 

Streptosaurus Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 111-117. 

Genotype. — Uta mearnsi Stejneger. 

Range. — Northern portion of Baja California and certain of the 
islands in the Gulf of California; in the United States: San Diego, 
Riverside, and Imperial Counties, Calif. 

Species. — Two. 

KEY TO FORMS OF STREPTOSAURUS 

Ij Throat with numerous round light spots; maximum length about 90 mm. 

raeamsi (p. 91) 

Throat mottled, without definite rounded spots; larger maximum length, 

about 104 mm slevini ^^ (p. 91) 

STREPTOSAURUS MEARNSI (Stejneger) 

Uta mearnsi Stejneger, Proc. U. S. Nat. Mus., vol. 17, 1894, pp. 589-591. — Van 
Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 191-194, pi. 15. 

[Streptosaurus] mearnsi, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, p. 
111.— Stejneger and Barbour, Check list of North American amphibians 
and reptiles, ed. 5, 1943, p. 90. 

TVpe.— U.S.N.M. No. 21882; E. A. Mearns collector. 

Type locality. — "Summit of Coast Range, United States and Mexico 
boundary line, California." 

Range. -Extreme southern California southward through the 
northern haK of Baja California. Reported from Baja California: 
Ensenada, San Salado, San Matias, Parral, Matomi, Coyote Bay. 

STREPTOSAURUS SLEVim (Van Denburgh) 

Uta slevini Van Denburgh, Occ. Pap. California Acad, Sci., No. 10, 1922, pp. 

194-196. 
Streptosaurus slevini, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, p. 112. 

Type. — California Acad. Sci. No. 50506; Joseph R. Slevin collector. 
Type locality. — Mejia Island, Gulf of California. 
Range. — Mejia Island and Angel de la Guarda Island, in the Gulf of 
California. 

Genus CROTAPHYTUS Holbrook 

Crotaphytus Holbrook, North American herpetology, ed, 2, vol. 2, 1842, p. 79. 

Genotype. — Agama coUaris Say. 

«' The senior author has unintentionally referred Pelrosaurus repens to this genus, owing to a lapsus; 
slevini was intended (Smith, Handbook of lizards, 1946, p. 175), 

861316—50 7 



92 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Range. — From Kansas south to Louisiana, west to the Pacific, as far 
north as central Oregon, and southward into northern Mexico. 
Absent from the northern Great Plains and some mountainous areas. 

Species. — Four, with five forms recognized, all but one {C. c. collaris) 
occurring in Mexico. 

KEY TO MEXICAN SPECIES OF CROTAPHYTUS 

1. Black ring or rings about neck complete or incomplete; body pattern not 

reticulated 2 

No black ring about neck, occasionally a black line present on shoulder; dorsal 
pattern boldly reticulated reticulatus (p. 93) 

2. Collar formed of a single bar or ring; snout more elongate insularis (p. 93) 

Collar formed of 2 rings, one or both incomplete 3 

3. Tail more distinctly compressed; 4 or 5 scale rows between suboculars and 

labials, and slightly enlarged scales on the middorsal line on tail. 

dickersonae (p. 93) 
Tail not or scarcely discernibly compressed; minimum scale rows between sub- 
oculars and labials 2 (rarely 3) ; scale rows between supraorbital areas 2 (or 
rarely 3) collaris baileyi (p. 92) 

CROTAPHYTUS COLLARIS BAILEYI " Stejneger 

Crotaphytus baileyi Stejneger, North Amer. Fauna, No. 3, 1890, p. 103, pi. 12, 

fig. 1. 
Crotaphytus collaris baileyi, Stone, Proc. Acad. Nat. Sci. Philadelphia, 1903, p. 

30. — Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 

104-114; pi. 7.— Smith, Handbook of lizards, 1946, pp. 170-172, pi. 33. 
Crotaphytus collaris Burt, Occ. Pap. Mus. Zool. Univ. Michigan, No. 196, 1928, 

pp. 1-19 (part). 
Crotaphytus collaris collaris, Smith (nee Say), Univ. Kansas Sci. Bull., vol. 22, 

1935, p. 143, pi. 24, fig. 1. 

Type.—U.S.^.M. No. 15821; C. Hart Merriam and Vernon Bailey 
collectors. 

Type locality. — Painted Desert, Little Colorado River, Ariz. 

Range. — Southern Idaho and Oregon southeastward through eastern 
California and northern Baja California to San Luis Potosl. Re- 
ported in Mexico from Baja Calijornia: San Salada, Trinidad, Las 
Palmas Canyon, Comondu, La Purisima, 6 miles and 33 miles north of 
Canipole, et cetera; Chihuahua: Chihuahua, Carmen, Progreso, San 
Buenaventura; Durango: 6 miles northeast of Pedricena; Coahuila: 
Villa Acuna, Monclova, 4 miles west of Saltillo, Cuatro Cienegas, 
Pueblo Nuevo, AUende; Nuevo Leon: Pesqueria Grande, Santa Catar- 
ina; Sonora: Hermosillo, Sierra Tule, Los Nogales, Rio Grande 61 
miles south of Los Nogales; San Luis Potosi: 30 miles north of Mate- 
huala, Presa de Guadalupe, 



M With sufficient series it is probable that certain populations now associated with baileyi and collaris will 
be recognized as other subspecies. For instance, certain specimens from San Luis Potosl have three scale 
series between the supraocular regions. C. collaris collaris may occur. 



CHECKLIST OF REPTILES OF MEXICO 93 

CKOTAPHYTUS INSULARIS Van Denburgh and Slevin 

Crotaphytus insularis Van Denburgh and Slevin, Proc. California Acad. Sci., 
ser. 4, vol. 11, 1921, p. 96. — Van Denburgh, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 114-116. 

Type. — California Acad. Sci. No. 49151; J. R. Slevin collector. 
Type locality. — Angel de la Giiarda Island, 7 miles north of Pond 
Island, Gulf of California, Baja California. 
Range. — Restricted to type locality. 

CROTAPHYTUS DICKERSONAE " Schmidt 

Crotaphytus dickersonae Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 

638, fig. 2. 
Crotaphytus collaris dickersonae, Allen, Occ. Pap. Mus. Zool. Univ. Michigan, 

No. 259, 1933, p. 7. 

TV^Je.— U.S.N.M. No. 64451; C. H. Townsend collector. 
Type locality. — Tibur6n Island, Sonora, 
Range. — Known only from the type locality. 

CROTAPHYTUS RETICULATUS Baird 

Crotaphytus reticulatus Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. — 
Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), p. 254, fig. 20.— Smith, 
Handbook of lizards, 1946, pp. 173-175, pi. 34. 

Type.— Two cotypes, U.S.N.M. No. 2692, Ringgold Barracks, 
John H. Clark collector; No. 2731, Laredo, Tex., Arthur C. B. Schott 
collector. 

Type locality. — Laredo and Ringgold Barracks, Tex., here restricted 
to Laredo. 

Range. — Extreme southern Texas, along the Rio Grande, as far west 
as Eagle Pass, and southward in the northern parts of adjacent states 
in Mexico. Reported in Mexico from Tamaulipas: Mier; Nuevo Leon: 
6 miles west of China, 4 miles west of China, 5 miles east of General 
Bravo, "Los Herrars"; Coahuila: 2 miles north of Nava. 

Genus GAMBELIA Baird 

Gamhelia Baird, United States and Mexican boundary survey, vol. 2, Rept., 
1859, p. 7.— Smith, Handbook of lizards, 1946, pp. 158-159. 

Genotype. — Gamhelia wislizenii Baird and Girard. 

Range. — Northern Idaho, Nevada, and Utah southward through 
southeastern California and Baja California, and into western Sonora 
and northern Coahuila. 

Species. — One, with two subspecies. Only the typical form occurs 
in Mexico. 



«' This species has been reduced to subspecific status on the basis of specimens reported by Allen from 
Hermosillo and 40 miles north of Guaymas. Since we have not seen these specimens, and since he did not 
take into consideration some of the more important characters, we retain the specific status for this form 
pending a more considered revision. 



94 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

GAMBEUA WISUZENII WISLIZENII (Baird and Girard) 

Croiaphytus wislizenii Baird and Girard, in Stansbury's Explorations and survey 

of the valley of the Great Salt Lake of Utah, 1852, pp. 340-341, pi. 3.— 

Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, vol. 1, 1922, 

pp. 116-128, pi. 8. 
Crotaphytus (Gambelia) wislizenii, Baird, United States and Mexican Boundary 

Survey, 1859, p. 7. 
Gambelia wislizenii wislizenii, Smith, Handbook of lizards, 1946, pp. 159-164, pi. 

30. 
Croiaphytus gamhelii Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 1852, 

p. 126 (California; U.S.N.M. No. 2722; W. Gambell, collector). 
Croiaphytus fasciatus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 206 

("Jornado del Muerte," Dona Ana County, N. Mex.; U.S.N.M. No. 2736; 

L. Sitgreaves collector). 
Crotaphytus copeii Yarrow, Proc. U. S. Nat. Mus., vol. 5, 1882, p. 441 (U.S.N.M. 

No. 12663; La Paz, Baja Cahfornia). 
Crotaphytus fasciatus (nan Hallowell), Mocquard, Nouv. Arch. Mus. Hist. Nat. 

Paris, ser. 5, vol. 1, 1899, p. 303, pi. 13, fig. 1 (Las Palmas, Baja California; 

Mus. Hist. Nat. Paris). 
Crotaphytus fasciolatus Mocquard, Bull. Mus. Hist. Nat. Paris, 1903, p. 209 (new 

name for C. fasciatus Mocquard) . 

Type.— U.S. l<iM. No. 2770; H. Mollhausen collector. 

Type locality. — Near Santa Fe, N. Mex. 

Range. — Northern Idaho, Nevada, and Utah southward through 
southeastern California and all of Baja California except northwestern 
corner, western Chihuahua and northern Coahuila. Reported from 
the states of Baja California: Cedros and Magdalena Islands, numer- 
ous mainland localities; Sonora: Tiburon Island, numerous mainland 
localities; Chihuahua: 15 miles south of Juarez, Santa Maria, Pro- 
greso. Lake Santa Maria, Chihuahua, 11 miles south of Ahumada, 2 
miles south of Moctezuma; Coahuila: Cuatro Cienegas. 

Genus PHRYNOSOMA Wiegmann 

Tapaja (or Tapaia, p. vi) Oken, Lehrbuch der Naturgeschichte, 1816, p. 295 
(type, Lacerta orbicularis Linnaeus by present restriction; suppressed). 

Tapayia Gray, Ann. Philos., new ser., vol. 26, 1825, p. 197 (type, Lacerta orbicu- 
laris Linnaeus, monotype; suppressed). 

Tapaya Fitzinger.ob Neue Classification der Reptilien, 1826, p. 17 (part only; 
type restricted to T. orbicularis C\x\\eT= Lacerta orbicularis Linnaeus, by 
Malcolm Smith, Fauna of British India, vol. 2, 1935, p. 211).— Bocourt, 
Mission scientifique au Mexique . . ., fitudes sur les reptiles, livr. 4, 1874, 
p. 220. 

Phrynosoma Wiegmann, Isis von Oken, 1828, p. 367. 

Batrachosoma Fitzinger, Systema reptilium, 1843, p. 79 (tj'pe, Phrynosoma coro- 
natum Blainville). 

Tropidogaster Fitzinger {nee Dum6ril and Bibron), loc. cit. (type, Phrynosoma 
cornutum Gray). 

»8 Cuvier (Le regne animal, vol. 2, 1817, p. 35) has been credited with this name by some authorities. He 
did not, however, use the name in latinized form anywhere in the first or 1817 edition of "Le Regne Animal"; 
he uses the name only as "les Tapayes." 



CHECKLIST OF REPTILES OF MEXICO 95 

Anota Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 182 (type, Anota 

m'calln HaUov/ell). 
Tapaya Girard (nee Fitzinger), in Wilkes, United States Exploring Expedition, 

Herpetology, 1858, p. 394 (type, Lacerta orbicularis Linnaeus). 
Doliosaurus Girard, op. cit. p. 407 (type, Phrynosoma platyrhinos Girard). 

Genotype. — Lacerta orbicularis Linnaeus. 

Range. — Nebraska, Kansas, Oklahoma, and Texas westward to the 
Pacific, northward to British Columbia and Montana, and southward 
through Mexico to Guatemala. 

Species. — Fifteen, with about 26 forms recognized; 21 occur in 
Mexico. 

KEY TO MEXICAN SPECIES OF PHRYNOSOMA 

1. Ventral abdominal scales smooth (a few scales may be keeled on breast of 

solare and a few specimens may show faint keeling on some of abdominal 

scales, but this form will have four occipital spines) 6 

Ventral abdominal scales distinctly keeled (rarely indistinctly keeled in 
cornutum) 2 

2. Horns (or horny spines) present on back of skull 3 

No horns or horny spines on back of skull, the scales that form spines in other 

species are low rounded protuberances; two flaring temporal expansions on 
back of head separated by a medial notch; lower jaw with great vertical 
development, the chin shields separated from the infralabials by 4 or 5 
scale rows; 76 mm ditmarsi (p. 104) 

3. Tail at least half length of body (usually more) ; large species with 2 lateral 

abdominal fringes of soft spines more or less clearly developed 4 

Tail shorter or but little longer than head; smaller species, with 1 series of 
lateral soft spines 5 

4. Ventral scales large, 30 to 35 across widest part of belly; scales surrounding 

spines of upper lateral fringe very prominent; greater part of gular scales 
arranged in slightly serrate longitudinal series; 2 very prominent series of 
large scales widely separated on back; tail about as long as body; length, 

170 mm asio (p. 102) 

Ventral scales smaller, 55 to 70 across widest part of abdomen; 3 temporal and 
2 occipital spines well developed; 3 brown bands across frontal area; 3 
radiating lines below eye; rarely ventral keeling is indistinct; length, 140 
mm comutum (p. 99) 

5. Outer temporal part of skull prolonged backward into a very large "horn" 

several times size of small occipital spines; tail as long as head or slightly 

longer; 9-10 femoral pores taurus (p. 104) 

Outer temporal region not prolonged into a large horn; temporal spines about 
the same size as occipital spines; tail shorter than head; femoral pores 
about 10-10 braconnieri (p. 100) 

6. No ear-opening present; tympanum superficial, covered with scales (very 

rarely unsealed in some Mexican specimens of modestum and some 

m'callii 7 

Ear opening present, tympanum usually more or less deeply sunk 10 

7. No lateral fringe of soft spines on sides of abdomen; occipital and temporal 

spines present but rather short; chin shields bordering labial series through- 
out; greater part of tail very slender and subcylindrical, not gradually 
tapering; venter white; chin usually speckled; 84 mm._raodesturQ (p. 101) 
At least 1 series of lateral soft spines 8 



96 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

8. Usually less than 12 femoral-preanal pores, the pore scales usually separated 

by 1 or more intervening scales 9 

Pore series 17-23, the pores surrounded by several small scales and usually 
contiguous; temporal and occipital horns very elongate; 2 series of lateral 
fringes on abdomen; an indistinctly enlarged series of longitudinal gular 
scales on each side of chin, more evident posteriorly; a dark dorsal streak; 
immaculate white on venter; tail much flattened; occasionally traces of 
abdominal keeling present m'callii (p. 99) 

9. Pore series about 8 on each side; 3 enlarged temporals and 2 enlarged occipital 

spines, tending to curve laterally somewhat; small species, 72 mm. snout to 

vent; venter immaculate white platyrhinos goodei (p. 102) 

Pore series 7-12, the temporal and occipital horn much reduced; superciliary 
spine short but heavy and prominent; a single series of scales in lateral 
fringe; venter usually punctate with dark; only rarely is the tympanum 
visible platyrhinos platyrhinos (p. 101) 

10. Four much-elongated, flattened occipital horns, contiguous with a like tem- 

poral series, all directed more or less in same general plane; breast scales 
keeled; 1 distinct lateral fringe and a small secondary fringe indicated. 

solare (p. 104) 

Never more than two strongly developed occipital horns (occasionally a very 

small median single scale may be present) 11 

11. Occipital horns at least twice as long as basal width; chin shields forming 

serrate series strongly visible from dorsal view of head; gulars with serrate 

series of enlarged scales coronatum group 17 

Occipital horns not twice as long as basal width; no serrate chin shields or 
specialized serrate series in gulars 12 

12. Head not strongly notched or emarginate between occipital horns, distance 

between bases of horns one-half to rarely one and one-half times basal 
width of horn; occipital horns distinctly larger than superciliary horns. 

orbiculare group 14 
Head strongly notched or emarginate between occipital horns; horns very 
short, their bases separated by a distance equal to 2 to 3)4 times basal width 
of a horn; occipital and superciliary horns subequal in size. 

douglassii group 13 

13. Tail length a little less than half snout-to-vent length but distinctly longer 

than width of head; gular scales not convex; chest scales smooth. 

douglassii hemandesi (p. 100) 

Tail very short, very slender, length less than width of head; gular scales 

convex, and chest scales keeled douglassii brachycercum (p. 100) 

14. Occipital spines nearly erect, while temporal spines approach the horizontal, 

all relatively long; top of head conspicuously concave, superciliary region 

elevated boucardi (p. 98) 

Occipital spines on same plane as temporals, approaching horizontal, all 
shorter; top of head flat 15 

15. Temporal horns not or barely extending backward as far as posterior tip of 

occipital horns 16 

Temporal horns extending backward farther than occipital horns. 

orbiculare cortezii (p. 98) 

16. Occipital horns extending posteriorly farther than posterior temporals; pre- 

anal area with preanal pores not confined to a single row (usually several 
irregularly placed pores or 2 longitudinal rows of 2 to 4 scales) . 

orbiculare orbiculare (p. 97) 



CHECKLIST OF REPTILES OF MEXICO 97 

Occipital and posterior temporal horns extending back an equal distance (or 
very nearly so); femoral pores 14-14, confined to single row. 

orbiculare dugesii (p. 98) 

17. Fourth (or fifth) temporal horn (counting forward) sharply pointed and 

greatly exceeding fifth (or fourth) in length 18 

Fourth and fifth temporal horns blunt, of nearly equal size, or fifth slightly 
larger 20 

18. Nasal opening large; distance between inner edges less than l^i times maxi- 

mum diameter of nostril cerroense (p. 103) 

Nasal opening moderate, the distance between inner edges more than 1% 
diameters of nostril 19 

19. Postrictal spine absent or rudimentary, .coronatum coronatum (p. 102) 
Postrictal spine present coronatum jamesi (p. 102) 

20. Frontal plates in adults smooth, convex coronatura blainvillii (p. 103) 

Frontal plates in adults rough, striated or rugose, and flat or peaked. 

coronatum. frontale (p. 103) 

PHRYNOSOMA ORBICULARE ORBICULARE " (Linnaeus) 

[Tapayaxin, Lacertus orbicularis Hernandez, Plantas y animales de la Nueva 
Espana, etc., 1651, C. xvi, p. 327, fig. (unnumbered).] 

Lacerta orbicularis Linnaeus, Systema naturae, ed. 12, vol. 1, 1789, p. 1062 (part, 
based on Hernd,ndez). 

Agama orbiculaire, Cuvier, Regne animal, vol. 2, 1817, p. 35. 

Phrynosoma orbiculare, Wiegmann, Isis von Oken, 1828, p. 367. 

Phrynosoma orbiculare orbiculare, Smith, Trans. Kansas Acad. Sci., vol. 37, 1934, 
p. 290, pi. 11, fig. 2, pi. 12, fig. 5. 

Phrynosoma wiegmanni Gray, The zoology of Captain Beechey's voyage . . ., 
1839, p. 96 (type unknown; type locality, "Mexico," here restricted to 
Mexico City, Distrito Federal). 

Tapaya orbicularis longicaudatus Duges, La Naturaleza, ser. 2, vol. 1, 1888, p. 117 
(Alf. Duges Mus., Guanajuato, Guanajuato; Valley of Mexico, here re- 
stricted to Mexico, D. F.). — Smith and Necker, Anal. Esc. Nac. Cienc. 
Biol., vol. 3, 1943, pp. 216-218, pi. 2, figs. 1, 3. 

Type. — Presumably the unnumbered figure in Hernandez, loc. cit. 

Type locality. — Mexico (by inference), here restricted to Mexico, 
D. F. 

Range. — The central plateau of Mexico from Chihuahua and 
Nuevo Le6n southward to Michoacan, Morelos, and Puebla. Re- 
ported from Mexico: Toluca, Lerma, San Andres, 11 miles south of 
Mexico (city), Teotiliuacan, Zumpango; Puebla: Puebla, 15 kilo- 
meters northwest of San Martin, Matamoros near Tezuitldn; Distrito 
Federal: Mexico, Tlalpam, Rio San Juan de Dios, Tlalnepantla, 
between Navitas and Chalco; Hidalgo: Guerrero near Real del 
Mineral, Actopan, El Chico Parque Nacional, 2-3 miles west of 
Tulancingo, near Paclmca; Morelos: near Tres Marias (Km. 57); 
Chihuahua: south of Chihuahua City; Samachique, 21 mUes south 
of Minaca; Durango: Coyotes, El Salto, Ciudad; Jalisco: North of 

w We are unable to agree to the idea of the conspecificity of the Mexican orbiculare group and the douglassil 
group. We feel that among other important distinguishing characters the posterior indentation of the 
skull, so marked in the latter group, has been overlooked in the pnst. 



98 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Rio Santiago, Guadalajara, Bolanos; Guanajuato: Guanajuato (city); 
San Luis Potosi: Jesus Maria, San Pedro; Queretaro: no specific 
locality; Nuevo Leon: Hacienda Pablillo; Tlaxcala: 13 kilometers 
northeast of Tlaxcala; Michoacdn: "above 3000 feet" (Jorullo?); 
Zacatecas: Plateado, Sierra Aladre. 

PHRYNOSOMA ORBICULARE CORTEZII (Aug. Duineril and Bocourt) 

Phrynosoma orbiculare, Wiegmann, Herpetologia Mexicana, 1834, p. 53, pi. 8, 

fig. 1. 
Tapaya orbicularis var. A, Bocourt, ibid, (infra), livr. 4, 1874, pp. 223-224, pi. 11, 

fig. 2. 
Tapaya Corfezii Aug. Dumeril and Bocourt,''^ Mission scientifique au 

Mexique . . ., fitudes sur les reptiles, livr. 1, 1870, pi. 9, fig. 2, 2a-g. 
Phrynosoma orbiculare cortezii, Bonlenger, Catalogue of the lizards in the British 

Museum, ed. 2, vol. 2, 1885, p. 242. — Smith, Trans. Kansas Acad. Sci., vol. 

37, 1934, pp. 291-292, pi. 11, fig. 1, and pi. 12, fig. 6. 

Type. — Mus. Hist. Nat. Paris; Lucien Biart collector. 

Type locality. — "Hacienda del Jasmin," between "Orisaba and 
Cordoba," Veracruz. 

Range. — Eastern half of Puebla and adjoining parts of Veracruz. 
Reported from Veracruz: 15 miles east of San Marcos, 5 kilometers 
east of Las Vigas, Perote, Jalapa, Orizaba; Puebla: San Diego south 
of Tehuacdn, Atzitzintla. 

PHRYNOSOMA ORBICULARE DUGESH (Ang. Dumgril and Bocourt) 

Tapaya Dugesii ^^ Aug. Dumeril and Bocourt, Mission scientifique au Mex- 
ique . . ., Etudes sur les reptiles, livr. 1, 1870, pi. 11, fig. 3, 3a-g. 

Phrynosoma orbiculare (var.) dugesii, Boulenger, Catalogue of the lizards in the 
British Museum, ed. 2, vol. 2, 1885, p. 243. 

Tapaya orbicularis var. B, Bocourt, ibid., livr. 4, 1874, pp. 224-225, pi. 17, 
fig. 3, 3a-g. 

Type. — Mus. Hist. Nat. Paris, two co types; Alfredo Duges collector. 

Type locality. — Colima. 

Range. — Known only from Colima. 

PHRYNOSOMA BOUCARDI (Aug. Dumgril and Bocourt) 

Tapaya Boucardii Aug. Dumeril and Bocourt, Mission scientifique au Mex- 
ique . . ., Etudes sur les reptiles, livr. 1, 1870, pi. 11, fig. 4, 4a-g. — Bocourt, 
ibid., livr. 4, 1874, pp. 225-226. 

Phrynosoma boucardii, Boulenger, Catalogue of the lizards in the British 
Museum, ed. 2, vol. 2, 1885, p. 243. — Smith, Journ. Washington Acad. Sci., 
vol. 39, 1949, p. 38. 

Type. — Mus. Hist. Nat. Paris, two males, two females; A, Boucard 
collector. 

Type locality. — "Plateau of Mexico," here restricted to Zimapan, 
Hidalgo. 

•' The name cortezii must be accredited to both authors of livr. 1. The name when it appears is Tapaya 
coTtezii (nobis), and when quoted by Bocourt both names are used. 

" In 1870 the name was used as a full species, when associated with the figure. The name is dropped in 
the livr. 4 of the same work where it is regarded as a variety B of Tapaya orbicularis. 



CHECKLIST OF REPTILES OF MEXICO 99 

Range. — Central plateau region. Reported from Guanajuato: 
Le6n; Hidalgo: Zimapan. 

PHRYNOSOMA CORNUTUM (Harlan) 

Agama cornuta Harlan, Journ. Acad. Nat. Sci. Philadelphia, vol. 4, 1825, p. 299, 
pi. 20. 

Ta-paya cornuta, Cuvier, Rfegne animal, vol. 2, 1829, p. 37. 

Phrynosoma cornutum, Gray, in Griffith, Cuvier's "Animal Kingdom" (Appen- 
dix), 1831, p. 9.— Smith, Handbook of Hzards, 1946, pp. 290-293, pi. 74. 

Phrynosoma bufonimn Wiegmann, Isis von Oken, vol. 21, 1828, pp. 367-369 (type 
locality uncertain (see Herpetologia Mexicana, 1834, p. 54), here restricted 
to Los Nogales, Sonora; collector unknown; Zool. Mus. Berlin). 

P[hrynosoma] harlanii Wiegmann, Herpetologia Mexicana, 1834, p. 54 (substitute 
name for Agama cornuta Harlan). 

Phrynosoma planiceps Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1852, 
p. 178 (western Texas near the Rio Grande, here restricted to El Paso). 

Phrynosoma cornutum planiceps, Boulenger, Catalogue of the lizards in the 
British Museum, vol. 2, 1885, p. 246. 

Type. — Unknown. 

Type locality. — Great Plains east of the Rocky Mountains, here 
restricted to Fort Riley, Geary County, Kans. 

Range. — Kansas to Texas, west to Arizona, south to Durango and 
San Luis Potosi. Reported from Tamaulipas: Alta Mira, Mier, Soto 
la Marina, Charco Escondido, Alatamoros, Tampico, Marmolejo; 
Nuevo Leon: Monterrey, Pesqueria Grande, 20 miles south of Nuevo 
Laredo, San Juan, 1 mile south of Nuevo Laredo; San Luis Potosi: 
Charco Largo; Coahuila: Castaiiuelos, Alamo de Parras, Monclova, 
San Pedro, Patos, Rio Nazas, El Zacate, Buena Vista, Muzquiz, et 
cetera; Durango: 7 miles south of La Loma, 25 miles north of Berme- 
jillo; Chihuahua: Batopilas, 5 miles north of Moctezuma, Progreso, 
Chihuahua (city), Huajuquilla (Jimenez), Sauz, Guzman, et cetera; 
Sonora: Los Nogales. 

PHRYNOSOMA M'CALLII Hallowell 

Anota m'callii Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 182.] 
Doliosaurus m'calli, Girard, Wilkes United States Exploring Expedition, Herpe- 

tology, 1858, p. 408. 
Phrynosoma m'calli, Stejneger, North Amer. Fauna, No. 7, 1893, p. 190. — Van 

Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 428-430, pi. 

37. 
Anota maccalli,'"^ Bryant, Univ. California Publ. Zool., vol. 9, No. 1, 1911, pp. 5, 

54, pi. 7. 

Type. — Unknown. 

Type locality. — Great Desert of the Colorado, between Vallecita 
and Camp Yuma, about 100 miles east of San Diego. 

Range. — Northwestern Sonora and northeastern Baja California; 
in United States : Southeastern California and southwestern Arizona. 

'« The senior author has expressed the opinion elsewhere that if the characters on which the genus Anota is 
based should prove constant this name should replace Phrynosoma for this species. 



100 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Reported from Sonora: East bank of Colorado River, 5 miles south of 
United States border. Baja California: "Yuhu Basin north of 
Petrified Forest." 

PHRYNOSOMA DOUGLASSH HERNANDESI (Girard) 

Tapaya hernandesi Girard, Wilkes United States Exploring Expedition, Herpe- 

tology, 1858, p. 395. 
Tapaya hernandezii, Baird, United States and Mexican Boundary Survey, vol. 2, 

Rept., 1859, p. 8. 
Phrynosoma hernandesi, Stejneger, North Amer. Fauna, No. 3, 1890, p. 112. 
Phrynosoma douglassii hernandesi, Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 

p. 413, fig. 70. — Van Denburgh, Occas. Pap. California Acad. Sci., No. 10, 

vol. 1, 1922, pp. 382-386, pi. 31. 
Phrynosoma orbiculare hernandesi, Klauber, Bull. Zool. Soc. San Diego, vol. 4, 

1939, pp. 91-93. 
Phrynosoma douglassii Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 302. 

Type.— v. S.N M. No. 197, a cotype. 

Type locality. — New Mexico and "Sonora," restricted to Santa Fe, 
N. Mex. (Cope, loc. cit., 1900). 

Range. — Northern Sonora, Chihuahua; in United States: Colorado, 
southern New Mexico, southeastern and central Arizona, south- 
western Texas. Reported from Sonora: ?"Mexican Boundary line"; 
Chihuahua: Ramos. 

phrynosoma douglassii BRACHYCERCUM Smith 

Phrynosoma douglassii brachycercum Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, 
pp. 361-363. 

Type.—V.S.l<i.M. No. 23993; Edward Palmer collector. 

Type locality. — ^"Durango." 

Range. — Known only from the type locality. 

PHRYNOSOMA BRACONNIERI Doineril and Boconrt 

Phrynosoma Braconnieri Aug. Dum:Sril and Bocourt, Mission scientifique au 
Mexique . . ., fitudes sur les reptiles, livr. 1, 1870, pi. 12, fig. 7, 7a-g. — 
BocouRT, livr. 4, 1874, pp. 233-234. — Boulenger, Catalogue of lizards in 
the British Museum, vol. 2, 1885, p. 248. — Smith, Trans. Kansas Acad. Sci., 
vol. 37, 1934, pp. 287-289. 

Type. — Mus. Hist. Nat. Paris, adult; collected by Aide-Major 
Jacob; several specimens collected by Alfredo Duges and A. Boucard. 

Type locality. — Oaxaca. 

Range. — Extreme southern edge of the central Mexican plateau, in 
semiarid portions of Puebla and Oaxaca (a specimen has been taken 
within a few hundred meters of the border of Veracruz, and very 
probably the species occurs in the mountains m the western part of 
that state). Reported from Puebla: Tehuacan, 14 miles northeast of 
Tehuacan, near Cacoloapam (Km. 226); Oaxaca: No specific locality, 
unless Oaxaca (city) is meant. 



CHECKLIST OF REPTILES OF MEXICO 101 

PHRVNOSOMA MODESTUM Girard 

Phrynosoma modestum Girard, in Stansbury, Exploration and survey of the 

valley of the Great Salt Lake of Utah, 1852, pp. 361-362 and 365, pi. 6, figs. 

4-8. — Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 

430-433, pi. 38. 
Doliosaurus modestus, Girard, Wilkes United States Exploring Expedition, 

Herpetology, 1858, p. 409. 
Anota modesta, Cope, Amer. Nat., vol. 30, 1896, p. 1014, 

Type. — U.S.N.M. (?), several cotypes; General Churcliill and "the 
party under Col. J. D. Graham" collectors. 

Type locality. — The Rio Grande west of San Antonio, Tex. (Churchill), 
and from between San Antonio and El Paso (Graham) ; here restricted 
to Las Cruces, N. Mex. 

Range. — Western Texas to Arizona, southward to San Luis Potosl. 
Reported from Nuevo Leon: Santa Catarina, Pesqueria Grande. 
Chihuahua: 11 miles south of Ahumada, Rio Santa Maria, Guzman, 
near Janos, Juarez, Chihuahua (city); Coahuila: 1.5 miles west of 
Saltillo, Musquiz, Jaral, Castanuelas, Palau, Alamo de Parras, San 
Pedro, Agua Nueva, Monclova, Buena Vista, Rio Nazas, Cuatro 
Cienegas, et cetera; Sonora: Nariz Temporal, Los Nogales, Sierra de 
la Nariz; Durango: between Lerdo and La Goma, Pedriceiia, 5 kilo- 
meters west of Torre6n, 25 miles north of Bermejillo, 7 miles south of 
La Loma; Zacatecas: 3 miles west of La Colorada; San Luis Potosi: 
near Charco Largo (130 kilometers north of San Luis Potosl city). 
Hacienda La Parada. 

phrynosoma PLATYRHINOS PLATYRHINOS Girard 

Phrynosoma platyrhinos Girard, Stansbury's Explorations and survey of the 

valley of the Great Salt Lake of Utah, 1852, pp. 361, 363-364, pi. 7, figs. 1-5.— 

Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 421- 

426, pi. 36. 
Doliosaurus platyrhinos, Girard, in Wilkes, United States Exploring Expedition, 

Herpetology, 1858, p. 409. 
Anota plattjrhina, Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), p. 443, fig. 81. 
Phrynosoma platyrhinos platyrhinos, Klauber, Bull. Zool. Soc. San Diego, vol. 14, 

1939, p. 13. 
Anota calidiarum, Cope, Amer. Nat., vol. 30, No. 358, 1896, p. 333 (Death Valley, 

Calif.). 
Phrynosoma calidiarum, Ditmars, The reptile book, 1907, p. 157, pi. 46, figs. 10, 

14, pi. 50, fig. 2. 

Type.— V.S.N. M. No. 189?; H. Stansbury collector. 

Type locality. — Great Salt Lake, Utah. 

Range. — Northeastern Baja California; widespread in Nevada, 
Idaho, Utah, Washington, California, and Arizona. Reported in Baja 
California: 20 miles north of San Felipe, San Felipe Bay, Pozo, San 
Agustin, Seven Wells on the Salton River. 



102 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

PHRYNOSOMA PLATYRHINOS GOODEI Stejneger 

Phrynosoma goodei Stejneger, North Amer. Fauna, No. 7, 1893, p. 191 (foot- 
note), pi. 2, figs. 3a-3c. 
Anota goodei, Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), p. 442, fig. 80. 
Phrynosoma platyrhinos goodei, Klauber, Copeia, 1935, pp. 178-179. 

Type.— U.S.N. M. No. 8567a; T. H. Streets collector. 

Type locality. — "Coast deserts of the state of Sonora, Mexico," here 
restricted to Puerto Libertad. 

Bange. — Known only from the gulf coast of Sonora. Reported 
from Puerto Libertad, 8 miles northwest of Alamo Muerto, west 
coast of Sonora. 

PHRYNOSOMA ASIO Cope 

Phrynosoma asio Cope, Proc. Acad. Nat. Sci. Philadelphia, vol. 16, 1864, pp. 

178-179. — Boulenger, Catalogue of lizards in the British Museum, vol. 2, 

1885, p. 244. 
Batrachosoma asio, Bocourt, Mission seientifique au Mexique • . . , Etudes 

sur les reptiles, livr. 4, 1874, pi. 17, fig. 9, 9a-c. 
Phrynosoma spinimentum Peters, Monatsb. Akad. Wiss. Berlin, 1873, pp. 742- 

743 (Tehuantepec; Mus. Berlin No. 7219). 

Ti/pe.— U.S.N.M. (?) ; John Xantus collector. 

Type locality. — Colima, Colima (by present restriction). 

Range.- — From Colima tlirough coastal Michoacan, Guerrero, Oaxaca 
to Chiapas, and in the Balsas Basin. In Central America, recorded 
from Guatemala. Reported from Oaxaca: San Pedro, TequesLxtlan, 
San Gerdnimo, Ranchero Poso Rio, vicinity of Tehuantepec, vicinity 
of Salina Cruz; Guerrero: Mexcala (on Balsas River); Michoacan: 
La Salad a, Cofradia, San Salvador, Apatzingan; Colima: Tecoman, 
Colima (city); Chiapas: no definite locality. 

PHRYNOSOMA CORONATUM CORONATUM (Blainville) 

Agama {Phrynosoma) coronata Blainville, Nouv. Ann. Mus. Hist. Nat. Paris, 

vol. 4, 1835, p. 284, pi. 25, figs. 1-lc. 
Phrynosoma coronatum coronaiiim, Linsdale, Univ. California Publ. Zool., vol. 

38, 1932, p. 368.— Klauber, Copeia, 1936, pp. 103-110, fig. 1.— Tevis, 

Copeia, 1944, pp. 13-15, fig. 2, map. 

Type.— Mus. Hist. Nat. Paris; P. E. Botta collector. 
Type locality. — "California," here restricted to Cape San Lucas, 
Baja California. 

Range. —Southern part of Baja California, north to about lat. 26° N. 

PHRYNOSOMA CORONATUM JAMESI Schmidt 

Phrynosoma jamesi Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 

668-669, pis. 65, 56. 
Phrynosoma coronatum jamesi, Klauber, Copeia, 1936, p. 110. — Tevis, Copeia, 

1944, pp. 13-15, map fig. 2. 

Type.— U.S. l^.M. No. 64450; H. Townsend collector. 

Type locality. — Shore of San Bartolome Bay, Baja California. 



CHECKLIST OF REPTILES OF MEXICO 103 

Range. — Central Baja California, between lat. 26° N. and lat. 
28°30' N. 

PHRYNOSOMA CORONATUM FRONTALE Van Denburgh 

Phrynosoma frontalis Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 4, 

1S94, pp. 296-298. 
Phrynosoma blainvillii frontale, Van Denburgh, Occ. Pap. California Acad. Sci., 

No. 5, 1897, p. 95; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 

395-401, pi. 33.— Klauber, Copeia, 1936, pp. 103-110. 
Phrynosoma cor onatum frontale, Linsdale, Univ. California Publ. Zool., vol. 38, 

1932, p. 368.— Tevis, Copeia, 1944, pp. 13-15, fig. 2 (map). 
Phrynosoma schmidti Barbour, Proc. New England Zool. Club, vol. 7, 1921, 

p. 113 (Mus. Comp. Zool. No. 15142; W. W. Brown collector; Cedros Is- 
land) . 
Phrynosoma nelsoni Schmidt, Bull. American Mus. Nat. Hist., vol. 46, 1922, 

p. 666 (Amer. Mus. Nat. Hist. No. 37585; E. W. Nelson and E. A. Goldman 

collectors; San Quintfn, Baja California). 

Ti/pe.— Stanford Univ. Mus. No. 93, adult male; C. H. Gilbert 
and W. W. Price collectors. 

Type locality. — Bear Valley, San Benito County, Calif. 

Range. — Discontinuous; central California east of the Sierra Nevada 
south to but not including extreme southwestern California; north- 
western coast of Baja California except a smaU area adjacent to the 
United States border; Cedros Island. 

PHRYNOSOMA CORONATUM BLAINVILLII Gray 

Phrynosoma blainvillii Gray, The zoology of Captain Beechey's voyage . . . , 

Reptiles, 1839, p. 96, pi. 29, fig. 1. 
Phrynosoma blainvillii blainvillii, Grinnell and Camp, Univ. California Publ. 

Zool., vol. 17, 1917, p. 164. — Van Denburgh, Occ. Pap. California Acad. 

Sci., No. 10, 1922, pp. 388-395, pi. 32.— Klauber, Copeia, 1936, pp. 103-110, 

fig. 2. 
Phrynosoma ochoterenai Terron, Anal. Inst. Biol., vol. 3, 1932, p. 109 (type 

formerly in Mus. Hist. Nat., Mexico, now lost; J. M. Gallegos collector; 

Tecate, Baja California). 

Type. — Brit. Mus. Nat. Hist., young specimen; de Blainville 
collector. 

Type locality. — "California," here restricted to San Diego. 

Range. — Southern California and extreme northeastern Baja 
California. 

PHRYNOSOMA CERROENSE Stejneger 

Phrynosoma cerroense Stejneger, North Amer. Fauna, No. 7, 1893, p. 187 (in 
footnote). — Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, 
pp. 401-403.— Klauber, Copeia, 1936, p. 108. 

Type.— V.S.N. M. No. 11977; L. Belding collector. 
Type locality. — ''Cerros" Island, Pacific coast of Baja California 
[= Cedros Island]. 

Range. — Known only from the type locality. 



104 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

PHRYNOSOMA TAURUS DugSs 

Phrynosoma laurus Duces, MS. 1868, fide Dumdril and Bocourt loc. cit., p. 234; 
and La Naturaleza, 1869, p. 143 (nomen nudum). — Aug. Dumeril and 
Bocourt, Mission scientifique au Mexique . . . , Etudes sur les reptiles, 
livr. 1, 1870, pi. 12, figs. 8, 8a-g.— Duofes, La Naturaleza, 1873, pp. 302-305, 
figs. 1-4 (this description bears the date Nov. 23, 1870, but was not pub- 
lished until 1873; there were two specimens, one from "CozcatMn," the other 
from Estado de Puebla). — Bocourt, Mission scientifique au Mexique . . . , 
fitudes sur les reptiles, livr. 4, 1874, pp. 234-235. — Boulenger, Catalogue 
of lizards in the British Museum, vol. 2, 1885, p. 249. — Smith and Necker, 
Anal. Esc. Nac. Cienc. Biol., vol. 3, 1943, pp. 208-210, pi. 3, figs. 1-2. 

Type.^'— Mus. Hist. Nat. Paris, 9 cotypes?; M. Boucard collector. 

Type locality. — "Matamoras Izucar," Puebla. 

Range, — Puebla and Guerrero. Reported from Puebla: Zapotitlan, 
San Diego near Tebuacan, Cozcatlan (Dr. Bedel collector), Izucar 
de Matamoros; Guerrero: 5 miles north of Chilpancingo, Amula 
8,000 feet. 

PHRYNOSOMA SOLARE Gray 

Phrynosoma Solaris Gray, Catalogue of the specimens of lizards in the collection 

of the British Museum, 1845, p. 229. 
Phrynosoma solare, Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 5, 

1895, p. 115; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 406-409, pi. 34. 
Phrynosoma regale Girard, in Wilkes, United States Exploring Expedition, 1858, 

p. 406 (U. S. N. M. No. 161; Sierra de la Nariz, near Zufii, Sonora). 

Type. — Brit. Mus. Nat. Hist, (not listed in Boulenger, Catalogue of 
lizards in the British Museum, vol. 2, unless it is spec, a, cf [Cali- 
fornia!, listed under Phrynosoma regale, p. 245); ? de Blainville col- 
lector. 

Type locality. — "California," here restricted to Yuma, Ariz. 

Range. — Southern Arizona southward into northeastern Baja 
California, western Sonora and into Sinaloa. Reported from Sonora: 
Santa Ana, Nogales, Hermosillo, 15 to 20 miles south and 45 miles 
southwest of Hermosillo, Tiburon Island, Empalme, 10 miles north- 
west of Guaymas, Alamos, Guirocoba; Baja California: Las Animas 
Bay; Sinaloa: Ahome, Sierra de Choix. 

PHRYNOSOMA DITMARSI Stejneger 

Phrynosoma diimarsi Stejneger, Proc. U. S. Nat. Mus., vol. 29, 1906, p. 565. — 
DiTMARB, The reptile book, 1907, p. 154, pi. 46, figs. 3, 7, pi. 48, fig. 1.— 
Smith, Handbook of lizards, 1946, pp. 297-299, pi. 97. 

Tii/^e.— U.S.N.M. No. 36022; Mr. Eustace collector. 



" Whether the specimens mentioned above are really the types may be open to question. A description 
in manuscript was made by Duges from a specimen in Mexico and presumably sent from Mexico to August 
Dumfiril in Paris. The Paris Museum had nine specimens of the same species. Drawings were made of 
one, and the drawings published under the name Phrynosoma taurus Duges. Since the specimen figured 
is the first description or indication, we believe that the original of the drawing becomes the type and this, 
we believe, is a specimen in the Museum d'Histoire Naturelle de Paris. Thus the type specimen was 
presumably never seen by the author of the name. The specimen described by Duges is in the Museum 
"Alfredo Duges," Colegio del Estado de Guanajuato. 



CHECKLIST OF REPTILES OF MEXICO 105 

Type locality. — State of Sonora, not far from the boundary of 
Arizona. 

Range. — Known only from the type locality. 

Genus SCELOPORUS Wiegmann 

Sceloporus Wiegmann, Isis von Oken, 1828, p. 369. 

Tropidolepis Cuvier, Rfigne animal . . . , ed. 1, vol. 2, 1817, p. 38 (type, Agama 

undulata Daudin). 
Lysoptychus Cope, Proc. U. S. Nat. Mus., vol. 11, 1888, pp. 397-398 (type, L. 

lateralis Cope = Sceloporus couchii Baird). 

Genotype. — Sceloporus torquatus Wiegmann. 

Species.— In all, 104 forms are recognized at present, consisting of 
54 species representing 15 groups. Eighty-nine forms of 52 species 
are known from Mexico. 

Range.— Southern Canada on the west and southern New "York on 
the east south throughout North and Central America to western 
Panama. 

KEY TO MEXICAN GROUPS OF SCELOPORUS 

1. Postfemoral dermal pocket present 2 

Postfemoral dermal pocket absent 4 

2. Postrostral scales absent; nasals and internasals in contact with rostral. 

maculosus (p. 133) 
Postrostral scales present; internasals and nasals separated from rostral. _ 3 

3. Tail strongly compressed in males, rounded and light pink in females; femoral 

pores 24 or more on each side pyrocephalus (p. 138) 

Tail rounded in both sexes, not pink in females; femoral pores 20 or less. 

variabilis (p. 129) 

4. Lateral body scales not imbricate 5 

Lateral body scales imbricate 6 

5. Preanal scales keeled in females; males with poorly developed postanals; no 

distinct belly patches in males; tail over twice length of body. 

utiformis (p. 135) 
Preanal scales smooth in females; males with well-developed postanals; 
lateral belly patches distinct in males; tail less than twice length of body. 

merriami (p. 132) 

6. No postrostrals; nasals and internasals in contact with rostral 7 

Postrostrals present; nasal and internasals separated from rostral 8 

7. Dorsal scales 50 or more; femoral pores 17 or more; preanals smooth in 

females scalaris (p. 136) 

Dorsal scales 46 or less; femoral pores 16 or less; preanals keeled in females. 

siniferus (p. 133) 

8. Ventral scales, at least laterally, pointed, not notched; preanal scales keeled 

in females; no lateral belly markings in males; femoral pore series widely 

separated medially; two postrostrals siniferus (p. 133) 

Ventral scales notched, or, if pointed, femoral pore series closely approxi- 
mated medially (separated by four scales or less); preanal scales smooth 
in females 9 

9. Males uniform white below; anterior section of frontal longitudinally divided; 

femoral pore series widely separated medially chrysostictus (p. 133) 

Males with lateral belly patches distinct or, if not, anterior section of frontal 
entire 10 



106 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

10. Femoral pore series closely approximated medially; two postrostrals; lateral 

scale rows parallel or nearly so; scales on posterior surface of thigh granular; 

small species scalaris (p. 136) 

Femoral pore series widely separated medially, or, if closely approximated, 
scales on posterior surface of thigh not granular; lateral scale rows strongly 
divergent; typically four postrostrals, sometimes three, rarely two 11 

11. Scales on posterior surface of thigh granular 12 

Scales on posterior surface of thigh not granular 15 

12. Dorsal scales highly irregular in size grammicus (p. 119) 

Dorsal scales subequal in size 13 

13. Lateral nuchal scales much smaller than and well differentiated from dorsal 

nuchal scales; scales of lateral row of dorsal nuchals enlarged, strongly 

keeled and mucronate grammicus (p. 119) 

Lateral nuchal scales not well differentiated from dorsal nuchal scales 14 

14. Throat light blue, barred or mottled with white; lateral belly patches confluent 

medially; preanal scales large, about seven in a row from a line between 

femoral pore series to anus grammicus (p. 119) 

Throat not as described or, if so, lateral belly patches not confluent medially 
and preanal scales smaller 15 

15. Scales on posterior surface of thigh granular, femoral pore series widely 

separated medially, the folds defining the preanal area passing between the 
femoral pore series; throat mottled with blue and white, never with a 
distinct, blue spot medially or on either side posteriorly, graciosus (p. 118) 
Scales on posterior siu-face of thigh not granular or, if so, femoral pore 
series more closely approximated medially, the folds defining preanal area 

not or barely passing between femoral pore series 16 

16; Dorsal, ventral, and lateral scales subequal in size; small species, 

megalepidurus (p. 121) 

Dorsal, ventral, and lateral scales distinctly differing in size; large or small 

species 17 

17. A distinct, dark, light-bordered nuchal collar torquatus (p. 121) 

No distinct, dark, light-bordered nuchal collar 18 

18. Gular region mottled, not barred nor with a median or two lateral blue spots 

posteriorly torquatus (p. 121) 

Gular region not mottled 19 

19. Gular region barred in males and lacking a median posterior dark blue or 

black spot spinosus (p. 110) 

Gular region not barred; or, if barred, with a median posterior dark blue or 
black spot 20 

20. Gular region barred in males, with a median posterior dark blue or black spot. 

undulatus (p. 117) 
Gular region not barred 21 

21. Supraoculars large, separated from superciliaries by no more than one com- 

plete and one incomplete row of small scales; and one or more of the posterior 

supraoculars in contact with median head scales 28 

Supraoculars large or small; if any of the posterior supraoculars are in con- 
tact with median head scales, the supraoculars are separated from super- 
ciliaries by three or more complete or incomplete rows of small scales 22 

22. Males lacking lateral belly patches vmdulatus (p. 117) 

Males with lateral belly patches 23 

23. A pair of dark blue or black spots on posterior part of gular region. 

undulatus (p. 117) 
No pair of dark spots on posterior part of gular region 24 



CHECKLIST OF REPTILES OF MEXICO 107 

24. Three or more rows of small scales between supraoculars and superciliaries; 

or, if two, the scales of inner row considerably larger than those of outer 

row 25 

One complete and one incomplete row of small, subequal scales between 
supraoculars and superciliaries 27 

25. One canthal, or, if two, the first forced above canthal ridge_form.osus (p. 107) 
Two canthals, the first not forced above canthal ridge 26 

26. A broad, black nuchal collar complete about neck dorsally, or anterior section 

of frontal longitudinally divided formosus (p. 107) 

Neither true undulatus (p. 117) 

27. Frontal ridges present forniosus (p. 107) 

Frontal ridges absent spinosus (p. 110) 

28. Adult males a uniform bright green above formosus (p. 107) 

Not so spinosus (p. 110) 

FORMOSUS GROUP 

Species. — Six, with a total of 11 forms. Extralimital are lunaei, m. 
malachiticus, and m. smaragdinus .'^ 

Range. — Alichoacan and Veracruz south to western Panama. 

KEY TO MEXICAN SPECIES OF THE FORMOSUS GROUP OF SCELOPORUS 

1. Median frontonasal separated from lateral frontonasals 2 

Median frontonasal in contact with lateral frontonasals 4 

2. Dorsals about 31; internasals large, keeled, 3 from rostral to median fron- 

tonasal prezygus '^a (p. 109) 

Dorsals 37 to 45 ; internasals smaller, not keeled, rugose or not 3 

3. A complete nuchal collar, sometimes narrowly interrupted medially. 

malachiticus salvini (p. 108) 
Nuchal collar incomplete malachiticus taeniocnemis (p. 107) 

4. Anterior section of frontal usually longitudinally divided; canthals sharply 

ridged; no nuchal collar; ventrals a fourth, laterals not more than two-thirds, 

size of dorsals asper (p. 109) 

Anterior section of frontal rarely longitudinally divided; canthals rounded; 
a nuchal collar or not; ventrals a half, laterals three-fourths size of dorsals. 5 

5. A broad, nearly or quite complete, nuchal collar 6 

Collar, if present, restricted to sides of neck 7 

6. Femoral pores 12-16; one or more supraoculars generally in contact with 

median head scales malachiticus acanthinus (p. 108) 

Femoral pores 16-21; supraoculars not in contact with median head scales. 

stejnegeri (p. 109) 

7. Males without yellow on throat; dorsal scale rows black edged; dorsal head 

scales with a light blue center formosus scitulus (p. 108) 

Males with yellow or orange on throat; dorsal surface uniform blue; head 
scales not light spotted formosus formosus (p. 108) 

SCELOPORUS MALACHITICUS TAENIOCNEMIS Cope 

Sceloporus taeniocnemis Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, p. 399. 
Sceloporus malachiticus taeniocnemis, Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, 
p. 356; Journ. Washington Acad. Sci., vol. 39, 1949, pp. 39, 40. 

" See Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. SSC), for a key to these forms. 
'2» Material examined since this key was originally prepared places this species in the torquatus group. 
861316 — 50 8 



108 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Type — U.S.N.M. No. 24768. 

Type locality. — Guatemala, here restricted to Coban. 

Range. — Highlands of Chiapas and northern Guatemala in Alta 
Verapaz, Known in Alexico only from Chiapas: Pinobete, Mount 
Ovando, Teopisca. 

SCELOPORUS MALACHITICUS SALVINI Gunther 

Sceloporus salvini Gather, Biologia Centrali-Americana, Reptilia and Batra- 
chia, 1890, p. 68. — Smith, Proc, Biol. Soc. Washington, vol. 49, 1936, p. 95; 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 55-58, pi. 3. 

Sceloporus malachiticus salvinii, Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 
356; Journ. Washington Acad. Sci., vol. 39, 1949, p. 39. 

Type. — Brit. Mus. Nat. Hist, cotypes: Jalapa, Hoege collector; 
Guatemala, Salvin collector. Lectotype specimen in Boulenger, 
Proc. Zool. Soc. London, 1897, p. 503. 

Type locality. — "Jalapa and Guatemala." Restricted (Smith, 
1939, p. 40) to Jalapa, Veracruz. 

Range. — Central western Veracruz to the Isthmus of Tehuantepec. 
Recorded only from the states of Veracruz and Oaxaca. 

SCELOPORUS MALACHITICUS ACANTHINUS Bocourt 

Sceloporus acanthinus Bocourt, Ann. Sci. Nat. Zool., ser. 5, vol. 17, No. 6, 1873, 
p. [24] (acathhinus) ; No. 10, 1873, p. 1; Mission scientifique au Mexique 
. . . , Etudes sur les reptiles, livr. 3, 1874, pp. 180-181, pi. 18, figs. 10, 10a, 
10b, pi. 19, figs. 4, 4a. — Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 74-78, pis. 6, 7. 

Sceloporus malachiticus acanthinus, Smith, Journ. Washington Acad. Sci., vol. 39, 
1949, pp. 39-40. 

Sceloporus guentheri Stejneger, Proc. Biol. Soc. Washington, vol. 31, 1918, p. 92 
(type, Brit. Mus. Nat. Hist.; "Mexico," here restricted to La Esperanza, 
Chiapas). 

Type. — Mus. Hist. Nat. Paris, male and female cotypes; Commis- 
sion Scientifique collector. 

Type locality. — San Agustin, near Volcan de Atitlan, Guatemala, 
610 meters. 

Range. — Pacific coast foothills, Chiapas to El Salvador. Known 
in Mexico only from the vicinity of Escuintla, Chiapas. 

SCELOPORUS FORMOSUS FORMOSUS Wiegmann 

Sceloporus formosus Wiegmann, Herpetologia Mexicana, 1834, p. 50, pi. 7. 
Tropidolepis formosus, Dumeril and Bibron, Erpetologie g6n6rale, vol. 4, 1837, 

pp. 303-304. 
Sceloporus formosus formosus, Wettstein, Sitzb. Akad. Wiss. Wien, math. nat. 

Kl., vol. 143, Abt. 1, 1934, p. 25.— Smith, Publ. Field Mus. Nat. Hist., zool. 

ser., vol. 26, 1939, pp. 34-41 (part). 
Sceloporus viviparus Cope, Proc. Amer. Philos Soc, vol. 22, 1885, p. 398 (Mirador, 

Veracruz; type, U.S.N. M. No. 25073). 



CHECKLIST OF REPTILES OF MEXICO 109 

Type. — ? Zool. Mus, Berlin; F. Deppe collector. 

Type locality. — Mexico, here restricted to Acultzingo, Veracruz. 

Range. — Central Veracruz south in mountainous regions to the 
Isthmus of Tehuantepec. Recorded from various localities in the 
states of Veracruz, Puebla, and Oaxaca. 

SCELOPORUS FORMOSUS SCITULUS Smith 

Sceloporus formosus scitulus Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 
352-354. 

Type.— EBT-HMS No. 26962; R. C. Taylor and E. H. Taylor 
collectors. 

Type locality. — Omilteme, Guerrero. 

Range. — Sierra Madre del Sur Mountains in Guerrero. 

SCELOPORUS ASPER Boulenger 

Sceloporus asper Boulenger, Proc. Zool. Soc. London, 1897, pp. 476, 497, 498, 
pi. 33 (colored).— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 51-55. 

Sceloporus obscurus Van Denburgh, Proc. Acad. Nat. Sci. Philadelphia, 1897 
(1898), p. 462 (Tepic, Nayarit; type, California Acad. Sci. No. 3213). 

Type. — Brit. Mus. Nat. Hist. 

Type locality. — La Cumbre de los Arrastrados, Jalisco. 

Range. — Mountainous regions of western Mexico, from Nayarit to 
Guerrero. Recorded from Nayarit, Jalisco, Michoacan, and Guer- 
rero.'^ 

SCELOPORUS STEJNEGERI Smith 

Sceloporus stejnegeri Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 350-352. 

Type.—V.S.lSi.M. No. 112635; E. H. Taylor, R. C. Taylor, and 
H. M. Smith collectors. 

Type locality. — Tierra Colorada, Guerrero. 

Range. — Guerrero, on southern low slopes of the Sierra Madre del 
Sur. 

SCELOPORUS PREZYGUS Smith '* 

Sceloporus prezygus Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 354-355. 

Type.— U.S.N.M. No. 46881; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — Conjab, 5,300 feet (between San Bartolom6 and 
Comitan), Chiapas. 

Range. — Known only from the type locality in Chiapas, 



'» The record from this state (Chilapa) (Ahl, Zool. Anz., vol. 106, 1934, p. 184) is open to doubt. The iden- 
tifieation needs verification. 

'* Group allocation of this species was not certain until after this account was in type. It belongs in the 
torguatus group. 



110 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

SPINOSUS GROUP 

Species. — Nine, with a total of 21 forms. 

Range. — Guatemala northward to southern Nevada, southwestern 
Colorado, southern Oklahoma, and central Texas. 

KEY TO THE SPINOSUS GROUP OF SCELOPORUS 

1. Femoral pores on both thighs total 7 or more 3 

Femoral pores on both thighs total 6 or less 2 

2. Ventral surfaces immaculate in adults of both sexes. 

liorridus albiventris (p. 116) 

Sides of belly blue, frequently dark-bordered, in males; gular region barred 

in males, sometimes in females horridus oligoporus (p. 116) 

3. Supraoculars completely in contact with superciliaries, or with one to four 

very small scales between; posterior superciliary greatly enlarged; venter 

immaculate in both sexes edwardtaylori (p. 112) 

Supraoculars separated from superciliaries by at least one row of small scales; 
posterior superciliary not enlarged; venter with distinct markings in 
males 4 

4. Femoral pores on both thighs total 12 or less; 4 supraoculars, the fourth in 

contact with median head scales horridus horridus (p. 116) 

Femoral pores on both thighs total 13 or more; or, if 12, all supraoculars 

separated from median head scales 5 

6. Femoral pores (one side) 17 or more 6 

Femoral pores (one side) 16 or less 10 

6. A black shoulder patch present, with a hght posterior border 10 

No black shoulder patch; or, if present, without a light posterior border. . 7 

7. Snout black; a broad, black bar across head at middle of supraocular region; 

chest and middle of belly orange in males; gular region in females white, 
with irregular, black marks; first canthal not in contact with lorilabials; 

ventral scales 44-54 8 

Snout not black, similar in color to rest of head; no black bar across head; 
chest and middle of belly not orange; gular region in both sexes bluish, with 
light lines following scale rows at anterior part of throat; first canthal in 
contact with lorilabials; ventral scales 38-42..inagister lineatulus (p. 115) 

8. Femoral pores usually (84 percent) 20 or less on each side; total pore counts 

usually (80 percent) 40 or less 9 

Femoral pores usually (78 percent) 21 or more on each side; total pore counts 
usually (83 percent) 41 or more; lateral belly patches usually not confluent 
medially in males melanorhinus melanorhinus (p. 112) 

9. Lateral nuchal pocket greatly reduced, with very few if any granules, and very 

little if any bare skin, shallow melanorhinus stuarti (p. 113) 

Lateral nuchal pocket with numerous granules, considerable bare skin, and 
considerable depth melanorhinus calligaster (p. 113) 

10. One or more supraoculars in contact with median head scales 14 

None of supraoculars in contact with median head scales 11 

11. Gular region barred 12 

Gular region not barred 13 

12. Dorsal scales usually more than 30; femoral pores usually more than 9 (one 

side); supraoculars usually 5 to 7 spinosus caeruleopunctatus (p. 116) 

Dorsal scales usually 30 or less; femoral pores usually 9 or less (one side); 
supraoculars usually 4 spinosus spinosus (p. 116) 



CHECKLIST OF REPTILES OF MEXICO 111 

13. Belly patches in males continuous across chest; prefrontals never in contact; no 

dorsolateral light lines in males; posterior surface of thigh mottled or with 

a broad light line, dark-bordered 14 

Belly patches small, confined to sides of belly in males; prefrontals usually in 
contact; dorsolateral light lines present in males; posterior surface of thigh 
nearly immaculate or with a narrow, short black line near insertion of hind 
leg olivaceus (p. 117) 

14. Gular region barred 15 

Gular region not barred 16 

15. Femoral pores 13 or more (one side) orcutti licki (p. 114) 

Femoral pores 12 or less (one side) spinosus spinosus (p. 116) 

16. Femoral pores (one side) 12 or less 17 

Femoral pores (one side) 13 or more 21 

17. Outer row of labiomental scales in contact with mental. 

magister magister (p. 114) 
Outer row of labiomental scales separated from mental 18 

18. First canthal in contact with lorilabials orcutti orcutti (p. 114) 

First canthal separated from lorilabials 19 

19. Lateral scales about one-half size of dorsal scales; median frontonasal not in 

contact with frontal; lateral belly patches in males not passing over 

chest 21 

Lateral scales considerably more than one-half size of dorsal scales; median 
frontonasal usually in contact with frontal; lateral belly patches in males 
passing over chest 20 

20. Dorsal markings absent or very dimly visible; a broad, light line on posterior 

surface of thighs lundelli gaigeae (p. 112) 

Dorsal markings present and distinct at least in females, consisting of narrow, 
dark cross bars; posterior surface of thigh mottled. 

lundelli lundelli (p. 112) 

21. Femoral pores (one side) 10 or less 22 

Femoral pores (one side) 1 1 or more 23 

22. First canthal rarely in contact with lorilabials; gular region never barred; 

outer row of labiomentals rarely in contact with mental. 

clarkii boulengeri (p. 113) 
First canthal usually in contact with lorilabials; gular region barred, or outer 
row of labiomentals sometimes in contact with mental (not in o. orcut- 
ti) 23 

23. First canthal separated from labiomentals; femoral pores 16 or less; lower fore- 

legs distinctly barred; gular region never barred clarkii clarkii (p. 113) 

First canthal in contact with lorilabials, lower forelegs not barred dis- 
tinctly 24 

24. Outer row of labiomentals in contact with mental; gular region not barred; 

shoulder patch distinct 26 

Outer row of labiomental scales separated from mental; gular region barred or 
shoulder patch indistinct 25 

25. Dorsal scales 32 to 40; femoral pores 13-19; usually one or more scales in con- 

tact with both subocular and supralabials; a distinct, light-bordered black 
shoulder patch; dorsolateral light lines present, indistinct posteriorly, the 

medial edge sometimes indistinct orcutti licki (p. 114) 

Dorsal scales 28 to 36; femoral pores 10-15; usually 2 complete rows of lorila- 
bials below subocular; shoulder patch indistinct; no dorsolateral light 
lines orcutti orcutti (p. 114) 



112 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

26. Supraoculars usually 5; femoral pores (one side) usually 15 or less. 

magister magister (p. 114) 
Supraoculars usually 6 or 7; femoral pores usually 16 or more 27 

27. No dark lines in adult males on sides of lateral scale rows; femoral pores (one 

side) usually less than 19; lateral belly patches confluent medially in adult 

males magister rufidorsum (p. 115) 

Dark lines on sides of lateral scale rows present in adult males; lateral belly 
patches not confluent mediallj' in adult males 28 

28. A distinct, narrow dorsal stripe about l}i scale rows wide; females and young 

with 2 rows of dark spots, one on each side of middorsal stripe. 

magister monserratensis (p. 115) 

A broad, light-colored dorsal area about 6 scale rows wide; dark spots on back 

absent or indistinct magister zosteromus (p. 115) 

SCELOPORUS LUNDELLI LUNDELU Smith 

Sceloporvs lundelli lundelli Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 66-71, pi. 4. 

Type.-AJmv. Michigan Mus. Zool. No. 80674; C. L. Lundell 
collector. 

Type locality. — Cohune Ridge (20 miles southeast of Benque Viejo), 
British Honduras. 

Range. — The southern portion of the Yucatan Peninsula. Recorded 
in Mexico only from Campeche. 

SCELOPORUS LUNDELU GAIGEAE Smith 

Sceloporus lundelli gaigeae Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 71-74, pi. 5. 

Ty^e.— Chicago Nat. Hist. Mus. No. 31524; H. M. Smith collector. 

Type locality. — Merida, Yucatan. 

Range. — Northern Yucatan; recorded only from Yucatan. 

SCELOPORUS EDWARDTAYLORI Smith 

Sceloporus edwardtaylori Smith, Herpetologica, vol. 1, 1936, pp. 6-8; Publ. Field 
Mus. Nat. Hist., zool. sen, vol. 26, 1939, pp. 78-82, fig. 9. 

Type.— EHT-HMS No. 8331 (EHT field No. 4221); E. H. Taylor 
and H. M. Smith collectors. 

Type locality. — Ixtepec (= San Ger6mmo), Oaxaca. 
Range. — Central and southern Oaxaca. 

SCELOPORUS MELANORHINUS MELANORHINUS Boconrt 

Sceloporus melanorhinus Bocourt, Ann. Sci. Nat., Zool., ser. 6, vol. 3, No. 12, 
1876, pp. 2-4.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 82-87 (part). 

Sceloporus melanorhinus melanorhinus, Smith, Proc. U. S. Nat. Mus., vol. 92, 
1942, p. 360. 

Type. — Mus. Hist. Nat. Paris, three females, one male, cotypes; 
F. Sumichrast collector. 



CHECKLIST OF REPTILES OF MEXICO 113 

Type locality. — Isthmus of Tehuantepec, Oaxaca, here restricted 
to Tehuantepec (City). 

Range. — Pacific slopes of Oaxaca. 

SCELOPORUS MELANORHINUS CALLIGASTER Smith 

Sceloporus melanorhinus, Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 

1939, pp. 82-87 (part), pis. 9, 10. 
Sceloporus melanorhinus calligaster Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, 

pp. 360-361. 

2>2?e.— U.S.N.M. No. 112201; H. M. Smith collector. 

Type locality. — Acapulco, Guerrero, 

Range. — Pacific slopes below about 3,500 feet, from Nayarit to 
Guerrero. Recorded from Nayarit, Jalisco, Colima, Michoacan, and 
Guerrero. 

SCELOPORUS MELANORHINUS STUARTI Smith 

Sceloporus melanorhinus calligaster, Smith, Rev. Soc. Mex. Hist. Nat., vol. 7, 

1947, p. 66. 
Sceloporus melanorhinus sluarti Smith, Nat. Hist. Misc., No. 20, 1948, pp. 1-3. 

Type. — Univ. Michigan Mus. Zool. No. 96759; L. C. Stuart collector. 

Type locality. — Finca Canibal, 3,000 feet, Huehuetenango, Guate- 
mala. 

Range. — The upper valley of the Rio Grijalva in Chiapas and Guate- 
mala. Known in Mexico only from Chiapas: Piedra Parada, 12 
miles north of Ocozucoautla. 

SCELOPORUS CLARKII CLARKH Baird and Girard 

Sceloporus clarkii Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 

1852, p. 127 (part). — Stejneger, North Amer. Fauna, No. 7, 1893, pp. 

178-181, figs, la-c, pi. 1. — Smith, Handbook of lizards of the United States, 

1945, pp. 206-208, pi. 45. 
Sceloporus clarkii clarkii, Cope, U, S. Nat. Mus. Bull. 1, 1875, pp. 49, 92 (part). — 

Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 118-128. 

Type. — U.S.N.M. No. 2940, three cotypes, of which two belong to 
S. m. magister, and one, with a white tag on its leg, is designated 
lectotype of S. clarkii; John H. Clark collector. 

Type locality. — "Province of Sonora" (= southern Arizona), here 
restricted to Santa Rita Mountains. 

Range. — Southern and central Arizona (except the western part of 
the State), southwestern New Mexico, and all of Sonora except the 
extreme western and southern parts. Recorded only from the states 
of Sonora and Chihuahua, and the islands of San Pedro Nolasco and 
Tiburdn. 

SCELOPORUS CLARKII BOULENGERI Stejneger 

Sceloporus boulengeri Stejneger, North Amer. Fauna, No. 7, 1893, p. 180, pi. 1> 

figs. 5a-c. 
Sceloporus clarkii boulengeri, Burt, Trans. Amer. Micr. Soc, vol. 54, 1935, 171-172 

(part). — Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 

128-133, pi. 15. 



114 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type. — U.S.N.M. No. 14079, three cotypes; A. Forrer collector. 

Type locality. — Presidio (about 50 miles south from Mazatlan), 
Sinaloa. 

Bange. — Pacific slopes, southern Sonora south to northern Jalisco. 
Recorded from Sonora, Sinaloa, Nayarit, and Jalisco. 

SCELOPORUS ORCUTTI ORCUTTI Stejneger 

Sceloporus orcutti Stejneger, North Amer. Fauna, No. 7, 1893, p. 181, pi. 1, 

figs. 4a-c. 
Sceloporus orcutti orcutti, Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 

1939, pp. 133-140, fig. 12. 
Sceloporus digueti Mocquard, Nouv. Arch. Mus. Hist. Nat. Paris, ser. 4, vol. 1, 

1899, pp. 311-313, pi. 12, fig. 2-2b (Mus. Hist. Nat. Paris; Santa Rosalia, 

Baja California). 

T2/pe.— U.S.N.M. No. 16330; C. R. Orcutt collector. 

Type locality. — Milquatay Valley, San Diego County, Calif. 

Bange. — California, from southern San Bernardino county south- 
ward into Baja California to the Sierra de la Gigante, and on some 
adjacent islands. 

SCELOPORUS ORCUTTI LICKI Van Denburgh 

Sceloporus lichi Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 5, 1895, 

pp. 79, 110-114, pi. 10. 
Sceloporus orcutti licki, Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 

1939, pp. 140-145, pi. 4. 

Type. — Stanford Univ. Mus. No. 2987a, lectotype (one of two 
paratypes; type in California Acad. Sci. destroyed in 1906); Gustav 
Eisen, and Vaslet collectors. 

Type locality. — Sierra San Lazaro, Baja California. 

Bange. — Cape region of Baja California, including Isla Espiritu 
Santo and Isla Partida. 

SCELOPORUS MAGISTER MAGISTER Hallowell 

Sceloporus magister Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 1854, 

p. 93.— Stejneger, North Amer. Fauna, No. 7, 1893, pp. 160, 178-183, 

pi. 1, figs. 2, a-c. 
Sceloporus magister magister, Linsdale, Univ. California Publ. Zool., vol. 38, 

1932, p. 365.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 

pp. 145-161, pi. 15. 

Type.— U.S.N.M. No. 2967; A. L. Heermann collector. 

Type locality. — Fort Yuma, Ariz. 

Bange. — Extreme western Texas to southern California (excluding 
western slopes), north to southern Nevada and southwestern Colorado, 
south to northern Durango and southern Sonora. Reported from 
Chihuahua, Coahuila, Durango, Sonora, and Baja California. 



CHECKLIST OF REPTILES OF MEXICO 115 

SCELOPORUS MAGISTER RUFIDORSUM Yarrow 

Sceloporus rufidorsum Yarrow, Proc. U. S. Nat. Mus., vol. 5, 1882, pp. 442-443 

(part) . 
Sceloporus viagister rufidorsum, Linsdale, Univ. California Publ. Zool., vol. 38, 

1932, p. 366.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 

pp. 161-165. 

r?/2?g.— U.S.N.M. No. 11981; L. Belding collector. 

Type locality. — "San Quentin Bay," Baja California. In Yarrow, 
U.S. Nat. Mus. Bull. 24, 1882, p. 64, the locality is listed "La Paz, 
California." 

Range. — Northern Baja California, exclusive of the northeastern 
section, southv^^ard to include the Vizcaino Desert and Cedros Island ; 
Coronados Islands. 

SCELOPORUS MAGISTER UNEATULUS Dickerson 

Sceloporus lineatulus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 

467-468. 
Sceloporus magister lineatulus, Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 

26, 1939, pp. 168-170, fig. 13. 

T?/2)e.— U.S.N.M. No. 64263; C. H. Townsend collector. 
Type locality. — Santa Catalina Island, Gulf of California. 
Range. — Known only from Santa Catalina Island. 

SCELOPORUS MAGISTER MONSERRATENSIS Van Denburgh and Slevin 

Sceloporus monserratensis Van Denburgh and Slevin, Proc. California Acad. 

Sci., ser. 4, vol. 11, 1921, p. 396. 
Sceloporus magister monserratensis. Smith, Publ. Field Mus. Nat. Hist., zool. ser., 

vol. 26, 1939, pp. 165-168. 
Sceloporus magister rufidorsum, Linsdale, Univ. California Publ. Zool., vol. 38, 

1932, p. 366 (part). 

Type. — California Acad. Sci. No. 50509; Joseph R. Slevin collector. 

Type locality. — Monserrate Island, Gulf of California. 

Range. — Southern edge of the Vizcaino Desert in Baja California 
southward to the southern end of the Sierra de la Gigante; islands 
adjacent to the coast in the gulf except Santa Catalina. 

SCELOPORUS MAGISTER ZOSTEROMUS Cope 

Sceloporus zosteromus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 105. — 
Stejneger, North Amer. Fauna, No. 7, 1893, pp. 178, 181, pi. 1, fig. 3, a-c. 

Sceloporus magister zosteromus, Linsdale, Univ. California Publ. Zool., vol. 38, 
1932, p. 366.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 170-172. 

Type.— U.S.N.M. Nos. 5298 (23 specimens), 69472-69488; John 
Xantus collector. 

Type locality. — Cape San Lucas, Baja California. 
Range. — Cape region of Baja California. 



116 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

SCELOPORUS HORRIDUS HORRIDUS Wiegmann 

Sceloporus horridus Wiegmann, Herpetologia Mexicana, 1834, p. 50. 
Sceloponis horridus horridus, Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, pp. 98-106, pi. 11. 

Type. — Zool. Mus. Berlin; F. Deppc collector. 
Type locality. — Mexico, here restricted to Cucrnavaca, Morelos. 
Range. — Known in southern Morelos, Guerrero, and Oaxaca in the 
Balsas Basin, and southern Pucbla. 

SCELOPORUS HORRIDUS OUGOPORUS Cope 

Sceloporus oligoporus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 177-178. 
Sceloporus horridus oligoporus, Taylor, Univ. Kansas Sci. Bull., vol. 24, 1938, 

p. 520. — Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 

106-108. 

Type.— U.S. 'N.M. Nos. 31386-31393, cotypes. 
Type locality. — Colima, Colima. 

Range. — Known from western Guerrero, central and southern 
Michoacan, Colima, and central Jalisco to Durango. 

SCELOPORUS HORRIDUS ALBIVENTRIS Smith 

Sceloporus horridus albiventris Smith, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 108-110, pi. 12. 

Type.— EKT-RMS No. 8519; E. H. Taylor collector. 
Type locality. — Tepic, Nayarit. 

Range. — Known from coastal regions from northern Jalisco through 
Nayarit to Sinaloa. 

SCELOPORUS SPINOSUS SPINOSUS Wiegmann 

Sceloporus spinosus Wiegmann, Isis von Oken, vol. 21, 1828, p. 370; Herpetologia 

Mexicana, 1834, p. 50, pi. 7, fig. 3. 
Sceloporus spinosus spinosus, MartIn del Campo, Anal. Inst. Biol. Mexico, 

vol. 8, 1937, p. 262.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 

1939, pp. 87-94, pi. 11. 

Type. — ? Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, here restricted to Puebla, Puebla. 

Range. — Durango and western Tamaulipas to northern Jalisco, 
Michoacan, Hidalgo, and Puebla on the Mexican Plateau. Recorded 
from Durango, Zacatecas, San Luis Potosi, Tamaulipas, Jalisco, 
Aguascalientes, Guanajuato, Hidalgo, Veracruz, Mexico, Michoacan, 
Puebla, and Distrito Federal. 

SCELOPORUS SPINOSUS CAERULEOPUNCTATUS Smith 

Sceloporus spinosus caeruleopunctatus Smith, Univ. Kansas Sci. Bull., vol. 24» 
1936 (1938), pp. 469-473; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26. 
1939, pp. 94-98, fig. 10. 

Type— ERT-RMS No. 8467; H. M. Smith collector. 



CHECKLIST OF REPTILES OF MEXICO 117 

Type locality. — Slopes of Cerro de San Luis, about 15 miles north of 
Oaxaca, Oaxaca. 

Range. — Kno^v^l only from the highlands of Oaxaca. 

SCELOPORUS OLIVACEUS Smith 

Sceloporus spinosus floridanus, Stejneger, North Amer. Fauna, No. 7, 1893, 
p. 181 (nee Baird). 

Sceloporus olivaceus Smith, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 263, 
277-279; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 110-118, 
pis. 13-14; Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 103-106. 

Type.— EUT-HMS No. 2508; E. H. Taylor and John S. Wright 
collectors. 

Type locality. — Arroyo Los Olmos, 3 miles southeast of Rio Grande 
City, Starr County, Tex. 

Range. — Extreme south-central Oklahoma, southward through 
central Texas to southern Tamaulipas, central Nuevo Le6n, south- 
eastern Coahuila, and San Luis Potosi. Recorded from each state 
named. 

UNDULATUS GROUP 

Species. — Four, with a total of 13 forms. Extralimital are 9 forms 
all restricted to the United States and Mexico. 

Range. — British Columbia and southern New York south to 
northern Baja California and central Zacatecas. 

KEY TO MEXICAN SPECIES OF THE UNDULATUS GROUP OF SCELOPORUS 

1. Supraoculars large, entire; femoral pore series separated by 9 or more scales; 

gular patches small, widely separated cautus (p. 117) 

Supraoculars smaller, divided; femoral pore series separated by 8 or fewer 
scales, or, gular patches large, often covering entire throat 2 

2. Femoral pore series separated by 9 scales or more; scales on posterior surface 

of thigh abruptly differentiated from dorsal scales of same member, the 
median posterior scales not distinctly larger than adjacent lateral posterior 

scales occidentalis biseriatus (p. 118) 

Femoral pore series separated by 8 scales or less; scales on posterior surface of 
thigh gradually merging with larger dorsals of same member, at least the 
median posterior scales distinctly larger than the adjacent lateral posterior 
scales 3 

3. Males without lateral belly patches; dorsolateral and lateral hght stripes very 

clearly defined undulatus virgatus (p. 118) 

Males with lateral belly patches; dorsolateral and lateral light stripes poorly 
defined undulatus consobrinus (p. 118) 

SCELOPORUS CAUTUS Smith 

Sceloporus cautus Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 387, 1938, 
pp. 2-7; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 173-174. 

Type.— EUT-HMS No. 13027; E. H. Taylor and H. M. Smith 
collectors. 



118 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type locality. — Thirty miles north of El Salado, San Luis Potosf, 
in the state of Coahuila, Mexico. 

Range. — Southern Coahuila, northern San Luis Potosi, and eastern 
Zacatecas. 

SCELOPORUS OCCIDENT AUS BISERIATUS Hallowell 

Sceloporus bi-senatus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 7; 

1S54, pp. 93-94. 
Sceloporus occidentalis bi-seriatus, Camp, Univ. California Publ. Zool., vol. 17, 

1916, p. 65. 

Type. — Acad. Nat. Sci. Philadelphia Nos. 8474-6, cotypes. 

Type locality. — "Borders of El Paso Creek and Tejon Valley," 
Calif., here restricted to El Paso Creek. 

Range. — Southern Idaho, southwest through Nevada and western 
Utah into California and the San Pedro Martir range in Baja Cali- 
fornia. 

SCELOPORUS UNDULATUS CONSOBRINUS Baird and Girard 

Sceloporus consobrinus Baird and Girard, in Marcy and McClellan, Exploration 

of the Red River of Louisiana, . . ., 1853, pp. 224-226, 237, pi. 10, figs. 

5-12. 
Sceloporus tindulatus consobrinus, Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 

p. 337, fig. 60.— Smith, Publ. Field Mus. Nat. Hist., zool ser., vol. 26, 1939, 

pp. 175-176. 

Type. — Destroj'-ed. 

Type locality. — Beckham County, Okla., near the confluence of the 
north fork of the Red River and Suydam Creek. 

Range. — Texas except eastern fourth, southwestern Oldahoma, ex- 
treme western and southern fourth of New Mexico, southeastern 
Arizona in vicinity of San Pedro River, south in Mexico to central 
Zacatecas, central Chihuahua on the west and central Tamaulipas 
on the east. Recorded only from Chihuahua, Coahuila, Durango, 
Nuevo Le6n, and Zacatecas. 

SCELOPORUS UNDULATUS VIRGATUS Smith 

Sceloporus undrilatus virgatus Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
3S7, 193S, pp. 11-14; Publ. Field Mus. Nat. Hist., zool ser., vol. 26, 1939, 
p. 176. 

Type.~Vmv. ]Mich. Mus. Zool. No. 81912; Berry Campbell 
collector. 

Type locality. — Above Santa jSIaria mine. El Tigre Mountains, 
Sonora. 

Range. — Southeastern Arizona and adjacent Alexico, at high ele- 
vations. Recorded in Mexico only from Sonora and Chiliuahua. 

GRACIOSUS GROUP 

Species. — One, with three subspecies, only one of which occurs in 
Mexico. 



CHECKLIST OF REPTILES OF MEXICO 119 

Range. — Montana and Washington south to extreme northwestern 
Baja California. 

SCELOPORUS GRAaOSUS VANDENBURGIANUS Cope 

Sceloporus vandenburgianus Cope, Amer. Nat., vol. 30, 1896, pp. 834-835; Ann. 

Rep. U. S. Nat. Mus. 1898 (1900), p. 390, fig. 64. 
Sceloporus graciosus vandenburgianus, Camp, Univ. California Publ. Zool., vol. 

17, 1916, p. 67. 

Type.— v. S.N. M. No. 21931; E. A. Mearns collector. 

Type locality. — Summit of the coast range, San Diego County, 
Calif. 

Range. — Mountains south from Ventura County, Calif., into 
northwestern Baja California. 

GRAMMICUS GROUP 

Species. — Two, one of which includes three subspecies. 

Range. — Extreme southern Texas and Chihuahua south to Oaxaca. 

KEY TO SPECIES OF THE GRAMMICUS GROUP OF SCELOPORUS 

1. Dorsal scales unequal; a series of enlarged scales on each side of middorsal 

line, separated from each other by small, flat scales. -heterolepis (p. 121) 
Dorsal scales more or less uniform in size 2 

2. Scales on sides of neck not abruptly differentiated from dorsal nuchal scales; 

no enlarged series of scales on sides of neck; dorsal scales 48-66. 

grammicus gramnaicus (p. 119) 

Scales on sides of neck abruptly differentiated from dorsal nuchal scales; two 

series of enlarged scales on sides of neck posterior to ear; dorsal scales 

52-93 3 

3. Dorsal scales usually less than 70 (52-74) grammicus disparilis (p. 120) 

Dorsal scales usually 70 or more (68-93) -grammicus microlepidotus (p. 120) 

SCELOPORUS GRAMMICUS GRAMMICUS Wiegmann 

Sceloporus grammicus Wiegmann, Isis von Oken, vol. 21, 1828, p. 370. — Smith, 

Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 179-182, fig. 16. 
Sceloporus grammicus grammicus, Smith and Laufe, Trans. Kansas Acad. Sci., 

vol. 48, 1945, pp. 332-333. 
Sceloporus pleurostictus Wiegmann, Isis von Oken, vol. 21, 1828, p. 370 (Zool. 

Mus. BerUn; F. Deppe collector; Mexico, here restricted to Chilpancingo, 

Guerrero). 
Sceloporus rubriventris Gunther, Biologia Centrali-Americana, Reptilia and 

Batrachia, 1890, p. 72, pi. 32, fig. C (Omilteme, Guerrero; Brit. Mus. Nat. 

Hist.). 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, here restricted to Chilpancingo, Guerrero. 

Range. — Southern Oaxaca (San Pedro Quiechapa, Ozolo tepee) and 
the Sierra Madre del Sur in Guerrero (Omilteme, Chilpancingo, 
Tamazulapan, Iguala). 



120 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

SCELOPORUS GRAMMICUS MICROLEPIDOTUS Wiegmann 

Sceloporus grammicus var, alpha Wiegmann, Isis von Oken, vol. 21, 1828, p. 370. 
Sceloporus microlepidotus Wiegmann, Herpetologia Mexicana, 1834, p. 51. — 

BocouRT, Mission scientifique au Mexique . . . , fitudes sur les reptiles, 

livr. 3, 1874, pp. 194-195, pi. 18 bis, fig. 13, a-d. 
Sceloporus microlepidotus microlepidotus, Smith, Publ. Field Mus. Nat. Hist., 

zool. ser., vol. 26, 1939, pp. 184-191 (part), fig. 17, 18. 
Sceloporus grammicus microlepidotus, Smith and Laufe, Trans. Kansas Acad. 

Sci., vol. 48, 1945, pp. 332-333. 
Sceloporus dispar Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 

1852, p. 127 (Veracruz, here restricted to Cruz Blanca; type lost). 
Sceloporus heterurus Cope, Proc. Acad. Nat. Sci. Philadelphia 1866 (1867), pp. 

322-323 (Mirador, Veracruz; type now lost). 
Sceloporus microlepidopterus Herrera, La Naturaleza, ser. 2, vol. 1, 1890, p. 331 

[lapsus calami). 
Sceloporus microlepis Boulenger, Proc. Zool. Soc. London, 1894, p. 731 {lapsus 

calami) . 

Type. — Zool. Mus. Berlin; F. Deppe, collector. 

Type locality. — Mexico, here restricted to Mexico, D. F. 

Range. — Southern part of the Mexican plateau from Jalisco, 
northern Guanajuato, and northern Hidalgo south to Oaxaca. Re- 
corded from the states of Colima, Distrito Federal, Guanajuato, 
Hidalgo, Jalisco, Oaxaca (Reyes), ^^ Puebla, Veracruz, Mexico, 
Morelos, Tlaxcala, and Michoacan. 

SCELOPORUS GRAMMICUS DISPARILIS Stejneger 

Sceloporus disparilis Stejneger, Proc. Biol. Soc. Washington, vol. 29, 1916, pp 

227-230. 
Sceloporus microlepidotus disparilis, Dunn, Proc. Acad. Nat. Sci. Philadelphia, 

vol. 88, 1936, p. 472.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 

1939, pp. 191-197, fig. 18, pi. 16. 
Sceloporus grammicus disparilis, Smith and Laufe, Trans. Kansas Acad. Sci., 

vol. 48, 1945, pp. 332-333. 
Sceloporus microlepidotus microlepidotus, Dunn, Proc. Acad. Nat. Sci. Phila- 
delphia, vol. 88, 1936, pp. 472-473, 474. 
Sceloporus pilsbryi Dunn, Proc. Acad. Nat. Sci. Philadelphia, vol. 88, 1936, pp. 

473-474 (Alvarez, San Luis Potosf; type, Acad. Nat. Sci. Philadelphia No, 

20085). 

Type.— U.S.N. M. No. 33041; WiUiam Lloyd collector. 

Type locality. — Lomita Ranch, 6 miles north of Hidalgo, Tex. 

Range. — Northern Hidalgo, Guanajuato, and southern Zacatecas 
northward to central Chihuahua and northern Coahuila and the 
southern tip of Texas. Recorded from Chihuahua, Coahuila, 
Durango, Guanajuato, Hidalgo, Nuevo Le6n, San Luis Potosf, 
Tamaulipas, and Zacatecas. 



" Intergrades of S. g. grammicus and S. g. microlepidotus are from Cerro San Felipe and Cerro San Juan> 
north of Oaxaca City. 



CHECKLIST OF REPTILES OF MEXICO 121 

SCELOPORUS HETEROLEPIS Boulenger 

Sceloporus heterolepis Boulenger, Proc. Zool. Soc. London, 1894, pp. 724, 731, 
pi. 48, fig. 4.— Smith, Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp, 
197-199, pi. 16. 

Type. — Brit. Mus. Nat. Hist., part of cotypes; one cotype, Mus. 
Comp. Zool. No. 32346. 

Type locality. — La Cumbre de los Arrastrados (to which the type 
locahty is here restricted), Real Alto,"Iliocho (=Rancho?)" La Ber- 
beria, Sierra de Bolaiios. 

Range. — Kno\vii only from western Jalisco (type localities and 
Cerro Tequila.) 

MEGALEPIDURUS GROUP 

Species. — Two. 

Range. — Western central Veracruz and eastern Puebla. 

KEY TO SPECIES OF THE MEGALEPIDURUS GROUP OF SCELOPORUS 

1. Dorsal scales 44 to 56; scales between femoral pore series 2 to 5; basal sub- 
caudals keeled in females; males with distinct blue areas on sides of belly. 

pictus (p. 121) 
Dorsal scales 54 to 62; scales between series of femoral pores 4 to 8; basal sub- 
caudals smooth in females; males immaculate below. 

megalepidurus (p. 121) 

SCELOPORUS MEGALEPIDURUS Smith 

Sceloporus megalepidurvs Smith, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 
272-277, pi. 9, figs. 7, 8, pi. 10, fig. 13; Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 204-209, fig. 20. 

Type.—EWT-RU^ No. 7543 (EHT field No. 2908); E. H. Taylor 
and H. M. Smith collectors. 

Type locality. — Near Totalco, Veracruz. 

Range. — Known only from northern Puebla, western Veracruz, and 
Tlaxcala (Apizaco). 

SCELOPORUS PICTUS Smith 

Sceloporus pictus Smith, Amer. Mus. Nov., No. 892, 1936, pp. 1-4; Publ. Field 
Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 200-204, pi. 17, two figs. 

Type.— Amer. Mus. Nat. Hist. No. 18744; Paul D. R. Riithling, 
collector. 

Type locality. — Santa Catarina, Puebla. 

Range. — Central Puebla and central western Veracruz. 

TORQUATUS '« GROUP 

Species. — Twenty-one forms, representing 10 species, are recognized. 
Range. — Arizona and Texas south to Guatemala. 

'• This group was in 1938 termed the poinsetlii group, inasmuch as Sceloporus poinsettii Baird and Qirard 
(1854) was the oldest name in the group after exclusion of S. torqiiatus Wiegmann, 1828. The latter name 
was considered unavailable because it was once (Wagler, 1830) placed in the genus Tropidurus, in 
which the combination Tropidurus torguatus (Wied, 1820) had already been made (Wied, 1824). Secondary 
homonyms, such as this, have not, however, been regarded in recent years as permanently suppressed: 
they are suppressed only as long as they actually remain homonyms. Accordingly, we revert to the 
name torquatut for this group. 



122 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

KEY TO SPECIES OF THE TORQUATUS GROUP OF SCELOPORUS '«» 

1. Supraoculars in a single series, with no scale divided 2 

Supraoculars in 2 series; or, if in one series, with one or more scales divided. 9 

2. Femoral pores 8-14; dorsal scales 25-35 3 

Femoral pores more than 14; or dorsal scales more than 35 5 

3. Dorsal nuchal collar broadly interrupted medially by a space about 5 scales 

wide; throat mottled torquatus binocularis (p. 126) 

Dorsal nuchal collar broad, complete; throat not mottled 4 

4. Femoral pore counts on both sides usually total more than 21; dorsal scales 

usually 31 or more serrifer plioporus (p. 124) 

Femoral pore counts on both sides usually total less than 22; dorsal scales 
generally less than 31 serrifer serrifer (p. 123) 

5. Dorsal scales 31 or less 6 

Dorsal scales more than 31 7 

6. Nuchal collar divided on each side of neck, the area between lighter in color; 

dorsal color light, with dark and light spots irregularly placed; size large 
(maximum snout-vent measurement 129 mm.); dorsal scales more strongly 

keeled and mucronate torquatus melanogaster (p. 126) 

Nuchal collar broad, complete; dorsal color dark, without light spots; maxi- 
mum snout-vent measurement 98 mm.; dorsal scales more weakly keeled 
and mucronate torquatus torquatus (p. 125) 

7. Lateral scales about one-half as large as median dorsals -lineolateralis (p. 126) 
Lateral scales as large as or larger than dorsals 8 

8. Dorsal scales 40 or more; nuchal collar narrow, with light borders broken; a 

light line on side of head and another on side of neck; each dorsal scale 

usually with a light median spot jarrovii jarrovii (p. 128) 

Dorsal scales usually less than 40; nuchal collar broad, with unbroken light 
borders; no light lines on sides of head and neck; no spots on dorsal scales. 

buUeri (p. 125) 

9. Lateral scales with the terminal point arising well within the free posterior 

margin; dorsal scales 41 or more 10 

Lateral scales with the terminal point arising at or very near the free posterior 
margin; head scales not microscopically rugose; no oblique dark blue lines 
on throat 11 

10. Dorsal scales 47 to 54; throat with very distinct, oblique, dark blue lines; 

head scales not microscopically rugose; oblique dark bands on sides of body 
distinct; maximum snout-vent measurement 79 mm. 

dugesii intermedius (p. 127) 

Dorsal scales 41 to 50; throat without or with very faint oblique lines; head 

scales microscopically rugose; oblique dark bands on sides of body indistinct 

or absent; maximum snout-vent measurement 87.5 mm; femoral pores not 

over 13 dugesii dugesii (p. 127) 

11. Dorsal scales 55 or more ornatus ornatus (p. 127) 

Dorsal scales less than 55 12 

12. Dorsal scales 47 to 53 ornatus caeruleus (p. 127) 

Dorsal scales less than 47 13 

13. Tail with very distinct, broad, alternating dark and light bands, most distinct 

toward tip, where they are complete; supraoculars in 2 complete rows; 
head scales very irregular; light borders of nuchal collar broad; a broad light 
band across neck behind occiput; preocular usually entire; inner row of 
labiomental scales generally terminating at a point posterior to suture be- 
tween second and third infralabials poinsettii (p. 125) 

"» For one other species of this group, see footnote 72a. 



CHECKLIST OF REPTILES OF MEXICO 123 

Tail without distinct alternating dark and light bands of nearly equal width; 
bands about tail not complete toward tip; dorsal scales 40 or less 14 

14. Dorsal scales 31 or less 15 

Dorsal scales 32 or more 16 

15. A middorsal series of very large, dark blotches, sometimes fused bandlike; 

all except males with a pair of parallel, closely placed dark lines down 
middle of throat; collar 4 scales long or less. 

mucronatus mucronatus (p. 124) 

No middorsal series of dark blotches; parallel throat lines absent; collar 

involving 4)4 scale lengths or more mucronatus omilteraanus (p. 124) 

16. Each dorsal scale row, in adult males, with a longitudinal light Hne; dorsals 

30 to 38 mucronatus aureolus (p. 124) 

Scale rows not with continuous lines 17 

17. Nuchal black collar 4 or 5 scales wide, bordered anteriorly and posteriorly 

by a light band 1 or 1J4 scales wide; each border interrupted medially by a 
light scale; snout-vent length often more than 100 mm., reaching 143 

mm cyanogenys (p. 125) 

Not so; maximum snout-vent length 100 mm 18 

18. Lateral body scales distinctly decreasing in size laterally, at a point halfway 

between axilla and groin, distinctly smaller than dorsal scales 19 

Lateral body scales not decreasing in size laterally at least up to a point half- 
way between axilla and groin, where they are still subequal to, or even a 
little larger than, middorsal scales 20 

19. Dorsal scales on lower foreleg about half size of those on upper foreleg; dark 

transverse streaks in lateral belly patches; nuchal collar covering six scales 

medially, or more jarrovii sugillatus (p. 128) 

Dorsal scales on lower foreleg but little if any smaller than those on upper 
foreleg; no dark streaks in lateral beUy patches; nuchal collar less than four 
scales long middorsally jarrovii immucronatus (p. 128) 

20. Supraoculars essentially in 1 row; if an outer row is evident, it is composed of 

scales much smaller than those of inner row, and usually number no more 

than two jarrovii jarrovii (p. 128) 

Supraoculars in 2 rows, those of outer row a little smaller than those of inner, 
usually numbering 3 or more 21 

21. Adult males black above and below, with orange areas and spots on sides of 

head, belly, and tail; only throat, underside of tail, and posterior surface of 
hind leg not black; females somewhat similar, very dark above, the collar 
poorly defined, young with poorly defined, narrow, light borders on neck 

collar; dorsal scales average 37.5 jarrovii oberon (p. 129) 

Adult males light brown above, with very broad, very well defined nuchal 
collar; a median area on belly white except in very largest males; sides of 
abdomen blue, black-edged; females and young with more distinct light 
borders on nuchal collar; dorsal scales average 40.6. jarrovii minor (p. 128) 

SCELOPORUS SERRIFER SERRIFER Cope 

Sceloporus serrifer Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 124. — 
Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 558-564, figs. 5, 6, 
pi. 47, fig. 2. 

Sceloporus serrifer serrifer, Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 
1939, p. 212, fig. 23. 
861316—50 9 



124 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type.— V.S.N M. No. 34868; Arthur Schott collector. 

Type locality. — Yucatan, here restricted to Merida. 

Range. — Northern part of the Yucatan Peninsula. Recorded only 
from the state of Yucatan; intergrades with S. s. plioporus are re- 
corded from Balchacaj, Campeche. 

SCELOPORUS SERRIFER PLIOPORUS Smith 

Sceloporus serrifer plioporus Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 
26, 1939, pp. 212-214, pi. 18. 

TV^e.— Chicago Nat. Hist. Mus. No. 32004; E. H. Taylor collector. 

Type locality. — Four miles east of Encero, Veracruz, 

Range. — Gulf coast of Mexico from southern Tamaulipas southward 

through the basal part of the Yucatan Peninsula to Peten, Guatemala. 

Recorded from the states of Tamaulipas, San Luis Potosi, Veracruz, 

and Tabasco (Tenosique) ; intergrades with S. s. serrifer are recorded 

from Balchacaj, Campeche, 

SCELOPORUS MUCRONATUS MUCRONATUS Cope 

Sceloporus torquatus mucronatus Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, p. 

402. 
Sceloporus mucronatus mucronatus, Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 

(1938), pp. 583-591, figs. 10, 11, pi. 49, fig. 2; Publ. Field Mus. Nat. Hist. 

zool. ser., vol. 26, 1939, pp. 218-220, figs. 26, 27. 

Type.— U.S.N. M. Nos. 25074-9; No. 25077 designated as lectotype 
(Smith, op. cit, 1938, p. 583). 

Type locality. — Mirador, Veracruz. 

Range. — Parts of Hidalgo, Veracruz, Puebla, and Mexico, at high 
elevations. 

SCELOPORUS MUCRONATUS AUREOLUS Smith 

Sceloporus mucronatus omiltemanus, Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 

(1938), pp. 591-598 (part), figs. 11, 12, pi. 50. 
Sceloporus m,ucronatus aureolus Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 

356-357. 

Type.— v. S.N. M. No. 112232; H. M. Smith collector. 
Type locality. — Two miles west of Acultzingo, Veracruz. 
Range. — Western central Veracruz and southern Puebla. 

SCELOPORUS MUCRONATUS OMILTEMANUS Gtinther 

Sceloporus omiltemanus Gtjnther, Biologia Centrali-Americana, Reptilia and 

Batrachia, 1890, p. 66, pi. 32, fig. A. 
Sceloporus t.[orquatus] omiltemanus, Taylor, Proc. Biol. Soc. Washington, vol. 

44, 1931, p. 129. 
Sceloporus mucronatus omiltemanus, Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 

(1938), pp. 591-598. 

Type. — ^Brit. Mus. Nat. Hist.; H. H. Smith collector. 



CHECKLIST OF REPTILES OF MEXICO 125 

Type locality. — Omilteme, Guerrero, 8,000 feet elevation. 
Range. — Central highlands of Oaxaca and the Sierra Madre del Sur 
in Guerrero .^^ 

SCELOPORUS POINSETTII Baird and Glrard 

Sceloporus poinsettii Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, vol. 
6, 1854, pp. 126-127.— Smith, Univ. Kansas Sci. BuU., vol. 24, 1936 (1938), 
pp. 606-617, pi. 51, fig. 1, text figs. 14, 15. 

Type.— U.S. N.M. No. 2948 cotypes, 2 specimens, and No. 2952, 2 
specimens, "Sonora"; J. H. Clark collector. 

Type locality. — "Rio San Pedro of the Rio Grande del Norte, and 
the province of Sonora," here restricted to the former locality. 

Range. — Southern New Mexico east to central Texas, south through 
western Nuevo Ledn and southern Coahuila to central Durango. 
Recorded in Mexico from Chihuahua, Coahuila, Durango, and Nuevo 
Le6n. 

SCELOPORUS CYANOGENYS Cope 

Sceloporus torquatus cyanogenys Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, 

p. 402. 
Sceloporus cyanogenys, Smith, Univ. Kansas Sci. BuU., vol. 24, 1936 (1938), 

pp. 599-606, pi. 51, fig. 2, text figs. 13-14. 
Sceloporus torquatus mucronatus, Boulenger, Catalogue of the lizards in the 

British Museum, 1885, vol. 1, p. 220 (part). 
Sceloporus torquatus poinsettii, Burt, Copeia, 1932, No. 3, p. 158 (part). 

7V2)e.— U.S.N. M. Nos. 31373-31377, cotypes; Acad. Nat. Sci. Phila- 
delphia, 11304-11305, cotypes. 

Type locality. — Monterrey, Nuevo Le6n. 

Range. — Southern Texas from Devils River and Starr County to 
central Tamaulipas and Nuevo Le6n. Recorded in Mexico only from 
the states of Tamaulipas and Nuevo Le6n. 

SCELOPORUS BULLERI Boulenger 

Sceloporus bulleri Boulenger, Proc. Zool. Soc. London, 1894, pp. 729-730, pi. 
48, fig. 3.— Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 579-583, 
figs. 8, 9, pi. 49, fig. 1. 

Type.—BTit. Mus. Nat. Hist. 

Type locality. — La Cumbre de los Arrastrados, Jalisco. 

Range. — Western and southern parts of Jalisco. 

SCELOPORUS TORQUATUS TORQUATUS Wiegmann" 

Sceloporus torquatus Wiegmann, Isis von Oken, 1828, p. 369. 

Sceloporus torquatus torquatus, Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, 

pp. 402, 403.— Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 

564-571, figs. 7, 8, pi. 48, fig. 1. 



" a record for Chiapas (Smith, Publ. Field Mus. Nat. Hist., zool. ser„ vol. 26, 1939, p. 221) is in error; 
the specimen referred to is an S. prezygus. 
" See footnote 76 for use of this name. 



126 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Agama torquata Peale and Green, Journ, Acad. Nat. Sci. Philadelphia, vol. 6, 
1830, pp. 231-234 (type Acad. Nat. Sci. Philadelphia; Mexico, here restricted 
to Mexico, D. F.). 

Sceloporus ferrariperezi Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, p. 400 
(cotypes, U.S.N.M. Nos. 9874, 9876, 9878, 9880, 9895, and No. 9897 which 
is an S. spinosus spinosus; Guanajuato, Guanajuato, by present restriction). 

Sceloporus ferrariperezi ferrariperezi, Smith, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 26, 1939, pp. 214-216, fig. 24. 

Type. — Zool. Mus. Berl.; F. Deppe collector. 

Type locality. — Mexico, here restricted to Mexico, D. F. 

Range. — Central Mexico, including Hidalgo, west-central Veracruz, 
Mexico, Distrito Federal, northern Puebla, eastern Morelos, southern 
Guanajuato, and northern Michoacan. Recorded only from the 
states cited. 

SCELOPORUS TORQUATUS BINOCULARIS Dunn 

Sceloporus binocularis Dunn, Proc. Acad. Nat. Sci. Philadelphia, vol. 88, 1936, 

pp. 474-475. 
Sceloporus ferrariperezi binocularis, Smith, Publ. Field Mus. Nat. Hist., zool. 

sen, vol. 26, 1939, pp. 216-218, pi. 18. 

Type.— Acad. Nat. Sci. Philadelphia No. 20032 ; Henry A. Pilsbry, 
Francis W. Pennell, and Cyril H. Harvey collectors. 

Type locality. — Trail from Pablillo to Alamar, Nuevo Leon. 
Range. — Known only from the general region of the type locality. 

SCELOPORUS TORQUATUS MELANOGASTER Cope 

Sceloporus melanogaster Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, pp. 400-401. 
Sceloporus iorquatus melanogaster, Boulenger, Catalogue of the lizards in the 

British Museum, vol. 2, 1885, p. 220. — Smith, Univ. Kansas Sci. Bull., vol. 

24, 1936 (1938), pp. 571-579, pi. 48, fig. 2, text figs 1, 8. 
Sceloporus ferrariperezi melanogaster, Smith, Publ. Field Mus. Nat. Hist., zool. 

ser., vol. 26, 1939, p. 216. 

T2/i?e.— U.S.N.M. No. 9877; Alfredo Duges collector. 

Type locality. — "Probably from Guanajuato"=Noria (Michoacan), 
or Tupataro, near Cueramaro, Guanajuato ^irfe Dug6s (La Naturaleza, 
ser. 2, vol. 1, 1887, pp. 114-115), here restricted to Tupataro. 

Range. — Northern Jalisco, west through most of Guanajuato, north 
through central and southern San Luis Potosi and Zacatecas. Re- 
corded from Aguascalientes, Guanajuato, Jalisco, Michoacan, San 
Luis Potosi, and Zacatecas. 

SCELOPORUS LINEOLATERALIS Smith 

Sceloporus lineolateralis Smith, Proc. Biol. Soc. Washington, vol. 49, 1936, pp. 
92-96; Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 617-624, pi. 52, 
figs. 16, 17; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 225-226, 
fig. 31. 



CHECKLIST OF REPTILES OF MEXICO 127 

Type.-ERT-UMS, No. 4323; E. H. Taylor and H. M. Smith 
collectors. 

Type locality. — Six miles northeast of Pedricena, Dm^ango, 
Range . — D ur ango . 

SCELOPORUS ORNATUS ORNATUS Baird 

Sceloporus ornatus Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858 (1859), p. 254. 
Sceloporus ornatus ornatus, Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 
pp. 647-651, figs. 21, 22, pi. 54, fig. 1. 

TV^e.— U.S.N.M. No. 2845; Darius Nash Couch collector. 
Type locality. — Patos, Coahuila. 
Range. — Southeastern CoahuUa. 

SCELOPORUS ORNATUS CAERULEUS Smith 

Sceloporus ornatus caeruleus Smith, Copeia, 1936 (1937), No. 4, pp. 227-230; 
Univ. Kansas Sci. Bull., vol. 24, 1936 (1938) pp. 652-657, figs. 22-23, pi. 54, 
fig. 2. 

Type.—Vmv. Kansas Mus. Nat. Hist. (DHD-HMS field No. 
350); D. H. Dunkle and H. M. Smith, collectors (lost?). 
Type locality. — Five mUes south of San Pedro, CoahuUa. 
Range. — Known only from the type locality. 

SCELOPORUS DUGESII DUGESn Bocourt 

Sceloporus dugesii Bocourt, Ann. Sci. Nat., Zool., ser. 5, vol. 17, No. 10, 1873, 
p. 2; Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 3, 
1874, pp. 188-190, pi. 18, figs. 7-7b. 

Sceloporus dugesii dugesii, Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 
pp. 657-663, figs. 22, 24, pi. 55, fig. 1. 

Type. — Mus. Hist. Nat. Paris; Alfredo Duges collector. 

Type locality. — Colima. 

Range. — Along the coastal ranges of western Mexico from southern 
Nayarit to Colima. Recorded from the states of Jalisco, Colima, 
and Nayarit. 

SCELOPORUS DUGESn INTERMEDIUS (Dugfes) 

Tr[opidolepis] intermedius Ducfes, La Naturaleza, vol, 1, 1869, p.l43 (nomen 

nudum). 
Sceloporus intermedius Ducfes, La Naturaleza, vol. 4, 1877, pp. 29-34, pi. 1, 

figs. 21-32. 
Sceloporus dugesii intermedius. Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 

(1938), pp. 663-670. — Smith and Necker, Anal. Esc. Nac. Cien. Biol., 

vol. 3, 1943, pp. 212-214, pi. 2, figs. 2, 3. 
Sc[eloporus] Westphalii Duofes, La Naturaleza, vol. 4, 1877, p. 30 (substitute 

name for *S. intermedius, which Duges retained for his description of the 

species in 1877, although he says "Querfa haberle dado el nombre de Sc. 

Westphahi"). 

Type. — Mus. Alf. Duges; two co types; collector unknown. 



128 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type locality. — La Noria, near Zamora, hacienda of D. Epifanio 
Jimenez (in Michoac^n). 

Range. — Known from Guanajuato and northern and central 
Michoacan. 

SCELOPORUS JARROVII JARROVII Cope 

Sceloporus jarrovii Cope, in United States Geological and Geographical Surveys 

West of the 100th Meridian, vol. 5, 1875, p. 569. 
Sceloporus jarrovii jarrovii, Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 

pp. 624-631, figs. 17, 18, pi. 47, fig. 1. 

Type. — U.S.N.M. Nos, 8494, two specimens, and 8495, cotypes. 

Type locality. — Southern Arizona. 

Range. — Central Arizona east to western New Mexico, and south 
through Chihuahua and western Sonora to extreme western Zacatecas 
and extreme northern Nayarit. Recorded in Mexico from Chihuahua, 
Durango, and Sonora. 

SCELOPORUS JARROVn SUGILLATUS Smith 

Sceloporus jarrovii sugillatus Smith, Proc. U. S. Nat. Mas., vol. 92, 1942, pp. 
357-359. 

T?/pe.— U.S.N.M. No. 112100; H. M. Smith collector. 

Type locality.— hake No. 4, Zempoala, Mexico. 10,000 feet 
elevation. 

Range. — High elevations in Ajusco range. Known only from the 
type locality. 

SCELOPORUS JARROVn IMMUCRONATUS Smith 

Sceloporus jarrovii immucronatus Smith, Copeia, 1936 (1937), pp. 223-227; Univ. 
Kansas Sci. Bull., vol. 24, 1936 (1938), pp. 640-646, figs. 17, 20, pi. 53, fig. 1. 

r?/^e.— EHT-HMS No. 9358A (EHT-HMS field No. 500); E. H. 
Taylor and H. M. Smith collectors. 

Type locality. — Ten miles north of El Pinalito, Hidalgo. 

Range. — Western Queretaro tln-ough Hidalgo, eastern San Luis 
Potosi, to south-central Veracruz. Recorded from each state cited. 

SCELOPORUS JARROVII MINOR Cope 

Sceloporus torquatus minor Cope, Proc. Amcr. Philos. Soc, vol. 22, 1885, p. 402. 
Sceloporus jarrovii minor, Smith, Univ. Kansas Sci. Bull., vol. 24, 1936 (1938), 
pp. 631-639, figs. 17-19, pi. 53, fig. 2. 

Type.— U.S.N.M. Nos. 26166-26167. 

Type locality.— Zacsit ecus, here restricted to Valparaiso Mountains. 

Range. — Northern Queretero and Guanajuato through Mexico, 
western Zacatecas, and most of San Luis Potosi, to western Nuevo 
Le6n. Recorded from each state cited. 



CHECKLIST OF REPTILES OF MEXICO 129 

SCELOPORUS JARROVU OBERON Smith and Brown 

Sceloporus jarrovii oberon Smith and Buown, Publ. Field Mus. Nat. Hist., zool. 
ser., vol. 24, 1941, pp. 253-257, fig. 24. 

Type.—V.S'NM. No. 105823; H. M. Smith and Rozella Smith, 
collectors. 

Type locality. — Arteaga, Coahiiila. 

Range. — Southeastern Coahuila. Recorded only from the type 
locality, from Diamond Pass, Mount Zapaliname, near Saltillo, and 
from Sierra Guadelupe. 

VARIABILIS GROUP 

Species. — Nine forms, representing five species, are recognized in 
this group. One form, S. variabilis olloporus Smith, is extralimital, 
occurring from Guatemala to Costa Rica. 

Range. — Southern central Texas south to Costa Rica, entirely on 
Atlantic slopes west of the Isthmus of Tehuan tepee. 

KEY TO SPECIES OF THE VARIABILIS GROUP OF SCELOPORUS 

1 . Series of femoral pores separated medially by no more than 6 scales 2 

Series of femoral pores separated medially by 10 or more scales 3 

2. Dorsal scales 69 to 76; scales around body 70 to 81; dorsal scale rows at nape 

18 to 21 parvus scutulatus (p. 131) 

Dorsal scales 58 to 69; scales around body 61 to 69; dorsal scale rows at nape 
15 to 18 parvus parvus (p. 131) 

3. Ventral interfemoral scales separated from ventral thigh scales by a group of 

small scales one-third or one-fourth size of adjacent scales; a rudimentary 
gular fold; lateral scales much less than half size of ventral scales; dorsal 

scales 69 to 83 couchii (p. 132) 

Ventral interfemoral scales more or less continuous with ventral thigh scales; 
no rudimentary gular fold; lateral scales more than one-half size of ventral 
scales; dorsals usually less than 69 4 

4. Dorsal scales 36 to 47; subnasal usually absent; shank and posterior surface of 

lower foreleg distinctly banded; spots on back distinct in both sexes; preocular 

usually divided; frontoparietals usually in contact medially _teapensis (p. 130) 

Dorsal scales 47 or more 5 

5. Males and feinales immaculate below; femoral pores usually 9 or less on each 

side (occasionally more in females); dorsal scales 48 to 55; frontoparietals 
usually separated by an azygous scale; preocular usually entire; subnasal 

rarely present 6 

Males with red, blue-bordered areas on sides of abdomen; subnasal usually 
present; frontoparietals usually in contact medially; preocular usually 
divided 7 

6. Postrostrals usually 2, never 4; scales around body usually 53 to 58. 

cozumelae (mainland) (p. 130) 
Postrostrals usually 4, rarely 2 or 3; scales around body usually 59 to 64. 

cozumelae (typical) (p. 130) 

7. Dorsal scales 59 or more 8 

Dorsal scales usually less than 59; dorsolateral light lines 1 and 2 half scale 

rows wide posteriorly; maximum snout-vent measurement about 74 mm. 9 



130 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

8. Dorsolateral light stripes very distinct, 2 and 2 half scale rows wide posteriorly; 

females with sides of belly marked as in males, but less distinctly; maxi- 
mum snout-vent measurement 71 mm variabilis smithi (p. 131) 

Dorsolateral light stripes not so distinct, 1 and 2 half scale rows wide posteriorly; 
spots between dorsolateral light stripes very distinct; females with sides of 
belly immaculate; maximum snout-vent measurement 53 mm. 

variabilis marmoratus (p. 131) 

9. Femoral pores 12 or more variabilis variabilis (p. 130) 

Femoral pores 11 or less variabilis oUoporus (p. 129) 

SCELOPORUS COZUMELAE Jones 

Sceloporus cozumelae Jones, Occ. Pap, Mus. Zool. Univ. Michigan, No. 186, 1927, 
pp. 1-4. — Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
246-251. 

Type.— V.S.N. M. No. 13904; "U. S. Fish Commission" collector. 

Type locality. — Cozumel Island, Quintana Roo. 

Range. — Northern half of Yucatan Peninsula and adjacent Islands. 
Recorded from Cozumel and Mujeres Islands and the states of 
Yucatan and Quintana Roo. 

SCELOPORUS TEAPENSIS Gunther 

Sceloporus teapensis GtJNTHER, Biologia Centrali-Americana, Reptilia and Ba- 
trachia, 1890, pp. 75-76. — Smith, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 256-262, pi. 21. 

Type. — Brit. Mus. Nat. Hist., six co types; H. H. Smith collector. 

Type locality. — Teapa, Tabasco. 

Range. — Restricted to Atlantic slopes from southern Veracruz and 
northeastern Oaxaca, eastward through Chiapas, Tabasco, and Cam- 
peche, and through the Peten region of Guatemala to British Hon- 
duras, south to Coban, Alta Verapaz. Recorded from each state 
cited in Mexico. 

SCELOPORUS VARIABILIS VARIABILIS Wiegmann 

Sceloporus variabilis Wiegmann, Herpetologia Mexicana, 1834, p. 51. 
Sceloporus variabilis variabilis, Smith, Proc. Biol. Soc. Washington, vol. 47, 1934, 

pp. 127-129; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 262-272, 

fig. 45, pi. 22. 

Type. — ? Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, here restricted to Veracruz, Veracruz. 

Range. — Southern Tamaulipas along the Atlantic coast to south- 
central Veracruz (Rio Blanco), inland to eastern Queretaro and 
Puebla; through eastern Oaxaca and western Guatemala, reaching 
the coast only in Chiapas. Recorded from each state mentioned, 
and from San Luis Potosi, Guanajuato, and Hidalgo. 



CHECKLIST OF REPTILES OF MEXICO 131 

SCELOPORUS VARIABIUS SMITHI Hartweg and Oliver 

Sceloporus variabilis smithi Hartweg and Oliver, Occ. Pap. Mus. Zoo!., Univ. 
Michigan, No. 356, 1937, pp. 1-5. — Smith, Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 278-282, fig. 45, pi. 23. 

Type.—lJniY. Mich. Mus. Zool. No. 81777; N. Hartweg and J. A. 
Oliver collectors. 

Type locality. — Guengola Mountain, 6 kilometers northwest of the 
city of Tehuantepec, Oaxaca. 

Range. — Eastern Oaxaca. 

SCELOPORUS VARIABILIS MARMORATUS Hallowell 

Sceloporns marmoratus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 

1852, p. 178. 
Sceloporus variabilis marmoratus, Smith, Proc. Biol. Soc. Washington, vol. 47, 

1934, pp. 125-128; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 

272-278. 
Sceloporus delicatissimus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 

6, 1852, p. 178 (San Antonio, Tex.; type, U.S.N. M. No. 16020). 

Type. — Presumably lost. 

Type locality. — San Antonio, Tex. 

Range. — Atlantic lowlands from San Antonio, Tex., south to 
southern Tamaulipas, west to Coahuila. Recorded in Mexico only 
from the states of Coahuila, Nuevo Le6n, and Tamaulipas. 

SCELOPORUS PARVUS PARVUS Smith 

Sceloporus parvus Smith, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 263-267, 

figs. 1, 3, 10. 
Sceloporus parvus parvus, Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 358, 

1937, pp. 3-4; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 251- 

255, fig. 44. 

Type.— EUT-HMS No. 7120 (EHT field No. 279); E. H. Taylor 
and H. M. Smith collectors. 

Type locality. — Hills 5 miles west of Sabinas Hidalgo, Nuevo Le6n. 

Range. — Northern Nuevo Leon to southern San Luis Potosi, west 
of the Sierra Oriental. Recorded only from the two states cited, and 
from Coahuila (Arteaga). 

SCELOPORUS PARVUS SCUTULATUS Smith 

Sceloporus parvus scutulatus Smith, Occ. Pap. Mus. Zool. Univ. Michigan., No. 
358, 1937, pp. 4-6; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 255-256. 

Type.— ERT-RMS No. 7129; E. H. Taylor and H. M. Smith 
collectors. 

Type locality. — Thirty miles north of Matehuala, San Luis Potosi. 

Range. — Northwestern San Luis Potosi south along the Sierra 
Oriental to southern Hidalgo. Recorded only from the two states 
cited. 



132 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

SCELOPORUS COUCHII Baird 

Sceloporus couchii Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858 (1859), p. 254. — 
Smith, Publ. Field Mus, Nat. Hist., zool. ser., vol. 26, 1939, pp. 240-246, 
pi. 19, fig. 43. 

Lysoptychus lateralis Cope, Proc. U. S. Nat. Mus., vol. 11, 1888, pp. 397-398, 
pi. 26, fig. 1 (San Diego, Duval Countv, Tex.; U. S. N. M. No. 14741). 

Sceloporus lateralis, Boulenger, Proc. Zool. Soc. London, 1890, p. 78. 

Type. — U.S.N.M. No. 2739 (9 specimens), cotypes; Lt. Darius 
Nash Couch collector. 

Type locality. — Santa Catarina, Nucvo Le6n (specimens collected 
by Lt. Couch were also at hand from ''Pesqueria Grande," Mexico). 

Range. — Central and northern Nuevo Le6n and eastern Coahuila. 
In the United States, reported from southern Texas.^^ 

MERRIAMI GROUP 

Species. — One, with two subspecies. 

Range. — Southwestern Texas and adjacent Coahuila, 

KEY TO SPECIES OF THE MERRIAMI GROUP OF SCELOPORUS 

1. Anterior section of frontal usually divided; frontoparietals usually divided; 
outer row of labiomental scales rarely terminating with the first scale wedged 
between first infralabial and first postmental; head scales rugose; sub- 
caudal surface distinctly banded; gular bars extensive, confluent medially. 

merriami annulatus (p. 133) 
Anterior section of frontal rarely divided; frontoparietals rarely divided; outer 
row of labiomentals terminating with the first scale wedged between first 
infralabial and first postmental; head scales smooth; subcaudal surface 
nearl}' or quite immaculate, not barred; gular bars short, usually separate 
medially merriami merriami (p. 132) 

SCELOPORUS MERRIAMI MERRIAMI Stejneger 

Sceloporus merriami Stejneger, Proc. Biol. Soc. Washington, vol. 17, 1904, pp. 

17-20. 
Sceloporus merriami merriami. Smith, Proc. Biol. Soc. Washington, vol. 50, 1937, 

pp. 83-86. 

Type.— v. S.'N.M. No. 33039; W. Lloyd collector. 

Type locality. — East Painted Cave, near mouth of the Pecos River, 
Tex. 

Range. — The Rio Grande and its immediate tributaries from western 
Brewster County to southeastern Val Verde County, Tex. The species 
occurs on the bluffs along the Mexican side of the river, but no specific 
records for either Coahuila or Chihuahua (in both of which states it 
should occur) are known. 



" Except possibly for the type specimen of Lysoptychus lateralis, no specimen of S. couchii has ever, to our 
knowledge, been taken in Texas. We consider it doubtful that the specimen came from there. 



CHECKLIST OF REPTILES OF MEXICO 133 

SCELOPORUS MERRIAMI ANNULATUS Smith 

Sceloporus merriami annulatus Smith, Proc. Biol. Soc. Washington, vol. 50, 1937, 
pp. 83-86; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 289-290, 
pi. 1. 

Type.— EUT-HMS No. A787; E. H. Taylor and J. S. Wright 
collectors. 

Type locality. — East slope of the Chisos Mountains, Brewster 
County, Tex. 

Range. — Southern and central Brewster County, Tex., and eastern 
Coahuila (Cuatro Cienegas, Jaral). 

MACULOSUS GROUP 

Species. — One, 

Range. — Eastern Durango. 

SCELOPORUS MACULOSUS Smith 

Scelo-porus maculosus Smith, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 267-269, 
pi. 8, figs. 2, 4, 5, pi. 10, fig. 11; Publ. Field Mus. Nat. Hist., zool. ser., vol. 
26, 1939, pp. 291-294, fig. 47. 

T?/2)e.-EHT-HMS No. 7638; E. H. Taylor and H. M. Smith 
collectors. 

Type locality. — Fourteen miles northeast of Pedricena, Durango. 
Range. — ICnown only from east-central Durango. 

CHRYSOSTICTUS GROUP 

Species. —One. 

Range. — Yucatan Peninsula. 

SCELOPORUS CHRYSOSTICTUS Cope 

Sceloporus chrysostictus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866 (1867), 
p. 125.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
295-300, pi. 24. 

Type.— U.S.N.M. No. 24865, six cotypes. 
Type locality. — Yucatan, here restricted to Chichen Itzk. 
Range. — The Yucatan Peninsula, and the area at its eastern base. 
Recorded in Mexico from Y'ucatan, Campeche, and Quintana Roo. 

SINIFERUS GROUP 

Species. — Four, one of which includes two subspecies. 
Range. — Pacific slopes from Guerrero to Nicaragua. 

KEY TO SPECIES OF THE SINIFERUS GROUP OF SCELOPORUS 

1 . Nasals and anterior internasals broadly in contact with rostral ; no postrostrals ; 

postanals enlarged in males; ventral scales notched ochoterenai (p. 135) 

Nasals and anterior internasals separated from rostral by two or more postros- 
trals; ventral scales not notched 2 



134 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

2. One canthal 3 

Two canthals 4 

3. Femoral pores 11-12; dorsal scales 38-44 carinatus (p. 135) 

Femoral pores 3-6; dorsal scales 28-37 squamosus (p. 135) 

4. Femoral pores 3-11; postanals not or slightly enlarged in males. 

siniferus siniferus (p. 134) 
Femoral pores 12-14; postanals distinctly enlarged in males. 

siniferus cupreus (p. 134) 

SCELOPORUS SINIFERUS SINIFERUS Cope 

Sceloporus siniferus Cope, Proc. Amer. Philos. Soc, vol. 11, 1869, pp. 159-160, 

161.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 313- 

319 (part), pi. 25. 
Sceloporus humeralis Bocourt, Ann. Sci. Nat. zool., ser. 5, vol. 17, No. 10, 1873, 

p. 2 (Mus. Hist. Nat. Paris; Oaxaca, here restricted to the city of Tehuante- 

pec). 

r2/i?e.— U.S.N .M. Nos. 30453-30471, co types. 

Type locality. — Pacific side of the Isthmus of Tehuantepec, here 
restricted to Tehuantepec (city), Oaxaca. 

Range. — Pacific slopes from western Guerrero to extreme western 
Guatemala, and inland to Morelos. Recorded in Mexico from 
Guerrero, Morelos, Oaxaca, and Chiapas. 

SCELOPORUS SINIFERUS CUPREUS Bocourt 80 

Sceloporus cupreus BocotJRT, Ann. Sci. Nat., Zool., ser. 5, vol. 19, No. 4, 1873, p. 3; 

Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 4, 1874, 

pp. 210-212, pi. 18 bis, fig. 2, 2a-b.— Smith, Occ. Pap. Mus. Zool. Univ. 

Michigan, No. 358, 1937, pp. 6-9; Publ. Field Mus. Nat. Hist., zool. ser., 

vol. 26, 1939, pp. 305-308. 
Sceloporus cochranae Smith, Proc. Biol. Soc. Washington, vol. 49, 1936, pp. 87- 

89, pi. 2 (Mount Zempoaltepec, Oaxaca; U.S.N. M, No. 47605). 
Sceloporus siniferus, Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 

p. 317 (part). 

Type. — Mus. Hist. Nat. Paris; Adolphe Boucard collector. 

" It is with some doubt that the name cupreus is applied to the highland form of siniferus. This was 
discussed in some detail previously (Publ Field Mus. Nat. Hist., zool, ser., vol. 29, 1939, p. 317). Speci- 
mens then available showed a higher femoral pore count for the types of cupreus and of cochranae (the latter 
from Mount Zempoaltepec, Oaxaca), for which the name cupreus was used, than for the few specimens 
referred to siniferus then available from the vicinity of Oaxaca City. However, new material from the 
latter region (including EHT-HMS 19195-19202, 19479-80) reveals a variation from 38 to 50 (av. 44.2) in 19 
counts of dorsal scales, and from 6 to 15 (av. 9.4) in 40 counts of femoral pores. 

From these data it is apparent that the highland population oi siniferus is recognizably different from the 
typical subspecies and deserves a name. Handicapped by lack of adequate material from Mount Zem- 
poaltepec and its environs, we find it diflBcult to determine whether the population from that area is the same 
as that from the Oixaca region, since the single specimen available has enlarged postanals and broad dor- 
solateral light stripes. Typically siniferus (including Oaxaca specimens) lacks enlarged postanals; none 
has been observed with them well developed. Likewise, distinct and broad dorsolateral stripes do not 
occur in male siniferus, although they are present in females. None of the other characters previously thought 
to be peculiar to the Mount Zempoaltepec specimen are actually so, as shown by data on the Oaxaca series. 
Moreover, there is some slight variation in the size of the basal caudal scales and in the width and distinct- 
ness of the dorsolateral light stripes in males. In view of this variability in the only two characters defining 
the Mount Zempoaltepec specimen (cupreus, sensu stricto), we believe it best under the circumstances to 
extend the concept of this name to include the Oaxaca specimens; it should then, of course, be treated as a 
subspecies, if the present interpretation of the name is correct. 



CHECKLIST OF REPTILES OF MEXICO 135 

Type locality. — Oaxaca, here restricted to Mount Zempoaltepec. 
Range. — Highlands of central Oaxaca. 

SCELOPORUS SQUAMOSUS Bocourt 

Sceloporus squamosvs Bocourt, Mission scientifique au Mexique . . ., fitudes 

sur les reptiles, livr. 4, 1874, pp. 212-214, pi. 18 bis, figs. 7, 7a-c, pi. 19, fig. 

3.— Smith, Publ. Field Mus. Nat, Hist., zool. ser., vol. 26, 1939, pp. 319-324, 

fig. 52, pi. 24. 
Sceloporus fulvus Bocoxjrt, ibid., pp. 214-215, pi. 18 bis, figs. 8, 8a-c (La Uni6n, 

El Salvador; Commission Scientifique collector; Mus. Hist. Nat. Paris). 

Type. — Mus. Hist. Nat. Paris; collected by (?) Commission Scien- 
tifique du Mexique. 

Type locality. — ^"Environs de Guatemala et de TAntigua" [=Volcan 
Antigua], 1,500 meters. 

Range. — Pacific slopes from eastern Chiapas to Costa Rica. Re- 
corded only from Chiapas in Mexico. 

SCELOPORUS CARINATUS Smith 

Sceloporus carinatus Smith, Proc. Biol. Soc. Washington, vol. 49, 1936, pp. 89- 
91, pi. 2, figs. 2, 3; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 303-305. 

Type.— ERT-RMS No. 15205 (EHT-HMS field No. 4866); E. H. 
Taylor and H. M. Smith collectors. 

Type locality. — Near Tuxtla Gutierrez, Chiapas. 
Range. — Plateau of Chiapas. 

SCELOPORUS OCHOTERENAI Smith 

Sceloporus ochoterenae Smith, Trans. Kansas Acad. Sci., vol. 37, 1934, pp. 269- 
272, pi. 9, figs. 6, 9, pi. 10, fig. 12; Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 26, 1939, pp. 308-313, text fig. 51. 

Type.— EHT-HMS No. 7158 (EHT field No. 1075); E. H. Taylor 
and H. M. Smith collectors. 

Type locality. — Two miles north of Mazatl^n, Guerrero (12 miles 
south of Chilpancingo, Guerrero). 

Range. — Morelos and Guerrero. 

UTIFORMIS GROUP 

Species. — One. 

Range. — Western Alexico, Sinaloa to Guerrero. 

SCELOPORUS UTIFORMIS Cope 

Sceloporus utiformis Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 177. — 
Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 325-330, 
fig. 53, pi. 26. 

Type.— U.S. ]<i.M. No. 42089; John Xantus collector. 
Type locality. — Near Colima, Colima. 



136 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Range. — Pacific slopes from southern Sinaloa to western Guerrero, 
inland about 125 miles in the southern part of its range. Recorded 
only from Sinaloa, Nayarit, Jalisco, Colima, Alichoacan, and Guerrero, 

SCALARIS GROUP 

Species. — Seven forms are recognized at present, representing four 
species. 

Bange. — High elevations from southeastern Arizona and central 
Nuevo Le6n southward to the edge of the plateau in Oaxaca. 

KEY TO SPECIES OF THE SCALARIS GROUP OF SCELOPORUS 

1. Nasals and internasals in contact with rostral; no postrostrals.jalapae (p. 136) 
Nasals and internasals separated from rostral by two postrostrals 2 

2. Dorsal scales 50 or more; lateral scales in slightly, although distinctly, oblique 

rows goldmani (p. 137) 

Dorsal scales less than 50; lateral scales in parallel rows 3 

3. One canthal 4 

Two canthals, the first occasionally forced above canthal ridge by contact of 

second canthal and subnasal 6 

4. Males with much black in ventral coloration; females suffused with black 

below; gular region never barred; black shoulder spot with a light blue spot, 
if present, on its anterior edge; tail with a continuous dark median dorsal 
stripe; tibia/head proportion usually less than 0.95; maximum snout-vent 

measurement 58 mm aeneus aeneus (p. 137) 

Black, if present on ventral surface, confined to bars in gular region, and a few 
dark, transverse bars on sides of abdomen; black shoulder spot with the light 
blue spot in its middle 5 

5. Tibia/head proportion usually less than 0.90; scales of second pair of post- 

mentals separated medially; dorsal scales usually more than 40; maximum 

snout-vent measurement 61 mm scalaris slevini (p. 138) 

Tibia/head proportion usually more than 0.90; scales of second pair of post- 
mentals usually in contact medially; dorsal scales usually less than 40; 
maximum snout-vent measurement 65 mm.. scalaris unicanthalis (p. 138) 

6. Males with much black in ventral coloration; black shoulder spot with the 

light blue spot, if present, on its anterior edge; tail with a continuous dark 
median dorsal stripe; tibia/head proportion usually less than 0.90; maximum 

snout-vent measurement 56 mm aeneus bicanthalis (p. 137) 

Black, if present on the ventral surface, confined to bars in gular region, and a 
few dark transverse bars on sides of abdomen; black shoulder spot with the 
light blue spot in its middle; tail with dark chevron-shaped bars; tibia/head 
proportion usually more than 0.90; maximum snout-vent measurement 
78 mm scalaris scalaris (p. 137) 

SCELOPORUS JALAPAE GUnther 

Sceloporus jalapae Gtjntiier, Biologia Centrali-Americana, Reptilia and Batra- 
chia, 1890, p. 74. — Smith, Proc. Biol. Soc. Washington, vol. 47, 1934, pp. 
121-125, fig. 1; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
333-338, figs, 55, 56, pi. 27. 

Type. — Brit. Mus. Nat. Hist.; Mr. Hoge collector. 
Type locality. — Jalapa, Veracruz. 



CHECKLIST OF REPTILES OF MEXICO 137 

Range. — Central Veracruz south through eastern Puebla to central 
Oaxaca. Recorded only from the states cited. 

SCELOPORUS AENEUS AENEUS Wiegmann 

Sceloporus aeneus Wiegmann, Isis von Oken, vol. 21, 1828, p. 370. 

Sceloporus aeneus aeneus, Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 

1937, p. 6; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 353-356, 

text fig. 57. 

Type.- — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, by inference, here restricted to Tres 
Cumbres. 

Range. — Western Puebla, to central western Michoacan, and 
Jalisco; north on the plateau to northern Guanajuato. Recorded 
from the states cited and from Distrito Federal, Mexico. Morelos, 
and questionably, Jalisco. 

SCELOPORUS AENEUS BICANTHALIS Smith 

Sceloporus aeneus bicanthalis Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
361, 1937, pp. 6-8; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 356-361, fig. 57, pi. 20. 

Type.— EJiT-HMS No. 7939; E. H. Taylor, collector. 

Type locality. — Cofre de Perote, near Cruz Blanca, Veracruz. 

Range. — From northern Hidalgo along the eastern escarpment of 
the plateau to central Oaxaca, including parts of Puebla, Mexico, 
and Veracruz, Recorded only from the states cited. 

SCELOPORUS GOLDMANI Smith 

Sceloporus goldmani Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 1937, 
p. 5; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 350-353. 

Type.— UniY. Michigan Mus. Zool. No. 80896; C. L. Lundell 
collector. 

Type locality. — Charcas, San Luis Potosi. 

Range. — Southern Coahuila southward in the central part of the 
plateau of Mexico to central San Luis Potosi. Recorded only from 
the states cited. 

SCELOPORUS SCALARIS SCALARIS Wiegmann 

Sceloporus scalaris Wiegmann, Isis von Oken, vcl. 21, 1828, p. 370. 

Sceloporus scalaris scalaris, Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 

1937, pp. 2-3; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 338- 

343, pi. 20. 

Type. — Zool. Mus. Berlin; F. Deppe collector. 
Type locality . -—^lexico , here restricted to Mexico, D. F. 
Range. — Central Durango, southward over the central and southern 
parts of the Mexican Plateau except (apparently) Guerrero. Re- 



138 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

corded from Distrito Federal and the states of Durango, Guanajuato, 
Hidalgo, Jalisco, Mexico, Michoacan, Puebla, and Zacatecas. 

SCELOPORUS SCALARIS UNICANTHALIS Smith 

Sceloporus scalaris unicanthalis Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No- 
361, 1937, pp. 4-5; Publ. Field Mus. Nat. Hist. zool. ser., vol. 26, 1939, 
pp. 349-350. 

T^/pe.— EHT-HMS No. 7699; H. M. Smith coUector. 

Type locality.- — Magdalena, Jalisco. 

Range. — Known only from the southern border of the Mexican 
plateau in central Jalisco, from the eastern border of Lake Chapala 
westward to Nayarit. Recorded only from the state of Jalisco. 

SCELOPORUS SCALARIS SLEVINI Smith 

Sceloporus scalaris slevini Smith, Occ. Pap. Mus. Zool. Univ. Michigan, No. 361, 
1937, pp. 3-4; Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
343-349. 

Type.— Cfilii. Acad. Sci. No. 48013; J. E. Slevin collector. 

Type locality. — Miller Peak, Huachuca Mountains, Cochise County, 
Ariz. 

Range. — Southern Arizona south to northern Durango, and east to 
to Nuevo Le6n in Mexico. Recorded from the states cited and from 
Sonora, Chihuahua, and Coahuila. 

PYROCEPHALUS GROUP 

Species. — Three. 

Range. — Pacific slopes from southwestern Chihuahua to north- 
western Oaxaca. 

KEY TO SPECIES OF THE PYROCEPHALUS GROUP OP SCELOPORUS 

1, Scales on posterior surface of thigh granular; postfemoral dermal pocket 

present gadoviae (p. 138) 

Scales on posterior surface of thigh larger, imbricate; no postfemoral dermal 
pocket 2 

2. Dorsal scales larger, 36-41; females not red-headed; males with unbroken 

lateral beUy patches; dark spot on interparietal not enclosing or touching 
light pineal spot — nelsoni (p. 139) 

Dorsal scales smaller, 41-50; females red-headed, conspicuously barred below 
on throat; males (and usually females) with a series of broad dark bars on 
each side of belly; a dark spot surrounding light pineal spot. 

pyrocephalus (p. 139) 

SCELOPORUS GADOVIAE Boulenger 

Sceloporus gadoviae Boulenger, Proc. Zool. Soc. London, 1905, vol. 2, pp. 246- 
247, pi. 7, fig. 1.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, 
pp. 373-377, fig. 53, pi. 31. 

Type.— Brit. Mus. Nat. Hist.; Hans Gadow collector. 

Type Zoca^i^?/. — Mezquititlan, north of Chilpancingo, Guerrero. 



CHECKLIST OF REPTILES OF MEXICO 139 

Range. — Southern Michoacan through Guerrero, Morelos, southern 
Puebla to northern and western Oaxaca. Recorded only from the 
states cited. 

SCELOPORUS PYROCEPHALUS Cope 

Sceloporris pyrocephalus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 177. — 
Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 368-373, 
pis. 29, 30. 

Sceloporus pyrrhocephalus Cope, Proc. Ainer. Philos. Soc, vol. 22, 1885, pp. 
394-397 (emendation). 

Type.—V.S.'N.M. No. 31495; John Xantus collector. 
Type locality. — Near Colima, Colima. 

Range. — Central Jalisco, south and east through Colima and 
Michoacan to Guerrero. Recorded only from states cited. 

SCELOPORUS NELSONI Cochran 

Sceloporus nelsoni Cochran, Journ. Washington Acad. Sci., vol. 13, 1923, pp. 
185-186.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 26, 1939, pp. 
364-368, pi. 28. 

Type.— U.S. ISl.M. No. 47676; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — Plomosas, Sinaloa. 

Range — Pacific slopes from southwestern Chihuahua to northwestern 
Jalisco. Recorded from Chihuahua, Sonora, Sinaloa, Nayarit, and 
Jalisco. 

Genus SATOR Dickerson 

Sator Dickerson, Bull. Anaer. Mus. Nat. Hist., vol. 41, 1919, pp. 468-469. 

Genotype. — Sator grandaevus Dickerson. 

Range. — Santa Cruz and Cerralvo Islands in the Gulf of California. 

Species. — Two. 

KEY TO SPECIES OF SATOR 

1. A poorly defined anterior gular fold, without granules; a posterior (true) gular 
fold immediately in front of arm insertions, but not continuous across throat; 
median lateral scales very small, often granular, abruptly differentiated from 

dorsals and ventrals grandaevus (p. 140) 

A well defined anterior gular fold, with granules; no evidence whatever, even 
at sides in front of arm insertions, of a posterior gular fold; median lateral 
scales larger, gradually merging with dorsals and ventrals. axigustus (p. 139) 

SATOR ANGUSTUS Dickerson 

Sator nngustus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 469. — 
Schmidt, BuU. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 665-666, fig. 10.— 
Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 256-258. 

Type. — Amer. Mus. Nat. Hist. No. 5712; C. H. Townsend collector. 
Type locality. — Santa Cruz Island, Gulf of California, Baja Cali- 
fornia. 

Range. — Restricted to the type locality. 

861316 — 50 10 



140 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

SATOR GRANDAEVUS Dickerson 

Sator grandaevus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 
469-470.— Schmidt, ibid., vol. 46, 1922, p. 665, figs. 8, 9.— Van Denburgh, 
Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 259-261. 

Type. — Amer. Mus. Nat. Hist. No. 5491; C. H. Townsend collector. 
Type locality. — Cerralvo Island, Gulf of California, Baja California. 
Range. — Restricted to the type locality. 

Genus UROSAURUS Hallowell 

Urosaurus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 1854, p. 92. — 
MiTTLEMAN, Bull. Mus. Comp. ZooL, vol. 91, 1942, pp. 103-181, pis. 1-16. 

Phymatolepis Dumeril, Arch. Mus. Hist. Nat. Paris, vol. 8, 1856, p. 548 (type, 
Phymatolepis bicarinatus Dumeril). 

Genotype. — Urosaurus graciosus Hallowell. 

Range. — Southern Texas west to California, entering Utah and 
Colorado, southward through Baja California and many adjacent 
islands, and through mainland Mexico from the western Texas border 
south to the plateau and along the western slopes to Chiapas. 

Species. — Ten species and 21 forms are recognized; all 10 species 
and 19 forms are laio\vn to occur or are to be expected in Mexico. 

KEY TO MEXICAN 8i SPECIES OF UROSAURUS 

1. Enlarged anterodorsal femoral scales smooth auriculatus (p. 146) 

Enlarged anterodorsal femoral scales strongly keeled 2 

2. Enlarged dorsals in a single broad band, uninterrupted by an intervening 

series of smaller scales 3 

Enlarged dorsals separated into two or more parallel series by the presence of 
a vertebral series of smaller scales 7 

3. Tail two or more times length of head and body combined. graciosus (p. 144) 
Tail less than twice length of head and body combined 4 

4. Dermal folds, when present, not heavilj'' crested with tubercles; blue abdominal 

patches only in males; enlarged dorsals comparatively smaU 5 

Dermal folds present, always crested with tubercles of fairly large size; 
abdominal blue patches sometimes in females as well as males 6 

5. Enlarged dorsals larger, from 17 to 24 in length of head from tip of snout to 

posterior edge of interparietal; gular region in males deep yellow or orange. 

nigricaudus (p. 145) 
Enlarged dorsals smaller, 32 to 36 in length of head from tip of snout to 
posterior edge of interparietal; gular region in males usually blue. 

microscutatus (p. 145) 

6. Four to seven rows of enlarged dorsal scales; abdomen of both sexes with a 

blue wash and/or blue patches; dorsolateral folds not converging in sacral 

region gadovi (p. 145) 

About three rows of enlarged dorsal scales; only males with a blue abdomen; 
dorsolateral folds converging in the sacral region to form prominent ridges. 

irregularis (p. 146) 

7. Three or fewer rows of lateral tubercles 12 

Four or more rows of lateral tubercles 8 



" Adapted from Mittleman, op. cit., pp. 127-133. 



CHECKLIST OF REPTILES OF MEXICO 141 

8. Enlarged dorsals commencing caudad of a line joining the anterior points of 

insertion of the forelimbs; dorsals weakly keeled, rounded posteriorly, 
prominently pavemented; general habitus not at all rugose .unicus (p. 147) 
Enlarged dorsals commencing craniad of a line joining the anterior points of 
insertion of the forelimbs, or else equal with such a line; dorsals prominently 
keeled, usually mucronate or spinose, imbricate; ventrals imbricate; general 
appearance rugose 9 

9. Form rugose; enlarged dorsals strongly carinate and prominently mucronate- 

tubercles of lateral and dorsolateral folds well developed; ventrals mucro- 
nate; gular surface generally stippled, with a light median area; blue ab_ 
dominal patches of males quite extensive, .bicarinatus bicarinatus (p. 146) 
General appearance somewhat less rugose; enlarged dorsals not so often 
mucronate; tubercles smaller, sometimes absent; ventrals less mucronate, 
occasionally rounded; gular surfaces evenly stippled; abdominal blue of 
males sometimes restricted to small sternal patches 10 

10. Ventrals rounded; dorsolateral and lateral tubercles very poorly developed; en- 

larged dorsals commencing on the nape bicarinatus nelsoni (p. 147) 

Ventrals submucronate to mucronate; dorsolateral and lateral tubercles well 
developed; enlarged dorsals commencing on the shoulders just craniad of a 
line joining the anterior points of insertion of the forelimbs 11 

11. Ventrals mucronate, prominently carinated laterally; gular scales with a 

tendency toward pavementation, especially anteriorly; gular surfaces evenly 
stippled; abdominal blue of males restricted to small sternal patches. 

bicarinatus anonymorphus (p. 146) 

Ventrals submucronate (occasionally rounded), only faintly keeled on the 

lateral portions of the belly, or else not at all; gular scales imbricate; gular 

region \\'ith an even blue wash, and only barely flecked if at all; abdominal 

blue of males evenly distributed bicarinatus tuberculatus (p. 147) 

12. Enlarged dorsals commencing on nape 13 

Enlarged dorsals commencing on shoulders or caudad of them 14 

13. Enlarged dorsals strongly keeled; scales of primary and secondary series 

almost equal in size; postfemoral dermal pocket absent or rudimentary; no 

prominent lateral pattern of dark whorls clarionensis (p. 142) 

Enlarged dorsals not so rugose, scales of primary series prominently larger 
than those of the secondary series; postfemoral dermal pocket regularly 
present; a distinct lateral pattern of dark whorls. omatus schottii (p. 144) 

14. Tail two or more times length of head and body combined. graciosus (p. 144) 
Tail less than twice length of head and body combined. 15 

15. Enlarged dorsals extending onto basal portion of tail for a distance equal to 

length of femur, or more; entire gular region, including sublabials, a uniform 

bright blue omatus caeruleus (p. 143) 

Enlarged dorsals extending onto basal portion of tail for a distance equal to less 
than length of femur; entire gular region including sublabials never com- 
pletely blue 16 

16. Enlarged dorsals often irregularly arranged; lateral tubercles not afifecting 

diagonal arrangements; average size less than 45 mm. from snout to 

vent 17 

Enlarged dorsals regularly arranged in parallel series on either side of the 
vertebrals; tubercles in parallel diagonal series; average size greater than 45 
mm. from snout to vent 18 



142 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

17. Scales of primary series not twice as large as those of the secondary series; 

largest of the dorsals inferior in size to enlarged femorals and tibials; ventral 
interhumeral and interfemoral areas immaculate, or but slightly stip- 
pled ornatus schmidti (p. 143) 

Scales of primary series almost twice as large as those of secondary series; 
largest of dorsals equal to, or larger than, enlarged femoral and tibial scales; 
ventral interhumeral and interfemoral areas heavily maculated. 

ornatus ornatus (p. 142) 

18. Largest of dorsals equal to, or larger than enlarged femorals; vertebrals extend- 

ing onto basal portion of tail for a distance equal to half, or slightly more, of 
length of femur; entire gular region in males, except sublabials, an intense 
blue; head length/head width ratio averaging 81 percent. 

ornatus chiricahuae (p. 143) 

Largest of dorsals inferior in size to enlarged femorals; vertebrals extending 

only onto rump, or but slightly farther; no uniform intense blue color present 

in male gular region; head length/head width ratio averaging 75 percent or 

less 19 

19. Enlarged dorsals separated into 2 parallel series by width of vertebral series, 

which is greater in width than broadest of enlarged dorsals; prefrontals and 
frontonasals usually 3 each; general coloration pallid, light tan above, 
whitish below, males with bright-blue abdominal patches; average head 
length/width ratio 75.4 percent; average length, snout to vent, 55.1 mm. 

ornatus symmetricus (p. 144) 
Enlarged dorsals separated by a vertebral series whose width is less than that 
of largest of dorsal scales; prefrontals 2, rarely 3 (by inclusion of an azy- 
gous); frontonasals 5; general color variable, but usually dark brown or 
gray with dark cross bands, and heavily stippled, spotted, or blotched ven- 
trally; abdominal patches in males dark blue to indigo; average head 
length/head width ratio 70.6 percent; average length, snout to vent, 46.4 mm. 

ornatus linearis (p. 143) 

ORNATUS GROUP 

Species. — Three species and 11 forms are included; all but two 
subspecies of ornatus occur or may be expected to occur in Mexico. 

Range. — Utah, central Texas, and southern Nevada southward to 
extreme northern Mexico, reaching farther southward only along the 
coast to southern Sinaloa; Clarion Island. 

UROSAURUS CLARIONENSIS (Townsend) 

Uta clarionensis Townsend, Proc. U. S. Nat. Mus., vol. 13, 1890, p. 143. 
Urosaurus clarionensis, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, p. 
149, pi. 8 (type). 

3Vpe.— U.S.N.M. No. 15904; C. H. Townsend collector. 
Type locality. — Clarion Island, Revillagigedo Islands, Mexico. 
Range. — Known only from the type locality (administered by 
Colima) . 

UROSAURUS ORNATUS ORNATUS (Baird and Girard) 

Uta ornata (part) Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 

1852, p. 126. 
Uta ornata ornata (part), Schmidt, Amer. Mus. Nov., No. 22, 1921, p. 6. 
Urosaurus ornatus ornatus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 

pp. 133-135. pi. 1 (cotypes). — Smith, Handbook of lizards, 1946, pp. 264- 

266, pi. 65. 



CHECKLIST OF REPTILES OF MEXICO 143 

Type. — U.S.N.M. No. 2750, two cotypes, male and female. 

Type locality. — Restricted to Rio San Pedro [=Devils River], Val 
Verde County, Tex. 

Range. — Central and southern Texas, south to northern Coahuila. 
Has been taken along the Rio Grande River on the American side, 
and very probably occurs in Coahuila. 

UROSAURUS ORNATUS CAERULEUS (Smith) 

Uta caerulea Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 172-178, pi. 26. 
Urosaurus ornatus caeruleus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 136-137, pi. 9, lower fig. 

Type.— Vniv. Kansas Mus. Nat. Hist., No. 19237 (Smith and Dun- 
kle field No. 132); Hobart M. Smith and David H. Dunkle collectors. 

Type locality. — Thirty mUes north of Chihuahua City, Chihuahua. 

Range. — Central Chihuahua. Reported only from the type locality 
and from 20 miles south of Chihuahua. 

UROSAURUS ORNATUS CHIRICAHUAE (Mittleman) 

Uta ornata chiricahuae Mittleman, Proc. Biol. Soc. Washington, vol. 54, 1941, 

p. 165. 
Urosaurus ornatus chiricahuae, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 

1942, pp. 139-142.— Smith, Handbook of lizards, 1946, p. 266, pi. 66. 

Type.— Mus. Vert. Zool. Univ. California, No. 7751. 

Type locality. — Pinery Canyon, Chiricahua Mountains, 6,000 feet, 
Cochise Co., Ariz. 

Range. — So far as known, restricted to the Chiricahua and Dos 
Cabezas Mountains, Ariz. ; of probable occurrence in Mexico. 

UROSAURUS ORNATUS LINEARIS (Baird) 

Uta ornata var. linearis Baird, United States and Mexican boundary survey, 

1859, vol. 2, pt. 2, p. 7. 
Uta ornata linearis, Schmidt, Amer. Mus. Nov., No. 22, 1921, p. 6. 
Urosaurus ornatus linearis, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 137, 139, pi. 3 (neotype).— Smith, Handbook of lizards, 1946, pp. 268-271, 
pi. 67. 
Type.— U.S.N.M. No. 2759 (now lost); Caleb Kennerly collector. 
Neotype: U.S.N.M. No. 62077, Los Nogales, Sonora, Mexico; F. J. 
Dyer, collector. 

Tijpe locality. — Los Nogales, Sonora, Mexico. 

i?a??^e.— Southern Aiizona and southern New Mexico, southward 
to northern Sonora and Chihuahua. 

UROSAURUS ORNATUS SCHMIDT! (Mittleman) 

Uta ornata schmidti Mittleman, Herpetologica, vol. 2, pt. 2, 1940, pp. 33-34, 

pi. 3, fig. 1 (1941). 
Urosaurus ornatus schmidti, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 

pp. 135-136, pi. 2 (type).— Smith, Handbook of lizards, 1946, pp. 271-272, 

pi. 68). 



144 BULLETESr 19 9, UNITED STATES NATIONAL MUSEUM 

Type.—IJ.S.'NM. No. 32929; V. Bailey collector. 
Type locality. — Fort Davis, Jeff Davis County, Tex. 
Range. — Southvvestorn Texas and northern Chihuahua. Reported 
in Mexico only from Chihuahua: Samalayuca. 

UROSAURUS ORNATUS SCHOTTII 8S (Baird) 

Uta schottii Baird, Proc. Acad. Nat. Sci. Philadelphia, vol. 10, 1858, p. 253. 

Uta ornata schottii, Mittleman, Copeia, 1941, pp. 136-138. 

Urosaurus ornatus schottii, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 

pp. 149-151. 
Uta {Phymatole'pis) lateralis Boulenger, Ann. Mag. Nat. Hist., ser. 5, vol. 11, 

1883, p. 342 (Brit. Mus. Nat. Hist.; type locality restricted to Presidio, 

Sinaloa). 
Uta lateralis, Boulenger, Catalogue of the lizards of the British Museum, vol. 

2, 1885, p. 214. 
Uta ornata lateralis, Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 

1922, pp. 199-201.— Oliver, Copeia, 1943, pp. 97-107, 1 fig. 
Uta gularis Cragin, Bull. Washburn Lab. Nat. Hist., vol. 1, 1884, p. 7 (type 

now lost, formerly in Washburn College collection, Topeka, Kans.; Guay- 

mas, Sonora; Prof. Lovewell collector). 

Type.— V.S.N. M. No. 2761 (now lost); A. Schott collector. 

Type localHy.—"Sisi. Madelina, Cal.-Mex. Boundary survey"= 
?Magdalena, bonora, Mexico. 

Range. — Central Sonora and southward to southern Sinaloa, and 
the Tres Marias Islands; Tibur6n Island. 

UROSAURUS ORNATUS SYMMETRICUS (Baird) 

Uta symmetrica Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 253. 
Uta ornata symmetrica, Schmidt, Amer. Mus. Nov., No. 22, 1921, p. 6. 
Urosaurus ornatus symmetricus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 

1942, pp. 142-144, pi. 4 (neotype). — Smith, Handbook of lizards, 1946, 

pp. 273-274, pi. 69. 

Type.— {]. S.N. M. No. 2760 (lost). Neotype: U.S.N.M. No. 2744; 
Fort Yuma, Imperial Count}'", Calif.; M. Thomas collector. 

Type locality. — Fort Yuma, Imperial County, Calif. 

Range. — In United States: southern California and adjacent parts 
of Arizona south to western Sonora and northern Baja California. 
Reported from Sonora: 2 miles south of Nogales, Pinetos Camp, 
32 miles south of Nogales, Duros Millos, Gran Desierto; Baja 
California: "Colorado River valley and desert to Delta." 

urosaurus GRACIOSUS Hallowell 

Uro-saurus graciosus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 7, 
1854, p. 92.— Smith, Handbook of lizards, 1946, pp. 259-262, pi. 63. 

Uta graciosa, Baird, United States and Mexican boundary survey, 1859, vol. 2, 
pt. 2, p. 7. — Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, 
pp. 212-216, pi. 17. 

M The status of this form is questioned by Oliver, loc. cit., who feels that the name should be Uta ornata 
lateralis Boulenger. 



CHECKLIST OF REPTILES OF MEXICO 145 

Urosaurus ornatus graciosus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 144-145, pi. 7 (cotypes). 

Type.— Ac2i^. Nat. Sci. Philadelphia Nos. 8550-8551. 

Type locality. — "Lower California" [=southem California], here 
restricted to Winterhaven (=Fort Yuma), Calif. 

Range. — Southern Nevada, western Arizona, southern California, 
and northern Baja California. Reported from Baja California: 85 
miles south of Mexicali, San Felipe. It may occur also m Sonora but 
is not recorded from there. 

NIGRICAUDUS GROUP 

Species. — Four. 

Range. — Jalisco, Michoacan, and Baja California. 

UROSAURUS NIGRICAUDUS (Cope) 

Uia nigricauda Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 17(3.^ — Van Den- 
burgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 216-219. 

Urosaurus nigricaudus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 
157-159, pi. 10. 

Type. — U.S.N.M. No. 5307, 12 cotypes; John Xantus collector. 
Type locality. — Cape San Lucas, Baja California. 
Range. — Southern tip of Baja California, with adjacent coastal 
islands, Espiritu Santo, Ballena, San Jose, and ?Magdalena. 

UROSAURUS MICROSCUTATUS (Van Denbnrgh) 

Uta microscutata Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 4, 1894, 
p. 298; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 219-221. 

Urosaurus microscutatus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 
pp. 159-163.— Smith, Handbook of lizards, 1946, pp. 262-264, pi. 64. 

Uta parviscutata Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 324-325, 
fig. 45 (substitute name for Uta microscutata Van Denburgh). 

T^/pe.— Stanford Univ. Mus. No. 1221; J. M. Stowell collector. 

Type locality. — San Pedro Martir Mountains, Baja California. 

Range. — Extreme southern California through all of Baja California 
except the southern fifth. Reported from Baja California: Puerto 
Escondido, San Xavier, San Quintin, San Pedro Martir, Mount San 
Matias, and the Islands of San Francisco, San Jose, Danzante, Coro- 
nado, Carmen, San Marcos, Santa Magdalena. 

UROSAURUS GADOVI (Schmidt) 

Uta gadovi Schmidt, Amer. Mus. Nov., No. 22, 1921, pp. 3-4. 
Urosaurus gadovi, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 154-156, 
pi. 11. 

Type. — Amer. Mus. Nat. Hist. No. 20355; Hans Gadow collector. 
Type locality. — Cofradia, Jalisco, Mexico. 

Range. — Pacific slopes of Michoacan and Jalisco. Reported from 
Michoacan: Apatzingan, Acahuato; Jalisco: Cofradia. 



146 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

UROSAURUS IRREGULARIS (Fischer) 

Phymatolepis (Uta) irregularis Fischer, Abh. Nat. Ver. Bremen, vol. 7, 1882, pp. 

232-234, pi. 17, figs. 1-4. 
Uta irregularis, Boulenger, Catalogue of the lizards in the British Museum, 

ed. 2, vol. 2, 1885, p. 216. 
Urosaurus irregularis, Mittleman, Bull. Mus. Comp. ZooL, vol. 91, 1942, pp. 

156-157, pi. 12 (upper fig.). 

Type. — Municipal Nat. Hist. Coll. Bremen, Germany, No. 437; 
collector unlaiown. 

Type locality. — "Aus dem Hochlande von Mexico." 
Range. — Mexico highlands. No specific locality known. 

BICARINATUS GROUP 

Species. — Three, with a total of six forms. 

Range. — Pacific slopes from southwestern Chihuahua south to 
Chiapas; Socorro Island. 

UROSAURUS AURICULATUS (Cope) 

Uta auriculata Cope, Proc. Boston Soc. Hist., vol. 14, 1871, p. 303. — Van 
Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 37, 197-199, 

Urosaurus auriculatus, Mittleman, BuU. Mus. Comp. Zool., vol. 91, 1942, pp. 
163-164, pi. 16 (type). 

Type.— U.S. ISl.M. No. 7027; Grayson collector. 
Type locality. — Socorro Island, Revillagigedo Islands. 
Range. — KJnown only from the type locality. 

UROSAURUS BICARINATUS BICARINATUS (Dumferil) 

Phymatolepis bi-carinatus Dum^ril, Arch. Mus. Nat. Hist. Nat. Paris, vol. 8, 

1856, p. 549, pi. 23, figs. 2, 2a, 2b. 
Uta bicarinata, Cope, Proc. Acad. Nat. Sci. Philadelphia, vol. 16, 1864, p. 177. 
Uta bi-carinata bi-carinata, Mittleman, Journ. Washington Acad. Sci., vol. 31, 

1941, pp. 70-71, figs. IG, 2. 

Urosaurus bicarinaius bicarinatus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 

1942, pp. 164-166, pi. 13. 

Type. — Mus. Hist. Nat. Paris, two cotypes; A. Boucard collector. 

Type locality. — Mexico, here restricted to Cuernavaca, Morelos. 

Range. — Pacific slopes from Michoacan to central Guerrero, and 
up the basin of the Rio Balsas to southern Puebla. Reported from 
Puebla: "Puebla," Izucar de Matamoros, Tlapanala, Chiautla; 
Morelos: Cuernavaca; Guerrero: Agua del Obispo, Cocoyul, Chil- 
pancingo, Mesquititlan, Iguala, Rio Balsas, etc.; Michoacan: San 
Bias, Jorullo. 

UROSAURUS BICARINATUS ANONYMORPHUS (Mittleman) 

Uta anonymorpha Mittleman, Herpetologica, vol. 2, 1940, pp. 34-38, pi. 3, fig. 2. 
Uta bicarinata anonymorpha, Mittleman, Journ. Washington Acad. Sci., vol. 
31, 1941, pp. 71-72, figs. IG, 2. 



CHECKLIST OF REPTILES OF MEXICO 147 

Urosaurus bicarinatus anonymorphus, Mittleman, Bull. Mus. Comp. Zool., 
vol. 91, 1942, pp. 166-168, pi. 14 (type). 

ri/2>e.— U.S.N.M. No. 46988; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — Tehuantepec, Oaxaca. 

Range. — Pacific slopes in eastern Guerrero, Oaxaca and western 
Chiapas. Reported from Chiapas: Tonala; Oaxaca: Tehuantepec, 
Juchitan, San Ger6nimo, Ixtepec, Totolapam, Tres Cruces, Mount 
Guengola, Cerro Arenal, etc.; Guerrero: Tierra Colorada. 

UROSAURUS BICARINATUS NELSONI (Schmidt) 

Uta nelsoni Schmidt, Amer. Mus. Nov., No. 22, 1921, p. 4. 

Uta hicarinata nelsoni, Mittleman, Journ. Washington Acad. Sci., vol. 31, 1941, 

pp. 72-73, figs. IH, 2. 
Urosaurus bicarinatus nelsoni, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, 

p. 168, pi. 15. 

Type.— U.S. ISl.M. No. 46836; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — Cuicatlan, Oaxaca. 

Range. — Known only from the type locality. 

UROSAURUS BICARINATUS TUBERCULATUS (Schmidt) 

Uta tuberculata Schmidt, Amer. Mus. Nov., No. 22, 1921, p. 4. 

Uta bi-carinata tuberculata, Mittleman, Journ. Washington Acad. Sci., vol. 31, 

1941, pp. 73-74, figs. IE, 2.— Oliver, Copeia, 1943, p. 105. 

Urosaurus bicarinatus tuberculatus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 

1942, pp. 169-170, pi. 12, lower fig. 

Type.— Amer. Mus. Nat. Hist. No. 13737; Paul D. R. Riithling 
collector. 

Type locality. — Colima, Colima, Mexico. 

Range. — Southern Sonora southward to Jalisco. Sonora: Guirocoba; 
Sinaloa: Presidio de Mazatlan; Jalisco: ?"North of Rio Santiago"; 
Colima: Colima, Villa Alvarez, Paso del Rio. 

UROSAURUS UNICUS (Mittleman) 

Uta unica Mittleman, Journ. Washington Acad. Sci., vol. 31, 1941, pp. 74-76, 

fig. 3. 
Urosaurus unicus, Mittleman, Bull. Mus. Comp. Zool., vol. 91, 1942, pp. 170- 

173. — Smith and Mittleman, Trans. Kansas Acad. Sci., vol. 46, 1943, pp. 

243-249. 

Type.— U.S.N.M. No. 14248; Edward Wilkinson collector. 
Type locality. — Chihuahua (Batopilas?). 
Range. — Known only from the type locality. 

Genus UTA Baird and Girard 

Uta Baird and Girard, in Stansbury, Exploration and survey of the valley of the 
Great Salt Lake of Utah . . ., 1852, pp. 344-345. 



148 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Genotype. — Uta stansburiana Baird and Girard. 

Range. — Western Texas to California, north to Washington, south 
through Baja Cahfornia and adjacent islands, and into the northern 
tier of Mexican states from Sonora to Coahuila. 

Species. — Nine are here listed, with a total of eleven forms; only one 
form (U. s. stansburiana) is extralunital, making a total for the genus 
of 12 forms. The entire genus is badly in need of careful revision, 
however; considerable shuffling of the nominal species and subspecies 
is to be expected eventually. 

KEY TO MEXICAN SPECIES OF UTA 

1. Dorsal scales shorter, not imbricate, not mucronate, often with intervening 

granules; a dark blotch behind axilla 2 

Dorsal scales larger, imbricate at least centrally, usually without intervening 
granules; caudal scales imbricate, keeled, and strongly mucronate 4 

2. Scales on base of tail not imbricate; not unicolor above 3 

Scales on base of tail imbricate, strongly keeled and mucronate; no dark or 

light markings above, except rarely a few pale blue dots. 

nolascensis (p. 151) 

3. Basal caudals weakly keeled, not or but shortly mucronate; about 110-125 

dorsals from interparietal to back of thighs; 26-30 of largest dorsals equal 
length of head to back of interparietal plate; gular region bluish 

stellata (p. 150) 

Basal caudals keeled and strongly mucronate; about 106-116 dorsals from 

interparietal to backs of thighs; 21-25 of largest dorsals equal length of head 

to back of interparietal plate; gular region blackish palmeri (p. 151) 

4. Dorsal scales from interparietal to rear of thighs 70-81 --squamata (p. 149) 
Dorsal scales 82 or more 5 

5. No stripes whatever in dorsal pattern of either young or adults; dorsals 

92-117, generally more than 100 taylori (p. 150) 

Stripes present in young and some adults; dorsals larger or smaller 6 

6. No distinct dark blue blotch behind axilla raannophorus (p. 149) 

A distinct dark blue spot or blotch behind axilla 7 

7. Two rows of postrostrals; both internasals separated by 2 scales from 

rostral 8 

One row of postrostrals; one or both internasals separated from rostral by 
only one scale 10 

8. Dorsal scales generally 17-23 in a head length (from snout to rear edge of 

interparietal) 9 

Dorsal scales generally 23-28 in a head length.stansburiana hesperis (p. 149) 

9. Size larger (64 mm. snout to vent) martinensis (p. 149) 

Size smaller (48 mm.) concinna (p. 150) 

10. Hind leg longer, 74-85 percent of body length, .stansburiana elegans (p. 148) 
Hind leg shorter, 65-79 percent of body length. 

stansburiana stejnegeri (p. 150) 

UTA STANSBURIANA ELEGANS Yarrow 

Uta elegans Yarrow, Proc. U. S. Nat. Mus., vol. 5, 1882, p. 442. — Schmidt, 
Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 656-657. 

Uta stansburiana elegans, Van Denburgh, Occ. Pap. California Acad. Sci., No. 
10, 1922, pp. 240-247 (part). 



CHECKLIST OF REPTILES OF MEXICO 149 

Type.— U.S. NM. No. 12666. 

Type locality .—hsi Paz, Baja California. 

Range. — Southern two-thirds of Baja California and on Smiths, 
Alejia, Angel de la Guarda, Isla Partida, Isla Raza, Sal Si Puedes, North 
San Lorenzo, South San Lorenzo, Tortuga, San Marcos, Ildefonso, 
East and West Las Galeras, Monserrate, San Jose, San Francisco, 
Espiritu Santo, and Ballena Islands. 

UTA MANNOPHORUS Dickerson 

Uta mannophorus Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, pp. 

470-471.— Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, 

pp. 252-254. 
Uta elegans, Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 656-657 

(part). 

Ty2)e.— U.S.N. M. No. 64260; C. H. Tomisend collector. 
Type locality. — Carmen Island, Gulf of California. 
Range. — Known only from type locality and Danzante and Coro- 
nado Islands, Baja California. 

UTA SQUAMATA Dickerson 

Uta squamata Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 471. — 
Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 249-251. — 
Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 656. 

Type.— U.S. ISl.M. No. 64259; C. H. Townsend collector. 
Type locality. — Santa Catalina Island, Gulf of California. 
Range. — Known only from type locality. 

UTA STANSBURIANA HESPERIS Richardson 

Uta stansburiana hesperis Richardson, Proc. U. S. Nat. Mus., vol. 48, 1915, pp. 

415-418.— Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, 

pp. 233-239.— Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 655.— 

Smith, Handbook of lizards, 1946, pp. 281-283, pi. 72. 
Uta parva Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 471 

(U.S.N.M. No. 64258; San Bartolom6 Bay, Baja California). 

Type.— Univ. California Mus. Vert. Zool. No. 892; J. Grinnell 
collector. 

Type locality. — Arroyo Seco Canon, near Pasadena, Los Angeles 
County, Calif. 

Range. — San Joaquin Valley of central California southward on the 
Pacific slope through northwestern Baja California to San Bartolome 
Bay; Los Coronados Islands. 

UTA MARTINENSIS Van Denbiirgh 

Uta martinensis Van Denburgh, Proc. California Acad. Sci., ser. 3, vol. 4, 
1905, pp. 18-19, pi. 6; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 
247-249.— Schmidt, Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, p. 655. 

Type. — California Acad. Sci. No. 4698; R. H. Beck collector. 



150 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type locality. — San Martin Island, Baja California. 
Range. — Known only from the type locality. 

UTA CONCINNA Dickerson 

Uta concinna Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 470. — 
Schmidt, Bull. Amer, Mus. Nat. Hist., vol. 46, 1922, pp. 655-656. 

Type.—U.S.N.M. No. 64257; C. H. Townsend collector. 
Type locality. — Cerros [= Cedros] Island, Baja California. 
Range. — KnoAvn only from the type locality, and the Natividad 
Islands, Baja California. 

UTA STELLATA Van Denbargh 

Vta stellata Van Denburgh, Proc. California Acad. Sci., Zool., ser. 3, vol. 5, 1905, 
pp. 21-22, pi. 8; Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 225-227. 

Type. — California Acad. Sci. No. 4704, R. H. Beck collector. 
Type locality. — San Benito Islands, Baja California. 
Range. — Known only from the type locality. 

UTA STANSBURIANA STEJNEGERI Schmidt 

Uta stansburiana stejnegeri Schmidt, Amer. Mus. Nov., No. 15, 1921, pp. 1-2; 

Bull. Amer. Mus. Nat. Hist., vol. 46, 1922, pp. 654-655.— Smith, Handbook 

of lizards, 1946, pp. 283-286, pi. 73. 
Uta stansburiana elegans, Van Denburgh, Occ. Pap. California Acad. Sci., No. 

10, 1922, pp. 240-247 (part), pi. 18. 

Type. — Amer. Mus. Nat. Hist. No. 384; A. G. Ruthven collector. 

Type locality. — Mouth of Dry Canon, Alamogordo, Otero County, 
N. Mex. 

Range. — Western Texas through southern Nevada to eastern 
California, southward into northeastern Baja California and the 
northern tier of Mexican states from Sonora to western Coahuila. 
Reported from Baja California: Cocopah Mountains, Volcano Lake, 
etc. ; Sonora: Costa Rica Ranch, Sonoyta, Pozo de Luis, Tepoca Bay, 
San Pedro Bay, Ortiz, Batamotal, Guaymas, Empalme; Islands of 
Tibur6n, Patos, Pelican, and San Esteban; Coahuila: 10 miles east of 
Torredn, San Pedro, 5 miles south of San Pedro, 30 miles west of La 
Rosa, Jaral, Torreon; Durango: Avilco, 5 miles north of Conejos, 
25 miles north of Bermejillo, Lerdo; Chihuahua: 15 miles south of 
Juarez. 

UTA TAYLORI Smith 8' 

Uta taylori Smith, Univ. Kansas Sci. Bull., vol. 22, 1935, pp. 158-166, pi. 26, fig. 3. 

Type.— EHT-RMS No. 10692 (originally field No. 320a), male; E. 
H. Taylor collector. 

Type locality. — Ten miles northwest of Guaymas, Sonora. 

" The possibility of intergradation between Uta taylori and Uta stansburiana, as recently suggested, is 
too remote to consider. Uta stansburiana extends throughout the known range of taylori. 



CHECKLIST OF REPTILES OF MEXICO 151 

Range. — Known only from the region near the type locality. 
Reported from Sonora: La Posa, 54 miles southwest of Hermosillo. 

UTA PALMERI Stejneger 

Uta palmeri Stejneger, North Amer. Fauna, No. 3, 1890, p. 106. — Van Den" 
BTJRGH, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 221-223. 

Type.— U.S. 'NM. No. 16002; Edward Palmer collector. 
Type locality. — San Pedro Martir Island, Sonora. 
Range. — Known only from the type locality. 

UTA NOLASCENSIS Van Denburgh and Slevin 

Uta nolascensis Van Denburgh and Slevin, Proc. California Acad. Sci., ser. 
4, vol. 11, 1921, pp. 395-396. — Van Denburgh, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 223-225. 

Type. — California Acad. Sci. No. 50508; Joseph R. Slevin collector. 
Type locality. — San Pedro Nolasco Island, Sonora. 
Range. — Known only from the type locality. 

Family XANTUSIIDAE Baird 

Xantusiidae Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 254. 

Genera. — Four genera Imown; the only extralimital genus is Cri~ 
cosaura of Cuba. 

Range. — Southwestern United States to Panama; Cuba. 

KEY TO MEXICAN GENERA OF THE XANTUSIIDAE 

1. Supraoculars present, as small scales above eyes (medial to superciliaries) ; 

dorsal scales uniformly granular Xantusia (p. 154) 

Supraoculars absent, no small scales above eyes (except superciliaries) ; dorsal 
scales heterogeneous in size 2 

2. Distinct, vertical rows of well-differentiated, enlarged, keeled scales on sides of 

body, separated from each other by granular area_-_Lepidophyina (p. 151) 
No distinct, vertical rows of enlarged scales on sides of body.Gaigeia (p. 153) 

Genus LEPIDOPHYMA A. Dumeril 

Lepidophyma A. Dumeril, in Dumeril and Dumeril, Catalogue m^thodique de la 

collection des reptiles, 1851, pp. 137-138. 
Poriodogaster Smith, in Gray, Proc, Zool. Soc. London, 1863, p. 154 (type, 

P. grayii Smith). 
Akleistops Muller, Verh. Naturf. Ges. Basel, vol. 6, 1878, p. 390 (type, 

A. guatemalensis MiiUer), 

Genotype. — Lepidophyma flavimaculatus A. Dumeril. 
Range. — From Hidalgo, Alexico, south to Costa Rica. 
Species. — Two, with five forms, one of which {fiavimaculata obscu- 
ra) is extralimital. 



152 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

KEY TO MEXICAN SPECIES OF LEPIDOPHYMA 

1. All (except basal) whorls of enlarged scales on tail separated from each other 

dorsally by 4 rows of scales; femoral pores 15 to 22; median prefrontal nor- 
mally present, sometimes absent flavimaculata flavimaculata (p. 152) 

All whorls of enlarged scales on tail separated from each other dorsally by 3 
rows of scales (rarely feeble evidence of a fourth row); femoral pores less 
than 14; median prefrontal present or absent 2 

2. A median prefrontal smithii smithii (p. 152) 

No median prefrontal 3 

3. Scales in posterior temporal region (anterior to ear) minute and very uniform 

in size, except for a series of relatively large, projecting auricular lobules and 
a row of larger scales beside the upper temporals; whorls on tail relatively 

little differentiated smithii occulor (p. 153) 

Scales in posterior temporal region (anterior to ear) larger, irregular in size; 
auricular lobules poorly defined or absent; no scales bordering upper tem- 
porals; tail whorls strongly differentiated smithii tehuanae (p. 152) 

LEPIDOPHYMA FLAVIMACULATA FLAVIMACULATA A. Dumeril 

Lepidophyma flavimaculatus A. DumiSril, in Dumeril and Dumeril, Catalogue 
m6thodique de la collection des reptiles (Paris Museum), 1851, pp. 138-139. 

Lepidophyma flavomaculaturn flavomaculatum, Smith, Proc. U. S. Nat. AIus., vol. 
92, 1942, p. 379. 

Type. — Mus. Hist. Nat. Paris; P. M. A. Morelet collector. 

Type locality. — Province de Pet6n, Guatemala, here restricted to 
Rio de la Pasion. 

Range. — Atlantic slopes from southern Veracruz to British Hondu- 
ras, excluding the Yucatan Peninsula. Recorded in Mexico from 
Veracruz: Rio de las Playas; Tabasco: Rio de las Playas; Chiapas: 
across the Usumacinta River from Piedras Negras, Peten, Guatemala. 

LEPIDOPHYMA SMITHII SMITHII Bocourt 

Lepidophyma Smithii Bocourt, Journ. Zool., Paris, vol. 5, No. 5, 1876, pp. 

402-403; Mission scientifique au Mexique . . ., Etudes sur les reptiles, livr. 

5, 1878, pp. 309-312, pi. 20F, figs. 3-3b, and pi. 20G, figs. 2-2b. 
Lepidophyma smithii smithii, Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 380. 
Akleistops guatemalensis Muller, Verh. Naturf. Ges. Basel, vol. 6, 1878, pp. 

390-398, pis. 1, 2 (Nat. Mus. Basel; Mazatenango, Guatemala). 

Type. — Mus. Hist. Nat. Paris, six co types; "Commission Scien- 
tifique" collector.^* 

Type locality. — "Tehuantepec, and west coast of Guatemala," re- 
stricted to "Guatemala," here restricted to Mazatenango. 

Range. — Pacific slopes of Chiapas and western Guatemala. Re- 
corded in Mexico only from Chiapas: La Esperanza near Escuintla. 

LEPIDOPHYMA SMITHU TEHUANAE Smith 

Lepidophyma smithii tehuanae Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp» 
377-378. 

" On the basis of the figures given of the types, Smith believes they represent a population characteristic 
of Chiapas and Guatemala, and not of the region of Tehuantepec city. 



CHECKLIST OF REPTILES OF MEXICO 153 

Type.— V.S.N. M. No. 111488; H. M. Smith collector. 

Type locality. — Cerro Arenal, 30 kilometers west of Tehuantepec, 
Oaxaca. 

Range. — Pacific slopes of Oaxaca and Chiapas (?) in the vicinity of 
the Isthmus of Tehuantepec. Reported from Tres Cruces, El Lim6n, 
Cerro Arenal, La Concepci6n, Santa Efigenia, mountains near Santo 
Domingo, near Tehuantepec, Cafetal Concordia; ? Chiapas: Tonala. 

LEPIDOPHYMA SMITHU OCCULOR Smith 

Lepidophyma smithii occulor Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 
378-379. 

TVpe.— U.S.N.M. No. 47133; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — Jalpan, Queretaro. 

Range. — Atlantic slopes presumably from northern Veracruz into 
southern Tamaulipas. Recorded only from San Luis Potosi: Tama- 
zunchale; Queretaro: Jalpan. 

Genus GAIGEIA Smith 

Gaigeia Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, p. 24. 

Genotype. — Lepidophyma gaigeae Mosauer. 
Range. — Hidalgo and Oaxaca. 
Species. — Four. 

KEY TO MEXICAN SPECIES OF GAIGEIA 

1. All whorls on tail complete, none restricted to dorsal surface -gaigeae (p. 154) 
Some of scale whorls on tail restricted to dorsal surface 2 

2. Only one row of scales on ventral surface between whorls of enlarged scales 

(i. e., every third whorl restricted to dorsal surface) 3 

Proximally 2 and distallj- 3 rows of scales on ventral surface between whorls 
of enlarged scales sylvatica (p. 154) 

3. Numerous, very closely approximated, enlarged, keeled scales, separated by 

small granules, present on sides of body; 2 or 3 rows of granules in vertebral 

region radula (p. 153) 

Dorsal scales practically uniform in size; 4 rows of granules in vertebral region. 

dontomasi (p. 153) 
GAIGEIA DONTOMASI Smith 

Gaigeia dontomasi Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 374-376. 

Type.— U.S.N. M. No. 111473; Thomas MacDougall collector. 
Type locality. — Lachiguiri, Oaxaca, 7,100 feet elevation. 
Range. — Known only from the type locality. 

GAIGEIA RADULA Smith 

Gaigeia radula Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 378-379. 
Type.— U. S.N. M. No. 111472; H. M. Smith collector. 



154 BULLETIN 199, UNITED STATES NATIONAL IMUSEUM 

Type locality. — San Jos6 Manteca, 5 kilometers from San Carlos 
Yautepec, Oaxaca. 

Range. — Known only from the type locality. 

GAIGEIA SYLVATICA (Taylor) 

Lepidophyma sylvatica Taylor, Copeia, 1939, pp. 131-133, figs. 1, 2. 
Gaigeia sylvatica, Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 380. 

Type.— ERT-HMS No. 16259; E. H. Taylor collector. 
Type locality. — Seven miles north of Zacualtipan, Hidalgo. 
Range. — Known only from the type locality. 

GAIGEIA GAIGEAE (Mosauer) 

Lepidophyma gaigeae Mosauer, Herpetologica, vol. 1, 1936, pp. 3-5, pi. 2. 
Gaigeia gaigeae. Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, p. 24. 

Type. — Mus. Comp. Zool. No. 42145; Walter Mosauer collector. 

Type locality. — Durango, Hidalgo. 

Range. — The immediate vicinity of the type locality. 

Genus XANTUSIA Baird 

Xantusia Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255. 

Zablepsis Cope, Amer. Nat., vol. 29, 1895, p. 758 (type, XantusiahenshawiStejne- 

ger). 
Amoebopsis Cope, Amer. Nat., vol. 29, 1895, p. 758 (type, Xantusia gilberti Van 

Denburgh). 

Genotype. — Xantusia vigilis Baird. 

Range. — Southern California, Arizona, southern Utah, Baja Cali- 
fornia, and adjacent islands. 

Species. — Five, one of which includes two subspecies; three species 
occur in Mexico. 

KEY TO MEXICAN SPECIES OF XANTUSIA 

1. Ventral plates in 12 series 2 

Ventral plates in 14 series henshawi (p. 155) 

2. A single frontal, eye large vigilis (p. 154) 

A pair of frontals, eye small gilberti (p. 155) 

XANTUSIA VIGILIS Baird 

Xantusia vigilis Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255. — Van 
Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 477-482, pi. 
49.— Smith, Handbook of lizards, 1946, pp. 330-333, pi. 91. 

Type. — U.S.N.M. No. 3063 (three cotypes); John Xantus collector. 
Type locality. — "Fort Tejon," California. 

Range. — Southwestern Utah and eastern California southward into 
northern Baja California as far as San Felipe Bay and San Matias Pass. 



CHECKLIST OF REPTILES OF MEXICO 155 

XANTUSIA HENSHAWI Stejneger 

Xantusia henshawi Stejneger, Proc. U. S. Nat. Mus., vol. 16, 1893, p. 467. — 
Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 484-486, 
pi. 50.— Smith, Handbook of lizards, 1946, pp. 325-327, pi. 88. 

Xantusia pida Cope, Amer. Nat., vol. 29, 1895, pp. 859, 939 (Tejon Pass, Calif.; 
probably Poway, San Diego County, Calif., fide Van Denburgh) . 

Type.—V.S.^M. No. 20339; H. W. Henshaw collector. 

Type locality. — Witch Creek, San Diego County, Calif., elevation 
2,700 feet. 

Range. — Extreme southwestern California and northern Baja Calif- 
fornia, in the San Pedro Martir Mountains. 

XANTUSIA GrLBERTI Van Denburgh 

Xantusia gilherii Van Denburgh, Proc. California Acad. Sci., vol. 5, 1895, p. 121, 
pi. 11; Occ. Pap. California Acad. Sci., No 10, 1922, pp. 482-484. 

Type. — California Acad. Sci. No. 401; Gustav Eisen collector. 
Type locality. — San Francisquito, Sierra Laguna, Baja California. 
Range. — Cape region of southern Baja California. 

Family SCINCIDAE Gray 

Scincidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 201, 

Genera. — About 60, of which three occur in Mexico. Of these three, 
none are confined to the Western Hemisphere. Only two other genera 
occur in the Americas: Cryptoblepharus (South America), a wanderer 
from the Pacific Islands, and Neoseps (Florida) . 

Range. — Southern Canada south through Central America and 
South America to Argentina; West Indies; Africa and Madagascar; 
south Asia; China; Japan; Philippine Islands; Dutch Indies; Pacific 
Islands and Australia. 

KEY TO MEXICAN GENERA OF SCINCIDAE 

1. Two supranasals between rostral and frontonasal; frontoparietal always di- 

vided; lower eyelid with a translucent disk or not; palatine bones in contact 
or not 2 

Supranasals absent, the single frontonasal in contact with rostral; frontoparietal 
divided or not; an undivided translucent disk on lower eyelid; palatine bones 
in contact on median line Scincella (p. 156) 

2. An undivided translucent disk on lower eyelid; palatine bones in contact on 

DQedian line Mabuya (p. 155) 

Lower eyelid without a translucent disk, or if so it is divided into several parts; 
palatine bones separated on median line Eumeces (p. 160) 

Genus MABUYA Fitzinger 

Mabuya Fitzinger, Neue Classification der Reptilien, 1826, pp. 23, 52. 

Genotype. — Lacertus mabouya Lacepede (by absolute tautonymy, 
Jide M. A. Smith, Fauna British India, etc., 1935, Reptilia and Am- 

861316—50 11 



156 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

phibia, vol. 2, p. 257). This is contrary to opinion 92 of the Inter- 
national Commission on Zoological Nomenclature, which states that 
Scincus sloanii of Daudin is the type. Fitzinger did not list that form.^^ 

Range. — Southern Mexico on both coasts; Central America, South 
America; Africa, Madagascar; southern Asia, Philippines, East 
Indies. 

Species. — About 85 species, only one of which occurs in Mexico. 

MABUYA MABOUYA MABOUYA (LacgpSde) 

Lacertus Mahouya LAci:p:feDE, Histoire naturelle des quadrupedes ovipares et des 

serpens, vol. 2, 1788, p. 378, pi. 24 (part). 
Mahuya mahouya mahouya, Dunn, Proc. Acad. Nat. Sci. Philadelphia, vol. 87, 

1935, pp. 537-546 (part). 
Scincus agilis Raddi, Mem. Soc. Ital. Modena, vol. 19, No. 18, 1823, p. 62 (type 

unknown; Rio de Janeiro). 
Mahuia alliacea Cope, Journ. Acad. Nat. Sci. Philadelphia, ser. 2, vol. 8, 1875, 

p. 115. pi. 6, fig. 1 (U. S. N. M. Nos. 30619-20; Costa Rica).88 

Type. — Apparently unknown. 

Type locality. — Restricted by Dunn to the Lesser Antilles; here 
restricted to St. Vincent. 

Range. — Coastal and foothill areas from southern Veracruz and 
Colima southward to Ecuador, Bolivia, and southern Brazil. Re- 
corded from numerous localities in the states of Colima, Michoacan, 
Guerrero, Oaxaca, Veracruz, Tabasco, Campeche, Yucatan, Quintana 
Roo (Cob a), and Chiapas, 

Genus SCINCELLA Mittleman 

Leiolopisma Auct. in part, nee Dumeril and Bibron, Erpetologie gen^rale, vol. 5, 

1839, p. 742 (type, Scincus telfairii Desjardines). 
Scincella " Mittleman, Herpetologica, vol. 6, 1950, p. 19. 

Genotype. — Scincus lateralis Say. 

Range. — Greater part of eastern United States east of the Rocky 
Mountains; Mexico from Hidalgo southward through Central America 
to Panama. Ceylon, southern Asia, China, Philippines, East Indies, 
Tasmania, Africa, New Zealand. 

Species. — About 50 species. Seven species and 10 forms are known 
in the Americas, all except S. incertum of Guatemala occurring in 
Mexico (even this exception may actually occur). 

KEY TO MEXICAN SPECIES OF SCINCELLA 

1. Frontoparietal divided; many median subcaudals in contact on each side with 

2 scales 2 

Frontoparietal single; median subcaudals in contact with only 1 scale on 
each side 6 

" Also see Travassos, Bol. Mus. Nae. Brasil, new ser., Zool., No. 37, 1945, pp. 1-7. 

" This is the only name based upon continental (as opposed to island) specimens north of South America. 
It may be revived, for imdoubtedly geographically segregated variants of taxonomic magnitude do exist 
within the enormous range now included within this supposedly single race. 

" Adoption here of this name is the decision of solely the senior author. 



CHECKLIST OF REPTILES OF MEXICO 157 

2. Usually 3 or more pairs of nuchals; upper tertiary temporal (occasionally 

split) in contact with parietal; 26-28 scale rows around middle of body; 

limbs short, separated by 10-12 scales when adpressed laterals (p. 159) 

Two pairs of nuchals, or fewer; upper tertiary temporal separated from parietal 
by contact of upper secondary temporal and nuchal 3 

3. Limbs, when adpressed, touching or overlapping in adults, forelimb reaching 

rear corner of eye; scale rows at middle of body often 30 or 32 4 

Limbs, when adpressed, separated from each other by 1 or more scale lengths; 
scale rows at middle of body rarely 30 and apparently never 32 5 

4. "Usually 1 pair of nuchals or less; lamellae under fourth toe 19 or fewer; 

dorsolateral light stripe continuous". (Smith) caudaequinae (p. 158) 

Usually 2 pairs of nuchals; lamellae under fourth toe 20 (constant?); dorso- 
lateral light stripe dark-spotted, interrupted silvicolum (p. 158) 

5. Usually nuchals on the two sides 1-2 or more (89 percent) ; lateral light line 

along head and neck poorly defined; axilla-groin/snout-vent percentage 59; 
65 mm. maximum snout-vent measurement. 

gemmingeri gemmingeri (p. 159) 
Usually nuchals on the two sides 1-1 or less (88 percent) ; lateral light line 
along head and neck fairl}' well defined; axilla-groin/snout- vent percentage 
56; 54 mm. maximum snout-vent measurement. 

gemmingeri f orbesorum (p. 159) 

6. Scale rows 30 or more 7 

Scale rows less than 30 9 

7. Tail blue or blue-gray, with traces of a cross-banded pattern evidenced by 

feeble dark bars visible laterally or by transverse series of light spots; legs 

longer, usually overlapping in adults 8 

Tail pink or reddish, with no traces of a cross-banded pattern, but instead with 
a longitudinal dark line on each side at least at base; legs shorter, never over- 
lapping in adults; dorsal scales 65 to 79, usually 69 or more. 

assatum assatum (p. 160) 

8. Dorsal scales 65 to 72, average 69.1 cherriei stuarti (p. 158) 

Dorsal scales 59 to 67, average 63.2 cherriei cherriei (p. 157) 

9. Tail blue or blue-gray, with traces of a cross-banded pattern evidenced by 

feeble dark bars visible laterally or by transverse series of light spots; legs 
longer, usually overlapping in adults; dorsal scales 54 to 60, average 57. 

cherriei izbaac (p. 158) 
Tail pink or reddish, with no traces of a cross-banded pattern, but instead with 
a longitudinal dark line on each side at least at base; legs shorter, never 
overlapping in adults; dorsal scales 58 to 72, average over 66. 

assatum taylori (p. 160) 

SCINCELLA CHERRIEI CHERRIEI (Cope) 

Mocoa cherriei Cope, Proc. Amer. Philos. Soc, vol. 31, 1893, p. 339. 

Lygosoma assatum cherriei, Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 

421, 1940, pp. 13-14. 
Lygosoma cherriei cherriei, Smith, Proc. Biol. Soc. Washington, vol. 54, 1941, pp. 

181-182. 
Leiolopisma cherriei cherriei, Smith, Herpetologica, vol. 3, 1946, p. 111. 
Scincella cherriei cherriei, Mittleman, Herpetologica, vol. 6, 1950, p. 20. 

Type. — Amer. Mus. Nat. Hist. No. 9531; George K. Cherrie 
collector. 

Type locality. — Palmar, Costa Rica. 



158 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Range. — Tabasco and northern Chiapas on Atlantic slopes, and 
Costa Rica on Pacific slopes, eastward to Panama. Recorded in 
Mexico only from Tabasco: Teapa; Chiapas: Palenque. 

SCINCELLA CHERRIEI IXBAAC (Stuart) 

Lygosoma assatum ixbaac Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 421, 

1940, pp. 8-10. 
Leiolopisma cherriei ixbaac, Smith, Herpetologica, vol. 3, 1946, p. 111. 
Scincella cherriei ixbaac, Mittleman, Herpetologica, vol. 6, 1950, p. 20. 

Type.— Vniv. Michigan Mus. Zool. No. 80820; Milton Trautman. 

Type locality. — Chichen Itza, Yucatan. 

Range. — The peninsula of Yucatan, southward as far as Campeche 
and northern Peten, Guatemala. Recorded in Mexico only from 
Yucatan: Chichen Itza; Campeche: Tres Brazos. 

SCINCELLA CHERRIEI STUARTI (Smith) 

Leiolopisma cherriei. Smith, Proc. Biol. Soc. Washington, vol. 52, 1939, pp. 

191-192, 193, pi. 2, fig. 7. 
Lygosoma cherriei stuarti Smith, Proc. Biol. Soc. Washington, vol. 54, 1941, 

pp. 81-82. 
Leiolopisma cherriei stuarti, Smith, Herpetologica, vol. 3, 1946, p. 111. 
Scincella cherriei stuarti, Mittleman, Herpetologica, vol. 6, 1950, p. 20. 

Ti/pe.— U.S.N.M. No. 115174; H. M. Smith collector. 

Type locality. — Potrero Viejo, Veracruz. 

Range. — Central Veracruz, in foothills, southward to the Isthmus 
of Tehuantepec; recorded only from the states of Veracruz and 
Oaxaca (between Rio Grande and La Gloria). 

SCINCELLA CAUDAEQUINAE (Smith) 

Leiolopisma caudaequinae Smith, Univ. Kansas Sci. Bull., vol. 34, 1950, p. — . 

Type.— Vniv. Ilhnois Mus. Nat. Hist. No. 10131; J. P. Craig. 

Type locality. — Salto Cola de Caballo, 25 miles south of Monterrey, 
Nuevo Le6n. 

Range. — Eastern foothills from central Nuevo Le6n southward to 
southern San Luis Potosi (known only from the type locality and 10 
miles west of Naranjo, San Luis Potosi). 

SCINCELLA SILVICOLA (Taylor) 

Leiopisma silvicolum Taylor, Copeia, 1937, No. 1, pp. 5-7. — Smith, Journ. 

Washington Acad. Sci., vol. 39, 1949, p. 41; Univ. Kansas Sci. Bull., vol. 

34, 1950, p. — . 
Scincella silvicola, Mittleman, Herpetologica, vol. 6, 1950, p. 20. 

T^/pe.— EHT-HMS No. 10033, adult female; E. H. Taylor collector. 
Typ>e locality. — Forested hiU about 10 miles southeast of C6rdoba, 
near San Lorenzo, Veracruz. 



CHECKLIST OF REPTILES OF MEXICO 159 

Range. — Atlantic slopes in central Veracruz and in the headwaters 
of the Rio Papaloapam in Oaxaca. Recorded from Veracruz: San 
Jos6 de Gracia; Oaxaca: Cuicatlan. 

SCINCELLA GEMMINGERI GEMMINGERI (Cope) 

Oligosoma gemmingeri Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 180. 

Lygosoma (Mocoa) Gemmingeri, Bocourt, Mission scientifique au Mexique. . . . 
fitudes sur les reptiles, livr. 7, 1881, pp. 449-450. 

Leiolopisma gemmingeri, Taylor, Copeia, 1937, pp. 7-8. — Smith, Journ. Washing- 
ton Acad. Sci., vol. 39, 1949, pp. 40, 41. 

Leiolopisma gemmingeri gemmingeri, Smith, Univ. Kansas Sci. Bull, vol. 34, 
1950, p. — . 

Lygosoma laterale (part), Boulenger, Catalogue of the lizards in the British 
Museum, vol. 3, 1887, p. 263. 

Type. — U.S.N.M. No. 6331, four cotypes; Frangois Sumichrast 
collector. 

Type locality. — Orizaba, Veracruz. 

Range. — Eastern Hidalgo, central and southern Veracruz, and 
Oaxaca south to Tehuantepec, on the slopes of the plateau and in 
lowlands. Recorded only from Veracruz: Tequeyutepec, La Perla, 
Rio Verde, Jalapa, Orizaba; Hidalgo; Zacualtipan (possibly referable 
to L. g. forbesorum) ; Oaxaca: Ixcuintepec, Tres Cruces, San Jos6 
Manteca, 17 miles north of Niltepec, Cafetal Concordia. 

SCINCELLA GEMMINGERI FORBESORUM (Taylor) 

Leiolopisma forbesorum Taylor, Copeia, 1937, pp. 8-11. 

Leiolopisma gemmingeri forbesorum, Smith, Univ. Kansas Sci. Bull., vol. 34, 
1950, p. — . 

Type.— EUT-HMS No. 10043, E. H. Taylor collector. 

Type locality. — La Placita, Hidalgo, 8 miles south of Jacala, elevation 
7,000 feet. 

Range. — Known only from higher mountains of Hidalgo, in the 
vicinity of the type locality. 

SCINCELLA LATERALE (Say) 

Scincus lateralis Say, in Long's Expedition to the Rocky Mountains, vol. 2, 1823, 
p. 234. 

Leiolopisma laterale, Jordan, Manual of the vertebrates of the northern United 
States, ed. 8, 1899, p. 324.— Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 
pp. 622-624, fig. 123.— Smith, Handbook of lizards, 1946, pp. 337-340, pi. 92. 

Scincella laterale, Mittleman, Herpetologica, vol. 6, 1950, p. 19. 

Type.— U.S.N.M. No. 3152. 

Type locality. — Mississippi River, below Cape Girardeau, Mo. 

Range. — New Jersey and southward to the Gulf and to central 
Coahuila, excluding the lower Rio Grande Valley. Reported from 
Coahuila: Nogales (Sabinas River near Miizquiz). 



160 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

SCINCELLA ASSATA ASSATA (Cope) 

LamprophoUs assatus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, pp. 179-180. 
Leiolopisma assatum, Burt, Trans. American Micr. Soc, vol. 54, 1935, p. 177. 
Leiolopisma assatum [assatum], Oliver, Occ. Pap. Mus, Zool. Univ. Michigan, 

No. 360, 1937, p. 12.— Smith, Herpetologica, vol. 3, 1946, p. Ill; Journ. 

Washington Acad. Sci., vol. 39, 1949, p. 40. 
Lygosoma assatum assatum, Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, 

No. 421, 1940, pp. 12-13. 
Scincella assata assata, Mittleman, Herpetologica, vol. 6, 1950, p. 20. 

Type.— Acad. Nat. Sci. Philadelphia No. 9465. 
Type locality. — Volcan Isalco, El Salvador. 

Range. — Pacific slopes, southeastern Chiapas southward to Hon- 
duras. Kecorded in Mexico from Chiapas: Huehuetan, Escuintla. 

SCINCELLA ASSATA TAYLORl (Oliver) 

Leiolopisma assatum taylori Oliver, Occ. Pap. Mus. Zool. Univ. Michigan, No. 

360, 1937, pp. 12-15. — Smith, Journ. Washington Acad. Sci., vol. 39, 1949, 

p. 40. 
Scincella assata taylori, Mittleman, Herpetologica, vol. 6, 1950, p. 20. 

Type. — Mus. Zool. Univ. Michigan No. 80107; A. Bakewell and 
J. A. Oliver collectors. 

Type locality. — Santiago, Colima. 

Range. — Colima to southwestern Chiapas, in lower areas. Recorded 
from Colima: Santiago, Paso del Rio; Guerrero: Between Rinc6n and 
Cajones, Mazatlan, Chilpancingo, Agua del Obispo, Tierra Colorada; 
Jalisco: Tenacatita; Chiapas: Tonala; Oaxaca: Matias Romero, Santa 
Efigenia, Tapanatepec. 

Genus EUMECES Wiegmann 

Eumeces Wiegmann, Herpetologia Mexicana, 1834, pt. 1, p. 36. 

Lamprosaurus Hallo well, Proc. Acad. Nat. Sci. Philadelphia, 1852, p. 206 

(type, Lamprosaurus guttulatus lia\\oweU. = Plestiodon obsoletum Baird and 

Girard). 
PlatyphoUs Duofes, La Naturaleza, ser. 2, vol. 1, 1887, p. 486 (type, Eumeces 

altamirani Dug5s). 

Genotype. — Scincus pavimentatus GeoSroy = Eumeces pavimentatus 
(see Wiegmann, Archiv fiir Naturg., vol. 2, 1835, p. 288). 

Range. — Southern Canada southward through the United States, 
Mexico, and Central America to Nicaragua; Bermuda; North Africa; 
southwestern Asia; China; French Indo-China; Japan. 

Species. — About 58 species and 76 forms are recognized; 23 forms 
of 22 different species occur in Mexico. 

KEY TO MEXICAN SPECIES OF EUMECES 

1. Median dorsal scales at least three times the width of adjoining scales 2 

Dorsal scales not greatly widened, rarely twice as wide as adjoining scales 3 

2. Scales in 21 rows about middle of body; 3 broad black stripes begin on snout, 

pass back and break up on middle of back schwartzei (p. 162) 



CHECKLIST OF REPTILES OF MEXICO 161 

Scales in 19 rows about middle of body; no stripes, but scales with black 
dots altamirani (p. 163) 

3. Three supraoculars; no postnasal; 1 postmental 4 

Four supraoculars ; postnasals and postmentals variable 5 

4. Parietals enclose interparietal; scale rows about middle of body, 24; seventh 

labial in contact with upper secondary temporal dugesii (p. 169) 

Parietals not enclosing interparietal; scale rows usually 24; seventh labial 
not in contact with upper secondary temporaL.lynxe furcirostris (p. 163) 

5. Postmentals normally 2 6 

Postmental normally 1 12 

6. Body without white stripes at any time; lateral scale rows generally oblique; 

young black with a blue tail; head with cream spots; adults oUve with 
darker areas on scales; limbs large, overlapping when adpressed; scales 

26-28 rows; 125 mm. snout to vent obsoletus (p. 165) 

Body with white stripes in young, retained in adult, or'approaching uniform 
ohve coloration in adults, especially old males; lateral scale rows parallel; 
young never black but usually with blue or pink (red) or orange tail 7 

7. Dorsolateral light stripes distinct and confined to third scale row. 

multivirgatus (p. 165) 

Dorsolateral light stripes not involving third scale row at all, or else including 

also second scale row 8 

8. Median light Une beginning between arms running forward bifurcating on 

head; or median hne lost and the bifurcating Unes only, remain on head. 9 

No trace of a median light line or bifurcating lines on head; usually 4 light 

lines, 2 lateral and 2 dorsolateral 10 

9. Parietals enclose interparietal; scale rows usually 28 at middle of body; post- 

nasal usually present; median white Une to shoulder; limbs short, not 

touching when adpressed callicephalus (p. 164) 

Parietals not enclosing interparietal; bifurcating lines on head, but median 
line obsolete; postnasal absent; limbs touch in young when adpressed, in 
adults separated by 3 or 4 scale lengths; postmental single or divided; 26-28 
scale rows; maximum size about 70 mm tetragrammus (p. 165) 

10. Parietals enclose interparietal; 24 scale rows; seventh labial broadly in contact 

with upper secondary temporal; tail orange in young; 4 light lines retained 

in adults; limbs fail to touch when adpressed lagtmensis (p. 167) 

Parietals do not enclose interparietal 11 

11. Snout-vent length greater than 75 mm., or interparietal nearly parallel- 

sided; specimens under 50 mm. snout-vent length with pink tail (no blue); 
8 supralabials on each side (98 percent) ...gilbertirubricaudatus (p. 167) 
Snout-vent length not more than 75 mm.; interparietal bluntly wedge-shaped; 
juveniles, usually adults, with blue tail; 7 supralabials on one or both sides 
(94 percent) skiltonianus (p. 167) 

12. A median light line bifurcating on frontal 13 

Median light line present or absent, if present bifurcating behind frontal. 14 

13. Median line from middle of body bordered with darker, and bifurcating on 

anterior half of frontal; limbs separated when adpressed; usually 24 scales 

about body lynxe lynxe (p. 163) 

Median light line bifurcating on the posterior part of frontal, or all lines lost 
in adult olive coloration; limbs overlapping when adpressed; subcaudals 
widened; maximum snout to vent length, 100 mm._.suinichrasti (p. 164) 

14. Parietals enclosing interparietal 15 

Parietals not enclosing interparietal 17 



162 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

15. Limbs long, overlapping when adpressed; strong, wide dorsolateral light lines; 

seventh labial broadly in contact with secondary upper labial; no primary 
temporal; frontoparietals narrowly separated; scale rows, 28; snout to 

vent, 65 mm colimensis (p. 169) 

Limbs shorter, separated when adpressed 16 

16. Seventh labial not touching upper secondary temporal; primary temporal as 

large as upper secondary temporal; 24 scale rows; maximum length snout to 

vent, 51 mm parvulus (p. 166) 

Seventh labial broadly in contact with upper secondary temporal; primary 
temporal present, smaller than upper secondary temporal; 22-24 scale rows 
around middle of body; maximum length snout to vent, 66 mm. 

indubitus (p. 168) 

17. Primary temporal absent; limbs short, not touching when adpressed; seventh 

labial in contact with upper secondary temporal; 22 scale rows; dorsolateral 

lines growing dim posteriorly dicei (p. 169) 

Primary temporal present 18 

18. Four narrow dark dorsal lines and narrow dorsolateral light lines; 22-24 

scale rows; no postnasal; limbs widely separated when adpressed; maximum 

snout to vent length, 76 mm copei (p. 166) 

Dorsal pattern not of 4 narrow black lines 19 

19. Bifurcating lines on head; lateral and dorsolateral light lines rarely reaching 

farther back than arm; scale rows, 26-28; maximum snout to vent length, 

66 mm brevilineatus (p. 164) 

No bifurcating lines on head 20 

20. A postnasal; usually bronze olive, with lateral brown stripe, with sometimes 

trace of a dorsolateral light line; 26 scale rows; the limbs adpressed, 

the toes may or may not touch humilis (p. 166) 

No postnasal 21 

21. Scale rows 20 around body; ear small, median dorsal scales widened. 

parviauriculatus (p. 166) 
Scale rows more than 20 22 

22. Scale rows 22 (rarely 24); body slender, dorsolateral lines distinct to tail; 

seventh labial usually touching upper secondary temporal; snout to vent 

56 mm ochoterenai (p. 169) 

Scale rows usually 24, rarely 22; dorsolateral lines to tail or not; parietal 
usually not enclosed but occasionally may be enclosed, variable. 

brevirostris (p. 168) 

EUMECES SCHWARTZEI Fischer 

Eumeces schwartzei Fischer, Abh. Nat. Ver. Hamburg, vol. 8, 1884, pp. 3-5, 
pi. 7, fig. 1.— Taylor, Uniy. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), 
pp. 94-101, figs. 5, 6, pi. 1. 

Type. — Naturh. Mus., Hamburg, Germany .^^ 

Type locality. — "Einer kleinen Insel in der Laguna de Terminos 
(Campeche Bai)." 

Range. — Campeche, Tabasco, Yucatan; in Central America it is 
known from Guatemala and British Honduras. Keported from 
Campeche: Tres Brazos, Encarnaci6n, Isla de Carmen; Tabasco: 
Tenosique; Yucatan: Chichen Itza. 

" This specimen was actually taken aboard ship in a cargo of dyewood and collected on board ship, alive, 
collector unknown. It was sent to the Zoologischen Garten In Hamburg. At its death it was sent to the 
Naturhistorischen Museum of Hamburg. 



CHECKLIST OF REPTILES OF MEXICO 163 

EUMECES ALTAMIRANl Dngfes 

Eumeces altamirani Ducfes, La Naturaleza, ser. 2, vol. 1, 1887-1890 (1891), 
pp. 485, 486, pi. 22, 6 figs.— Taylor, Proc. Biol. Soc. Washington, vol. 49, 
1936, pp. 55-58; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 102-103, 
fig. 6, pi. 2. — Smith and Neckeu, Anal. Esc. Nac. Cienc. Biol., vol. 3, 1943, 
pp. 190-192. 

Type. — In Museo "Alfredo Duges" in Colegio del Estado de 
Guanajuato in Guanajuato (without number) ; Federico Altamirano 
collector. 

Type locality. — "Regiones Calidas del Estado de Michoacan" 
Gater designated by Duges as Apatzingan de la Constituci6n, Mich- 
oacan) . 

Range. — Known only from Michoac§,n, on the southern slopes of 
the plateau. Reported from El Sabino, Apatzingan. 

EUMECES LYNXE LYNXE (Wiegmann) 

Scincus quinquelineatus var., Wiegmann, Isis von Oken, 1828, p. 373 {non 

Linnaeus). 
Plestiodon quinquelineatum, Dumeril and Bibron, ErpcStologie g^n^rale, vol. 5, 

1839, pp. 707-708 (part). 
Euprepes lynxe Wiegmann, Herpetologia Mexicana, 1834, pp. 36-37. 
Eumeces lynxe, Peters, Monatsb. Akad. Wiss. Berlin, 1864, p. 484. — Smith, 

Journ. Washington Acad. Sci., vol. 39, 1949, p. 41. 
Eumeces lynxe lynxe, Taylor, Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), 

pp. 163-173, pi. 41, figs. 18, 19, fig. B. 
Plestiodon Bellii Gray, Catalogue of the specimens of lizards in the collection of 

the British Museum, 1845, p. 92 (type locality not designated, here restricted 

to El Chico, Hidalgo). 

Type.— Zool. Mus., Berlin; F. Deppe collector. 

Type locality. — "Specimena nostra prope Chico invenit Deppe." 
\erj probably (and here restricted to) El Chico (or Mineral El Chico) 
near Pachuca, Hidalgo, where the species has been found to be abun- 
dant. 

Range. — Known definitely from Hidalgo, Veracruz, San Luis 
Potosi and Puebla; recorded, probably erroneously, from Guerrero, 
Guanajuato and Michoacan. Reported from Hidalgo: Zacualtipan, 
San Miguel, Guerrero, Durango, El Chico; Vtracruz: Jalapa, Mount 
Orizahsi; San Luis Potosi: Alvavez; Puebla: Zacatlan. 

EUMECES LYNXE FURCIROSTRIS Cope 

Eumeces furcirostris Cope, Proc. American Philos. Soc, vol. 22, 1885, pp. 169- 
170.— Taylor, Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), pp. 173- 
178, figs. 18, 20; Univ. Kansas Sci. Bull., vol. 29, 1943, pp. 272-273. 

Type.— Acad. Nat. Sci. Philadelphia No. 11327; Dr. Flohr collector. 

Type locality.— J aXsiipSi, Veracruz. 

Range. — Puebla, central northern Veracruz, and possibly eastern 
Hidalgo. Reported from Puebla: Tezuitlan; Veracruz: Toxtlacuaya, 
Jalapa, Las Vigas, Cofre de Perote; Hidalgo: Zacualtipan. 



164 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

EUMECES SUMICHRASTI (Cope) 

Plistodon sumichrasli Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 321. 

Eumeces sumichrasli, Bocourt, Mission scientifique au Mexique . . ., fitudes sur 
les reptiles, Hvr. 6, 1879, p. 422. — Taylor, Univ. Kansas Sci. Bull., vol. 23, 
1935 (1936), pp. 178-186, figs. 21-23, pi. 12.— Smith and Necker, Anal. Esc. 
Nac. Cienc. Biol., vol. 3, 1943, pp. 192-194, pi. 3, fig. 4.— Smith, Journ. 
Washington Acad. Sci., vol. 39, 1949, p. 41. 

Eumeces rovirosae Duces, La Naturaleza, ser. 2, vol. 2, 1895-1896 (1895), pp. 
298-299, pi. 13 (type locality, Mineral de Santa Fe, Chiapas; type in Alfredo 
Duges Museum, Colegio del Estado de Guanajuato). 

Eumeces schmidti Dunn, Proc. Acad. Nat. Sci. Philadelphia, vol. 84, Mar. 22, 
1932, pp. 30-31 (type locality, Lancetilla, Honduras; Acad. Nat. Sci. Phila- 
delphia No. 19877). 

T?/pe.— U.S.N. M. No. 6601; Fran9ois Sumichrast collector. 

Type locality. — "Orizava" ex errore, = Potrero, Veracruz. The 
specimen bears Sumichrast's original tag, "Potrero, No. 4, F. Sumi- 
chrast." Later Sumichrast published notes that he had found the 
species "en los encinales de Potrero, cerca de Cordoba a una altura 
de 590 metros." 

Range. — Veracruz to Chiapas in lowlands; Central America; Hon- 
duras, British Honduras, and Guatemala. Reported from Veracruz: 
Potrero, Jalapa; Chiapas: Palenque, La Esperanza, Mineral de Santa 
F6. 

EUMECES BREVILINEATUS Cope 

Eumeces brevilineatus Cope, U. S. Nat. Mus. Bull. 17, 1380, pp. 18-19, 44, 46. — 
Taylor, Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 283-290, figs. 
41, 42, 43, pi. 22. 

Type. — Lectotype, designated by Taylor, loc. cit., Q.S.N.M. No. 
10159, one of four cotypes; G. W. Marnock collector. 

Type locality. — Helotes, Bexar County, Tex. 

Range. — Tamaulipas and Nuevo Leon; in the United States: 
southern Texas. Reported from Tamaulipas: 26 kilometers north of 
El Lim6n; Nuevo Leon: 4 miles west of Sabinas Hidalgo, 31 miles 
south of Sabinas Hidalgo. 

EUMECES CALUCEPHALUS Bocourt 

Eumeces callicephalus Bocourt, Mission scientifique au Me.xique . . ., fitudes sur 
les reptiles, livr. 6, 1879, pp. 431-433, pi. 22D, figs. 2-2c, pi. 22E, fig. 2.— 
Taylor, Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 290-298, figs. 44, 
45, pi. 23. 

Type.— Mus. Nat. Hist. Nat. Paris; Alfredo Duges collector. 

Type locality. — Guanajuato [Guanajuato]. 

Range. — Sonora (Alamos), Chihuahua, Durango, Zacatecas, Jalisco, 
Guanajuato, Michoacan, Queretaro (Huaxteca Potosina), Nayarit; in 
United States: Arizona (southeastern). 



CHECKLIST OF REPTILES OF MEXICO 165 

EUMECES TETRAGRAMMUS (Baird) 

Plestiodon tetragrammus Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 256. 
Eumeces tetragrammus, Cope, U. S. Nat. Mus. Bull. 1, 1875, p. 45 — Tayloe, 

Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), pp. 298-304, fig. 46; 

ihid., vol. 29, 1943, p. 274. 
[Eumeces tetragrammus] var. funebrosus Cope, Ann. Rep. U. S. Nat. Mus., 1898. 

(1900), p. 661 (type locality, Matamoros, Tamaulipas; U. S. N. M. No. 3120, 

two cotypes). 

Type.— JJ. S. N. M. No. 3124; Darius Nash Couch collector. 

Type locality. — "Lower Rio Grande." (Later indicated by Baird, 
as "Matamoras, Mex." A second specimen was at hand collected by 
Dr. Kennerly at "Salado River," U. S. N. M. No. 3139.) 

Range. — Tamaulipas, San Luis Potosi, and Veracruz, in lowlands; 
in the United States: southern Texas. Reported from Tamaulipas: 
Sji miles west of El Forl6n, San Jose, Matamoros; San Luis Potosi: 
Antiguo Morelos, Ebano; Veracruz: south of Tampico. 

EUMECES OBSOLETUS (Baird and Girard) 

Plestiodon obsoletum Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, vol. 

6, 1852, p. 129. 
Eumeces obsoletus, Cope, U. S. Nat. Mus. Bull. 1, 1875, p. 45. — Taylor, Univ. 

Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), pp. 305-320, figs. 47, 48, pi. 24, 
Lamprosaurus guttulatus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1852, 

pp. 206-207 (type locality, "Jornada del Muerte," New Mexico; Acad. Nat. 

Sci. Philadelphia). 

Type.—V. S. N. M., No. 3133; John H. Clark collector. 

Type locality. — "Valley of the Rio San Pedro of the Rio Grande del 
Norte," now Devils River, Tex. 

Range. — Tamaulipas, Nuevo Le6n, Coahuila, and Chihuahua. In 
United States: Arizona, New Mexico, and Texas, north to Utah, 
Colorado, Nebraska, and Kansas. Reported from Tamaulipas: 
Matamoros; Nuevo Leon: Santa Catarina; Chihuahua: Chilmahua 
(city); Coahuila: Cuatro Cienegas. 

EUMECES MULTIVIRGATUS (Hallowell) 

Plestiodon multivirgatum Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1857, 

p. 215. 
Eumeces multivirgatus, Cope, U. S. Nat. Mus. Bull. 1, 1875, p. 45. — Taylor, Univ. 

Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 341-353, figs. 54-56, pis. 27, 28. 

Type. — Acad. Nat. Sci. Philadelphia No. 9371; Dr. Hammond 
collector. 

Type locality. — "Posa Creek, 460 miles west of Fort Riley, Kan- 
sas" (=Cow Creek,^^* Larimer County, Colo.). 

Range. — The central high plains and proplateau areas south from 
southwestern Nebraska to extreme western Texas, northern Arizona, 

««• The original tag attached to the specimen definitely states Cow Creek. The "Posa" is a bad interpreta- 
tion of poor chirography. 



166 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

and presumably western Chihuahua. Recorded in Mexico only from 
Chihuahua: no specific locality .^^ 

EUMECES HUMILIS Boulenger 

Eumeces Bocourtii (non Brocchi) Boulenger, Ann. Mag. Nat. Hist., ser. 5, vol. 11, 
1883, p. 342. 

Eumeces humilis Boulenger, Catalogue of the lizards in the British Museum, 
vol. 3, 1887, p. 377 (new name for Eumeces Bocourtii). — Taylor (part), 
Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 358-363, figs. 58, 59.— 
Smith, Proc. New England Zool. Club, vol. 21, 1942, pp. 94, 95. 

Type.—Bvit. Mus. Nat. Hist. Nos. 83,4,5,33-34 (two cotypes); 
Alfonso Forrer collector. 

Type locality. — Presidio [Sinaloa]. There is a strong presumption 
that the locality is in Sinaloa although not known positively to be so. 

Range. — Sinaloa. Known only from the type locality. (Specimens 
previously referred to this species from Texas and New Mexico have 
been referred to another form, Eumeces taylori, by Smith.) 

EUMECES PARVULUS Taylor 

Eumeces parvulus Taylor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp. 175- 
178, fig. 1; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 363-367, figs. 
59, 60, pi. 31, figs. 3, 4.— Smith, Copeia, 1943, p. 250. 

r?/^?^.— U.S.N.M. No. 56903; collector unknown. Originally in the 
Julius Hurter collection. 

Type locality. — Tepic, Nayarit. 

Range. — Colima, Nayarit and Sinaloa. Reported from Colima: 
Paso del Eio; Nayarit: Tepic, Compostela, Miniman; Sinaloa: Plomo- 
sas. 

EUMECES PARVIAURICULATUS Taylor 

Eumeces parviauriculatus Taylor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp* 
178-181. fig. 2; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 368-371, 
fig. 59, pi. 31, fig. 5. 

TVpe.— U.S.N.M. No .^47536; E. A. Goldman collector. 
Type locality. — Near Alamos, Sonora. 

Range. — Sonora and western Chihuahua. Reported from Sonora: 
Near Alamos; Chihuahua: Mojarachic. 

EUMECES COPEI Taylor 

Eumeces copei Taylor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp. 133-137; 
Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 387-394, figs. 64, 65, pi. 33. 

Tj/pe.—ERT-UMS No. 29717 (EHT field No. 3859); H. M. Smith 
and E. H. Taylor collectors. 

Type locality. — Ten miles southeast of Asuncion, in the western part 
of the state of Mexico. 



M This specimen, U. S. N. M. No. 30833, very likely represents a distinct species (see Taylor, loc. eft., 
p. 353) , but we hesitate to attempt a characterization of it because of certain obvious abnormalities the speci- 
men possesses, and because of its intimate and entirely uncertain relationship with populations farther to 
the north In Arizona and New Mexico. 



CHECKLIST OF REPTILES OF MEXICO 167 

Range. — Puebia, Distrito Federal, Morelos, Mexico, and Michoacan. 
Reported from Puebia: Near Rio Frio (Mexico); Mexico: Rio Frio, 
near Salazar, 10 miles southeast of Asuncidn, 8 miles west of Villa 
Victoria, 15 kilometers west of Toliica, 3 miles west of Zinacatepec; 
Morelos: Tres Marias, Lagunas de Zempoala; Distnto Federal: Santa 
Lucia; Michoacdn: Cerro de Tecolote, Mount Tancftaro. 

EUMECES SKILTONIANUS (Baird and Girard) 

Plestiodon skiltonianum Baird and Girakd, Proc. Acad. Nat. Sci. Philadelphia, 

1852, p. 69. 
Eumeces skiltonianus, Cope, U. S. Nat. Mus. Bull. 1, 1875, p. 45. — Rodgers and 

Fitch, Univ. California Publ. Zool., vol. 48, 1947, pp. 193-198, pi. 8a (col.). 
Eumeces skiltonianus skiltonianus, Taylor, Univ. Kansas Sci. Bull., vol. 23, pt. 

1. 1935 (1936), pp. 415-428, fig. 68, pis. 35, 36, figs. 2, 3, 4. 
Eumeces skiltonianus amhlygrammus Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 

p. 643 (type locality. Fort Humboldt, Calif.; U.S.N.M. No. 3166). 

Type. — U.S.N.M. No. 3172, two cotypes, smaller one designated as 
lectotype; Rev. George Gary (or Geary) collector. 

Type locality. — Oregon, here restricted to The Dalles. 

Range. — Southern British Colmnbia to northwestern Baja California 
and adjacent islands, and eastward to western Montana and Utah. 
Reported in Baja California from Los Coronados Islands, Todos Santos 
Islands; Ensenada, San Jose, Rancho San Jos^, Alcatraz, San Pedro 
Martir Mountains, Arroyo Encantada, San Quintin, etc. 

EUMECES GILBERTI RUBRICAUDATUS Taylor 

Eumeces quadrilineatus Hallowell (nee BIyth\ Reports of explorations and 
surveys, to ascertain the most practicable and economical route for a railroad 
from the Mississippi River to the Pacific Ocean, 1853-6, vol. 10, pt. 4, Zool., 
Rept., 1859, p. 10, pi. 9, figs. 3a, b, c, d (type locality, "Upper California, 
near Mohave river and in San Bernardino Valley," here restricted to Mojave 
River; U. S. Nat. Mus.). 

Eumeces gilberti rubricaudatus Taylor, Univ. Kansas Sci. Bull., vol. 23, 1935 
(1936), pp. 446-451, figs. 72-73, pi. 39, vol. 29, 1943, p. 277; Rodgers and 
Fitch, Univ. California Publ. Zool., vol. 48, 1947, pp. 203-205, pi. 8e (col.). 

Type.— Caliiornia. Acad. Sci. No. 39002. 

Ty2)e locality. — Tehachapi Mountains, Calif. 

Range. — Southern California and extreme northwestern Baja 
California. Reported in Mexico only from Baja Calijornia: San 
Antonio del Mar, North Coronado Island. 

EUMECES LAGUNENSIS Van Denburgh 

Eumeces lagunensis Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 5, 
1895, pp. 79, 13^^-135, pi. 13.— Taylor, Univ. Kansas Sci. Bull., vol. 23, 
1935 (1936), pp. 431-437, figs. 69, 70, pi. 36, fig. 1.— Rodgers and Fitch, 
Univ. California Publ. Zool., vol. 48, 1947, pp. 205-206. 



168 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Plesiiodon lagunensis, Van Denburgh and Slevin, Proc. California Acad. Sci., 

ser. 4, vol. 11, 1921, pp. 28, 40, 44, 52. 
Plesiiodon skiltonianus lagunensis, Nelson, Mem. Nat. Acad. Sci., vol. 16, 1921, 

pp. 114-115. 
Eumeces skiltoniamis lagunensis, Linsdale, Univ. California Publ. Zool., vol. 38, 

1932, p. 374. 
Eumeces skiltonianus, Loveridge, Copeia, No. 173, 1930, pp. 111-112. 

Type. — Originally California Acad. Sci. Nos. 400 and 402, cotypes 
(destroyed in the earthquake and fire in 1906); Gustav Eisen collector. 
Neotype.— U. S. N. M. No. 67398; W. M. Mann collector; February 
1924, "on the Trail between Loreto and Comondu." 

Type locality. — San Francisquito, Sierra de la Laguna, Baja 
California. 

Range. — Confined to the southern third of Baja California in the 
mountains. Reported from Baja California: Comondu; San Francis- 
quito, Sierra de la Laguna; between Loreto and Comondu. 

EUMECES BREVIROSTRIS (Gunlher) 

Mabouia brevirostris Gunther, Proc. Zool. Soc. London, 1860, pp. 316-317. 

Eumeces brevirostris, Bocotjrt, Mission scientifique au Mexique . . ., fitudes sur 
les reptiles, livr. 6, 1879, pp. 439-440, pi. 22A, figs. 7, 7a, 7b, and pi. 22E, 
fig. la.— Taylor, Univ. Kansas Sci. Bull., vol. 23, pt. 1, 1935 (1936), pp. 
459-466, figs. 76, 77, pi. 41. 

Type. — Brit. Mus. Nat. Hist.; Auguste Salle collector. 

Type locality. — Oaxaca, here restricted to the city of Oaxaca. 

Range. — Durango to Oaxaca and east to Veracruz, in highlands. 
Reported from Guerrero: Omilteme; Oaxaca: Cerro San Felipe, La 
Parada, San Jose Lachiguiri, Tehuantepec; Veracruz: Laguna, Perote, 
San Bernardino, Totalco, Orizaba; Puebla: Near Rio Frio (Mexico), 
TexTuelucan, El Seco (Km. 205); Durango: Ciudad, El Sal to, Coyote; 
Jalisco: La Cumbre de los Arrastrados, Talpa, Mascota, Sierra de 
Juanocatlan, La Laguna; Michoacdn: No specific record. 

EUMECES INDUBITUS Taylor 

Eumeces indubitus Taylor, Univ. Kansas Sci. Bull., vol. 21, 1933 (1934), pp 
257-267, pis. 24, 25; vol. 23, pt. 1 (1936), pp. 466-472, figs. 76, 78, pi. 42. 

T^^e.— EHT-HMS No. 29715 (EHT field No. 1731); E. H. Taylor 
and H. M. Smith collectors. 

Type locality. — Kilometer 63 (Mexico-Cuernavaca highway), near 
Cuernavaca, Morelos. 

Range. — Known from the states of Morelos, Mexico, and Michoacan. 
Reported from Morelos: near Cuernavaca, Tepoztlan, Lagunas de 
Zempoala, Huajintlan; Mexico: Asunci6n; Michoacdn: 15 miles south- 
east of Zitacuaro, Puerto Hondo. 



CHECKLIST OF REPTILES OF MEXICO 169 

EUMECES DUGESII Thominot 

Eumeces (Plestiodon) Dugesii Thominot, Bull. See. Philom. Paris, ser. 7, vol. 7 
(1882-1883), 1883, pp. 138-139.— Taylor, Univ. Kansas Sci. Bull., vol. 23, 
pt. 1, 1935 (1936), pp. 472-478, figs. 76, 79, pi. 43. 

Tyjje. — Mus. Hist. Nat. Paris; Alfredo Duges collector. 

Type locality. — ^''Province Guanajuato," here restricted to the city 
of Guanajuato. 

Range. — Guanajuato, Jahsco (?), and Michoacan. Reported from 
Guanajuato: Guanajuato; Michoacdn: Carapa, Rancho San Jose, 9 
miles west of Zacapii, Tangancicuaro, Patamban, El Soledad (Tan- 
cltaro), Apatzingan; Jalisco: Nevado de Colima.®" 

EUMECES COUMENSIS Taylor 

Eumeces colimensis Taylor, Publ. Field Mus. Nat. Hist., zool. ser., vol. 20, 1935, 
pp. 77-80, fig. 7; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 478-482, 
figs. 76-80, pi. 40, fig. 3. 

Type. — Chicago Nat. Hist. Mus. No. 1649; collector unknown. 

Type locality. — Colima, Colima. 

Range. — Known only from the type locaUty. 

EUMECES DICEI Ruthven and Gaige 

Eumeces dicei Ruthven and Gaige, Occ. Pap. Mus. Zool. Univ. Michigan, No. 
260, 1933, pp. 1-3.— Taylor, Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), 
pp. 482-485, figs. 76-81; vol. 29, 1943, pp. 280-282. 

Type.— Mus. Zool. Univ. Michigan No. 69253; Lee R. Dice 
collector. 

Type locality. — Marmolejo, Tamaulipas. 

Range. — Tamauhpas and Nuevo Leon. Reported from N'uevo 
Leon: Pablillo, Cieneguillas south of Galeana; Tamaulipas: Mar- 
molejo. 

EUMECES OCHOTERENAI Taylor 

Eumeces ochoterenae Taylor, Proc. Biol. Soc. Washington, vol. 46, 1933, pp. 
129-133, 2 figs.; Univ. Kansas Sci. Bull., vol. 23, 1935 (1936), pp. 485-489, 
figs. 76, 82, pi. 43. 

Type.— ERT-B.MS No. 29716 (EHT Field No. 1015); E. H. 
Taylor and H. M. Smith collectors. 

Type locality. — Mazatlan; "4 mi. north of Chilpancingo, Guerrero" 
(actually about 15 km. south of Chilpancingo). 

Range. — Known only from Guerrero; probably confined to the 
Sierra Madre del Sur. Reported from Agua del Obispo, 7 miles east 
of Chilpancingo, Mazatlan, vicinity of Chilpancingo, between Rincon 
and Cajones, Chilapa. 

M Gadow (Proc. Zool. Soc. London, 1905, pp. 195, 218-219) records a "fuscirostris" (=furciTOstTis) from 
this locality. Inasmuch as the most distinctive feature of d. fuTcirostris is the possession of three supra- 
oculars, it seems highly probable that Gadow observed this character in his specimen from the Nevado. 
In such case the specimen would be dugesii, the only species in that area that possesses this character. 



170 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Family ANELYTROPSIDAE Cope 

Anelytropsidae Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 228; Ann. 
Rep. U. S. Nat. Mus. for 1898 (1900), p. 665. 

Genera. — A single genus, Anelytropsis Cope, is known. 
Range. — Foothills of the plateau in central eastern Mexico. 

Genus ANELYTROPSIS Cope 

Attelytropsis Cope, Proc. Amer. Philos. See, vol. 22, 1885, p. 380; Ann. Rep, 
U. S. Nat. Mus. for 1898 (1900), p. 667. 

Genotype. — Anelytropsis papillosus Cope. 
Bange. — Central eastern part of Mexico. 
Species. — One. 

ANELYTROPSIS PAPILLOSUS Cope 

Anelytropsis papillosus Cope, Proc. Amer. Philos. Soc, vol. 22, 1885, pp. 380-381, 
pi. (unnumbered), fig. 9 (opposite p. 184). — Smith, Univ. Kansas Sci. Bull., 
vol. 22, 1935, p. 146, pi. 24, figs. 2, 3. 

Type. — Unknown; originally two cotype specimens. 

Type locality. — Near Jalapa, Veracruz. 

Range. — Eastern San Luis Potosi and eastern central Veracruz. 
Reported from San Luis Potosi: 20 miles south of Valles; Veracruz: 
Jalapa, Motzorongo. 

Family TEIIDAE Gray 

Teiidae Gray, Philos. Mag., ser, 2, vol. 2, 1827, p. 55. 

Genera. — About 40 genera, 3 of which, Cnemidophorus, Ameiva, 
and Gymnophthalmus, occur in Mexico. 

KEY TO MEXICAN GENERA OF TEIIDAE 

1. Anterior nasal plates not separated by a frontonasal; frontoparietal present; 

inner finger well developed; scales granular; eyelids present 2 

Nasal plates widely separated by a frontonasal; inner finger absent; no eyelids; 
scales cycloid, quincuncial; prefrontals present but no frontoparietals. 

Gymnophthalmus (p. 192) 

2. Central gular scales not or scarcely enlarged as compared with adjacent lateral 

scales; tongue somewhat widened posteriorly, and notched behind; no basal 

sheath between base of tongue and larynx Cnemidophorus (p. 174) 

Central gular scales enlarged, often abruptly so; tongue not, or less widened 
posteriorly, a basal sheath evident between the base of tongue and larynx. 

Ameiva (p. 170) 
Genus AMEIVA Meyer 

Ameiva Meyer, Synopsis reptilium . . . sistema generum methodum . . ., 
1795, p. 27. 

Genotype. — Lacerta americana Seba [=Ameiva ameiva (Linnaeus)]. 
Range. — Tamaulipas and Jalisco to Brazil; West Indies. 
Species. — About 13, with some 28 forms. Two species with 11 
subspecies occur in Mexico. 



CHECKLIST OF REPTILES OF MEXICO 171 

KEY TO MEXICAN SPECIES OF AMEIVA 

1. Outer row of ventrals considerably smaller than the others. 

f estiva edwardsii (p. 174) 
Outer row of ventrals as large as the others 2 

2. Preanal scales in 2 rows 3 

Preanal scales in one row or with no more than 1 posterior scale divided. _ 9 

3. Two rows of granules between third supraocular and superciliaries; third 

supraoculars generally completely separated from frontoparietals by 

granules undulata parva (p. 173) 

A single row of granules between third supraoculars and superciliaries; third 
supraoculars in contact with frontoparietals anteriorly 4 

4. Median gulars abruptly enlarged 6 

Median gulars little enlarged, gradually merging with lateral gulars 5 

5. Upper lateral vertical light lines from axilla to groin, 12 or more light lines. 

iindulata gaigeae (p. 172) 
Upper lateral vertical lines from axilla to groin, 1 1 or less light lines. 

iindulata hartwegi (p. 171) 

6. Median gulars irregular or no more than 2 regular (87 percent) ; lamellae 

under fourth toe 28 or more; no upper lateral light stripe, although large 

spots take its place in adult males undulata podarga (p. 172) 

At least 3 median gulars regular (100 percent in all except u. aviphigramma, 
with 87 percent); lamellae variable; upper lateral light stripe present or 
absent 7 

7. Dorsolateral dark stripes present except in some large adults, in which the 

upper lateral light spots if present are much narrower than the spaces between 
them; lamellae on the fourth toe usually (88 percent) 27 or less. 

undulata stuarti (p. 173) 

No dorsolateral dark stripes; upper lateral light spots as wide as or wider 

than spaces between, or represented by a continuous upper lateral light 

stripe; lamellae variable 8 

8. Upper lateral light spots or lines not contacting dorsolateral light area or 

line in adult males, separated by a narrow dark area; lateral gulars not or 

scarcely enlarged undulata amphigramma (p. 172) 

Upper lateral light spots merged with dorsolateral light line in adult males; 
lateral gulars markedly enlarged undulata thoraasi (p. 173) 

9. Last preanal scale generally (86 percent) divided; lateral markings showing 

little tendency to be arranged vertically; middorsal markings greatly re- 
duced undulata dextra (p. 173) 

Last preanal scale generally entire; lateral markings tending to be arranged 
vertically; middorsal markings well developed 10 

10. Upper lateral light spots in adult males large, wider than intervening dark 

spaces; generally (95 percent) 5 or less rows of preanals. 

undulata sinistra (p. 174) 

Upper lateral light spots in adult males small, narrower than intervening 

dark spaces in all males; frequently (65 percent) 6 or more rows of preanals 

luidulata undulata (p. 174) 

AMEIVA UNDULATA HARTWEGI Smith 

Ameiva undulata hartwegi Smith, Proc. Biol. Soc. Washington, vol. 53, 1946, 
p. 55. —Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 35-30, 
pi. 2, fig. b. 
861316—50 12 



172 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Type.— U.S.N. M. No. 108600; H. M. Smith collector. 

Type locality. — "Across the Rio Usumacinta from Piedras Negras, 
Guatemala, in Chiapas, Mexico." 

Range. — Atlantic slopes of Mexico and Guatemala from the vicinity 
of the southeastern end of Laguna de Terminos south and eastward 
across the base of the Yucatan Peninsula to northwestern Honduras. 
Recorded in Mexico only from Chiapas (the type locality) ; it probably 
occurs also in Campeche and Quintana Roo. 

AMEIVA UNDULATA GAIGEAE Smith and Laufe 

Ameiva undulata gaigeae Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31, 
1946, pp. 37-39, fig. IC, pi. 2C. 

IV^6— EHT-HMS No. 11927; Progreso, Yucatan; Hobart M. 
Smith collector. 

Type locality. — Progreso, Yucatan. 

Range. — Northern half of the Yucatan Peninsula and southward 
to the island Carmen along the extreme eastern coast. Reported 
from Yucatan: Chichen Itza, La Vega, Tunkas; Quintana Roo: 
Coba, Mujeres Island; Campeche: Champot6n. 

AMEIVA UNDULATA PODARGA Smith and Laufe 

Ameiva undulata podarga Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31, 
1946, figs. ID, 2A, pp. 40-43. 

Ti/pe.— EHT-HMS No. 14471; Hobart M. Smith and David H. 
Dunlde collectors. 

Type locality. — Seven miles west of Ciudad Victoria, Tamaulipas. 

Range. — Known from southern Tamaulipas and eastern San Luis 
Potosi and probably extends into Veracruz and Hidalgo lowlands. 
Reported from Tamaulipas: Alta Mira, Victoria, Hacienda La 
Clementina near Forl6n, Antiguo Morelos; San Luis Potosi: near 
Ciudad del Maiz, Rio Guayala near Magiscatzin, near Valles, Huichi- 
huayan, Tamazunchale. 

AMEIVA UNDULATA AMPHIGRAMMA Smith and Laufe 

Ameiva undulata amphigramma Smith and Laufe, Trans. Kansas Acad. Sci., 
vol. 48, 1945, pp. 338-344; Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 43-47, 
pi. 1, figs. C, D. 

Type.— EUT-UMS No. 11983; H. M. Smith collector. 

Type locality. — San Andres Tuxtla, Veracruz. 

Range. — Northern Veracruz southward at low elevations to the 
Isthmus of Tehuantepec, westward into valleys extending into extreme 
eastern Oaxaca and probably northeastern Puebla. Reported from 
Veracruz: Atoyac, Boca del Rio, Cuatotolapam, Lake Catemaco, 
Potrero Viejo, Rodriguez Clara, San Andres Tuxtla, etc.; Oaxaca: 
Cosolapa, Matias, Agua Fria, Tuxtepec; Tabasco: La Venta; Puebla: 
1 mile northeast of Huauchinango. 



CHECKLIST OF REPTILES OF MEXICO 173 

AMEIVA UNDULATA THOMASI Smith and Lanfe 

Ameiva undulata thomasi Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31 
1946, pp. 47-50. 

Ty^e.— EHT-HMS No. 15327, H. D. Thomas, collector. 

Type locality. — La Libertad, Chiapas, near Rfo Cuilco where it 
crosses the Guatemalan border. 

Range. — Upper tributaries of Rio Grijalva in the interior of Chiapas 
and adjacent Guatemala. Recorded in Mexico only from the type 
locality. 

AMEIVA UNDULATA STUARTI Smith 

Ameiva undulata stuarti Smith, Proc. Biol. See. Washington, vol. 53, 1940, pp. 
55-56. —Smith and Latjfe, Univ. Kansas Sci. Bull., vol. 31, 1946, pp. 50-51, 
fig. IB, pi. 1, fig. B. 

r?/2)e.— U.S.N.M. No. 108601, H. M. Smith collector. 

Type locality. — Palenque, Chiapas. 

Range. — Atlantic slopes of Mexico from the middle of the Istlimus 
of Tehuantepec eastward in the lowlands to the southern borders of 
Laguna de Terminos and to Tenosique, Tabasco; southward up the 
valley of the Rio Grijalva at least as far as Tuxtla Gutierrez, Chiapas. 
Reported from Chiapas: Palenque, San Ricardo, Tuxtla Gutierrez; 
Campeche: Balchacaj, Tres Brazos; Tabasco: Tenosique, Frontera, 
Teapa; Oaxaca: Isthmus of Tehuantepec (intergrades?). 

AMEIVA UNDULATA PARVA Barbonr and Noble 

Ameiva undulata parva Barbour and Noble, Bull. Mus. Comp. Zool., vol. 59, 
1915, pp. 476-477.— Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31, 1946, 
pp. 51-54, fig. lA, pi. 2, fig. A. 

Tijpe. — Mus. Comp. Zool. No. 5831; Van Patten collector. 

Type locality. — Guatemala (restricted by Smith and Laufe to 
Mazatenango). 

Range. — Pacific slopes from the Isthmus of Tehuantepec in Oaxaca 
to Costa Rica. Reported in Mexico from Oaxaca: near Niltepec; 
Chiapas: Tapachula, Tonala, La Esperanza, Huehuetan. 

AMEIVA UNDULATA DEXTRA Smith and Laufe 

Ameiva undulata dextra Smith and Laufe, LTniv. Kansas Sci. Bull., vol. 31, 1946 
pp. 54-59, fig. 2C. 

Type.— EB.T-B.MS No. 11966; E. H. Taylor and H. M. Smith 
collectors. 

Type locality. — Near Rinc6n, Guerrero. 

Range. — Southern slope of the Sierra Madre del Sur. Known from 
Oaxaca and Guerrero. Reported from Oaxaca: Juquila; Guerrero: 
Chilpancingo, Cocoyul, Los Cajones, Rincon, Organos, El Treinta, 
Acapulco, 8 miles east of Coyuca. 



174 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

AMEIVA UNDULATA SINISTRA Smith and Laufe 

Ameiva undulaia sinistra Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31, 1946, 
pp. 59-62. 

Type.—EUT-RMS No. 11908; H. M. Smitli collector. 

Type locality. — Manzanillo, Colima. 

Range. — Pacific slopes from Jalisco to the arid Balsas basin in 
Michoac^n, thence inland along the northern drainage of the Rio 
Balsas to Puebla. Reported from Colima: Colima, Queseria, Hda. 
Paso del Rio, Salvador, Pascuales, Periquillo, Manzanillo; Jalisco: 
Tenacatita, Ixtapa, Tenacatita Bay; Michoacdn: Uruapan; Puebla: 
Chiautla; Morelos: Puente de Ixtla. 

AMEIVA UNDULATA UNDULATA (Wiegmanti) 

Cnemidophorus undulatus Wiegmann, Herpetologia Mexicana, 1834, p. 27. 
Ameiva undulata, Gray, Catalogue of the specimens of lizards in the British 

Museum, 1845, p. 20. 
Ameiva undulata undulata, Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31, 

1946, pp. 62-64, fig. 2B. 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, by inference. Restricted by Smith (Proc. 
Biol. Soc. Washington, vol. 53, 1940, p. 56) to Tehuantepec. 

Range. — Pacific slopes of the Isthmus of Tehuantepec as far west 
as Puerto Angel, and eastward to Niltepec. Known only in the state 
of Oaxaca: El Lim6n, Palmar, Tres Cruces, Cafetal Concordia, Juchi- 
tin, Puerto Angel, Totontepec, Tehuantepec, Rancheria La Manga. 

AMEIVA FESTIVA EDWARDSII Boconrt 

Ameiva edwardsii Bocourt, Ann. Sci. Nat., ser. 5, vol. 17, 1873, art. 17, pp. 1-2. 
Ameiva f estiva edwardsii, Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 471, 

1943, p. 21, fig. 7. — Smith and Laufe, Univ. Kansas Sci. Bull., vol. 31, 1946, 

pp. 64-66. 

Type. — Mus. Hist. Nat. Paris; Commission Scientifique collector. 

Type locality. — Izabal and Santa Maria de Panzos, near Rio 
Polochic, Guatemala; here restricted to Panz6s. 

Range. — Atlantic foothills from the Isthmus of Tehuantepec 
through Honduras, in heavy forests. Recorded in Mexico only from 
Chiapas: Ixtacomitan, Palenque, across the Rio Usumacinta from 
Piedras Negras (Peten, Guatemala). 

Genus CNEMIDOPHORUS Wagler 

Cnemidophorus Wagler, Natiirliches System der Amphibien . . ., 1830, p. 154. — 
Burt, U. S. Nat. Mus. Bull. 154, 1931. — Burger, Chicago Acad. Sci. Nat. 
Hist. Misc., No. 65, 1950, pp. 1-9. 

Aspidoscelis Fitzinger, Systema reptilium, 1843, p. 20 (type, Lacerta sexlineata 
Linnaeus) . 

Verticaria Cope, Proc. Amer. Philos. Soc, vol. 11, 1869, p. 158 (type, Cnemi- 
dophorus hyperythrus Cope). 



CHECKLIST OF REPTILES OF MEXICO 175 

Genotype. — SejJS murinus Lauren ti. 

Range. — North America south from Oregon, Wisconsin, and Mary- 
land through Central America to southern Brazil and Bolivia. 

Species. — About 18, and a total of about 47 species and subspecies; 
15 species and 41 forms, as here recognized, occur in Mexico. These 
18 species represent 5 distinct groups, all but one of which (the 
lemniscatus group, restricted to Central and South America, of 3 
species and 5 forms as recognized by Burt, op. cit.), occur in Mexico. 

KEY TO MEXICAN SPECIES OF CNEMIDOPHORUS 

1. Only one frontoparietal scale " 2 

Two frontoparietals 9 

2. Dorsal surface unicolor, not striped or reticulated. hyperythrus pictus (p. 186) 
Dorsal surface striped or reticulated, not unicolor 3 

3. Lateral stripes more or less broken; at least a few spots or cross bars present; 

body often tessellated; adults large ceralbensis (p. 188) 

Lateral stripes distinct and in even, continuous lines; spots and cross bars 
absent; size always small 4 

4. Usually 2 or 3 longitudinal light lines in vertebral area 5 

Usually 1 median dorsal light line, forked anteriorly or not 6 

5. Usually 3 light lines along middle of back; second supraocular usually without 

granules between it and frontal hyperythrus hyperythrus (p. 187) 

Usually 2 light lines; second supraocular usually at least in part separated 
from frontal by granules hyperythrus beldingi (p. 18G) 

6. Dorsal stripe not of same intensity and width as lateral stripes but fainter and 

usually narrower 7 

Dorsal stripe of same intensity and width as lateral stripes 8 

7. More than half of second supraocular usually in contact with frontal. 

hyperythrus schmidti (p. 187) 
Half or less than half of second supraocular usually in contact with frontal. 

hyperythrus beldingi (p. 186) 

8. Ground color of sides (between the 2 distinct lateral stripes) reddish gray; 

dorsal ground color gray or black, abruptly contrasted with lateral ground 

color hyperythrus danheimae (p. 186) 

Ground color of sides not usually reddish gray, but often reddish or brownish; 
dorsal ground color of same general shade as that of the sides and not in 
sharp contrast to it h3^erythrus caeruleus (p. 186) 

9. Supraoculars normally 3; striped at least in the young 10 

Supraoculars normally 4, or, if 3, no stripes at any stage, but a pattern of small, 

scattered light spots on a dark background 15 

10. Scales between midventral scutes and anus 4 to 8; snout- vent measurement 

never exceeding 92 mm 12 

Scales between midventral scutes and anus 9 to 15; snout-vent measurement 
reaching 138 mm 11 

11. Adults and subadults spotted, the spots often dim.guttatus guttatus (p. 179) 
Adults and subadults (as well as young) striped, the stripes sometimes broken 

up into spots guttatus immutabilis (p. 180) 

" The species possessing this character were formerly regarded as members of the genus Vertkaria Cope. 
Burt (U. S. Nat. Mus. Bull. 154, 1931, p. 14) has concluded, however, that the group is not a natural one, 
since one species (ceralbensis) is related to the tesselatus group, while the others are related to the sezlineatus 
group. The character is, furthermors, subject to a slight variation. 



176 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

12. An accessory scute between frontoparietal and parietal. 

deppii cozumelus (p. 179) 
No accessory scute 13 

13. Femoral pores usually 17 or fewer (81-82 percent) 14 

Femoral pores usually 18 or more (94 percent) deppii deppii (p. 178) 

14. Lower preocular generally (79 percent) in contact with loreal; a broad mid- 

dorsal light band deppii lineatissimus (p. 179) 

Lower preocular generall (73 percent) not in contact with loreal; narrow light 
stripes on back as on sides deppii oligoporus (p. 179) 

15. Enlarged scales immediately preceding gular fold (mesoptychials) small (not 

more than 3 times diameter of smallest scales in median part of pregular 
fold) ; or, if larger, no evidence whatever of longitudinal light lines, or else 

at least a few dark spots on throat below level of ear openings 16 

Mesoptychials larger; a lined pattern always present in the young, generally 
in adults; no dark spots on throat below level of ear openings; or, if less than 
3 times diameter of smallest scales in median part of pregular fold, 7 light 
lines and intervening dark spaces absolutely straight, continuous and 
disconnected 28 

16. Mesoptychials moderately or considerably enlarged, more than 3 times 

diameter of smallest scales in median part of pregular fold 17 

Mesoptychials smaller 19 

17. No evidence of a longitudinal arrangement of Unes on back 18 

Dorsal surface distinctly lined tesselatus (p. 188) 

18. No spotting in area above level of tympanum and anterior to insertion of 

forearm, spots relatively well defined, very small, not yellowish. 

bacatus (p. 187) 
Spots present in area above tympanum and anterior to insertion of forearm; 
spots often obscure and diffuse, usually orange or yellowish. 

catalinensis (p. 188) 

19. Ventral surfaces of tail and limbs more or less suffused with red or pink. 20 
Not so „ 21 

20. Black markings on temporal regions absent or few or ill-defined; dorsal and 

lateral markings on body showing both transverse and longitudinal arrange- 
ment in at least large specimens; each dark unit or spot square or rectangular 

in outline, not noticeably rounded tigris rubidus (p. 191) 

Black markings on temporal region well defined; dorsal and lateral markings 
on body showing a predominantly longitudinal arrangement; back covered 
by alternate longitudinal chains of black and white, individual spots in 
black chain rarely square or rectangular tigris celeripes (p. 190) 

21. Dorsal markings consisting of either fine reticulations or light spots, and these 

markings without longitudinal arrangement 22 

Dorsal markings with at least some indication of longitudinal arrange- 
ment 23 

22. Labials deep or moderate brown or black tigris martyris (p. 191) 

Labials hght gray, slate or light brown tigris canus (p. 190) 

23. Dorsal light lines only 3 on each side (exclusive only of a pair of lateral lines), 

twice as wide as interspaces mazimus (p. 188) 

Dorsal light lines more numerous or narrower, or both 24 

24. Dorsal pattern of 6-8 persistent longitudinal light stripes 25 

Dorsal pattern of fewer than 6 unbroken light stripes; at least lower lateral 

stripe on each side broken by dark vertical bars 27 



CHECKLIST OF REPTILES OF MEXICO 177 

25. Dark fields between dorsal light stripes unspotted and unbroken throughout 

life; entire ventrum black in adults tigris aethiops (p. 189) 

At least some of dark fields broken into rows of spots; ventrum not entirely 
black in adults 26 

26. Middorsal dark field or row of spots distinctly wider than paravertebral light 

stripes; throat white, spotted or barred with black. 

tigris multiscutatus (p. 191) 
Middorsal dark field or row of spots as wide as or slightly narrower than 
paravertebral light stripes; throat and chest black; belly spotted. 

tigris gracilis (p. 188) 

27. Four longitudinal light stripes persisting unbroken in middorsum of adults, 

the dark fields between represented by rows of black spots; sides with 

bold vertical bars tigris tigris (p. 189) 

Six longitudinal light stripes persisting unbroken in adults, or, if any broken, 
dorsal pattern generally light and without any bold markings. 

tigris marmoratus (p. 190) 

28. Anterior nasal in contact with second upper labial; femoral pores never more 

than 14 42 

Anterior nasal usually separated from second upper labial; if in contact, 
femoral pores usually more than 14 29 

29. Enlarged scales preceding gular fold relatively small, frequently grading into 

granular scales of fold 30 

Enlarged scales preceding gular fold relatively large, abruptly differentiated 
from granular scales of throat 31 

30. Postantebrachials enlarged; ventral surfaces light bluish in adults; juvenile 

striped pattern retained throughout life; maximum snout- vent length 70 
mm.; dorsal scales relatively large, 50 to 70 in a transverse row between 

enlarged belly plates inomatus (p. 184) 

Postantebrachials not enlarged; belly, chest, or gular region dark (black) in 
adults; snout-vent length reaching 95 mm.; dorsal scales 85 or more in a 
transverse row tigris aethiops ^^ (p. 189) 

31. In addition to a dorsolateral light stripe on tail, a lateral light stripe present, 

bordered below by a black line (near base of tail) continuing onto otherwise 
unicolor posterior surface of thigh; no light spots in dark field between 
stripes in adults; postantebrachials not enlarged in either sex; neither sex 

dark blue or orange below sexlineatus (p. 184) 

No lateral light stripe on tail, at least not distinguishable from ventral tail 
color; thighs mottled or unicolor, no posterior light stripe except sometimes 
feebly mdicated at insertion of leg; light spots often present in dark fields 
between stripes; postantebrachials distinctly enlarged generally, sometimes 
less so in females; males frequently dark blue on belly or chest, or else cream, 
not at all bluish 32 

32. Only 5 light lines, including a vertebral, 2 dorsolateral and 2 lateral lines 

through the ear; ventral surfaces cream even in males, never blue; light lines 
very broad, more than half width of intervening dark spaces in adults; no 
marking between light lines, except series of small spots in dark spaces in 

juveniles (disappearing in adults) burti (p. 185) 

Not so 33 



"Young only key out here; adults key out through couplet 16 to couplet 25. 



178 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

33. Space between paravertebral light stripes (i. e., those originating even with 

parietal scales, not with median interparietal) narrower or at least no wider 
than space between dorsolateral and upper lateral light stripes; if stripes not 

visible, they are replaced by dorsal pattern of dark cross bars 34 

Space between paravertebral light stripes wider; if stripes not visible, they are 
replaced by numerous scattered light spots 37 

34. Adults without a cross-barred pattern 35 

Adults cross-barred 36 

35. Dark interspaces or rows of dark spots continuing to area above hind legs. 

sackii stictogrammus (p. 183) 

Dark interspaces or rows of spots disappearing anterior to area above hind 

legs sackii semifasciatus (p. 184) 

36. Postantebrachials generally scutelike sackii so alaris (p. 182) 

Postantebrachials generally polygones, little enlarged_sackii australis (p. 181) 

37. Hind legs mottled in adults, but not with small light spots; dorsal pattern 

cross-barred in adults sackii sackii (p. 180) 

Hind legs with small light spots in adults; dorsal pattern not cross-barred. 38 

38. Stripes absent in adult, replaced by numeroiis, scattered light dots especially 

toward rear half of body sackii bocourti (p. 181) 

Stripes present in adults 39 

39. Collar black in adults; stripes obsolete except for lateral stripes; keels on 

caudal scutes nearly parallel sackii raariarum (p. 183) 

Collar light; stripes not obsolete; keels on caudal scales strongly oblique. _ 40 

40. Ventral surfaces of tail and limbs pink sackii angusticeps (p. 183) 

Ventral surfaces of tail and limbs cream or white 41 

41. Stripes becoming rows of light spots in adults.- sackii communis (p. 182) 
Stripes persistent sackii gularis (p. 183) 

42. Femoral pores 11-15 in known specimens labialis (p. 185) 

Femoral pores 19-20 in known specimens gadovi (p. 185) 

DEPPn GROUP 

Species. — Six forms, belonging to two species, now recognized. 
Range. — Central Veracruz on the Atlantic coast and Nayarit on the 
Pacific southward to Costa Rica. 

CNEMIDOPHORUS DEPPII DEPPH Wiegmann 

Cnemidophorus deppii Wiegmann, Herpetologia Mexicana, 1834, p. 29. — Bo- 
court, Mission scientifique au Mexique . . . , fitudes sur les reptiles, livr, 

3, 1874, p. 281, pi. 20c, fig. 5, pi. 20d, fig. 1. 
Cnemidophorus deppei [deppei], Gadow, Proc. Zool. Soc. London, 1906, pp. 309- 

316, figs. 71 D-G, 72 A-G. 
Cnemidophorus deppii deppii, Burt, U.S. Nat. Mus., Bull. 154, 1931, pp. 56-63 

(part), fig. 14, right. — Hartweg and Oliver, Occ. Pap. Mus. Zool. Univ. 

Michigan, No. 359, 1937, pp. 1-3. — Smith, Ann. Carnegie Mus., vol. 30, 

1944, pp. 90-91. 
Cnemidophorus decemlineatus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 

1860. p. 482 (U.S.N.M. No. 6058, 12 cotypes; Nicaragua; C. Wright collector). 
Cnemidophorus laiivittis Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 94 

(U.S.N.M. No. 30227; Juchitdn, Oaxaca; F. Sumichrast collector). 
Cnemidophorus alfaronis Cope, Proc. Acad. Nat. Sci. Philadelphia, 1894, p. 199 

(Amer. Mus. Nat. Hist. No. 16315; San Mateo, Costa Rica; A. Alfaro 

collector). 



CHECKLIST OF REPTILES OF MEXICO 179 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico, restricted to Tehuantepec, Oaxaca. 

Range. — Southern Guerrero on Pacific slopes and the Salama Basin 
in Guatemala on Atlantic slopes, south to Costa Rica. Recorded in 
Mexico only from the states of Guerrero, Oaxaca, and Chiapas. 

CNEMIDOPHORUS DEPPII COZUMELUS Gadow 

Cnemidophorus deppei cozumela Gadow, Proc. Zool. Soc. London, 1906, p. 136. 
Cnemidophorus deppii cozumelus, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 

63-65, fig. 14 left.— Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 292, 

1934, pp. 12-13, fig. 1. 

Type.— Qvii. Mus. Nat. Hist. Nos. 1886.4.15.17-20, four cotypes. 
Type locality. — Cozumel Island, east coast of Quintana Roo. 
Range. — The eastern portion of the Yucatan Peninsula. Recorded 
in Mexico only from Quintana Roo (Mujeres and Cozumel Island) . 

CNEMIDOPHORUS DEPPII LINEATISSIMUS Cope 

Cnemidophorus lineatissimus Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 94. 
Cnemidophorus deppii lineatissimus, Hartweg and Oliver, Occ. Pap. Mus. Zool. 
Univ. Michigan, No. 359, 1937, p. 2, 

Type.— U.S. '^.M. Nos. 24937-40, 32299-32314, 20 cotypes; Hoge 
collector. 

Type locality. — Colima and Guadalajara (Jalisco), here restricted 
to Colima, Colima. 

Range. — Pacific slopes from Nayarit to central and perhaps southern 
Guerrero. The range interdigitates with that of C. d. deppii in central 
Guerrero. Recorded only from Nayarit, Jalisco, Colima, Michoac^n, 
Guerrero, and Puebla (Chiautla). 

CNEMIDOPHORUS DEPPII OLIGOPORUS Smith 

Cnemidophorus deppii oligoporus Smith, Publ. Field Mus. Nat. Hist., zool. ser., 
vol. 24, 1939, pp. 26-27; Smith and Burger, Trans. Kansas Acad. Sci., vol. 
53, 1950, pp. 173-174. 

Type. — Chicago Nat. Hist. Mus. No. 29145; Julius Friesser. 

Type locality. — Perez, Veracruz. 

Range. — Atlantic slopes from northern Veracruz to Campeche. 
Recorded only from the states of Veracruz and Campeche (Ciudad del 
Carmen) . 

CNEMIDOPHORUS GUTTATUS GUTTATUS Wiegmann 

Cnemidophorus guttatus Wiegmann, Herpetologia Mexicana, 1834, p. 29. — 
Bocourt, Mission scientifique au Mexique . . ., Etudes sur les reptiles, 
livr. 3, 1874, p. 285, pi. 20c, fig. 4.— Burt, U.S. Nat. Mus. Bull. 154, 1931, 
pp. 66-74 (part). 

Cnemidophorus guttatus guttatus, Cope, Trans. Amer. Philos. Soc, vol. 17, 1892, 
p. 32.— Gadow, Proc. Zool. Soc. London, 1906, pp. 309, 325-326, fig. 75.— 
Hartweg and Oliver, Occ. Pap. Mus. Zool. Univ. Michigan, No. 359, 1937, 
p. 3. 



180 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Cnemidophorus unicolor Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 93 
(U. S. Nat. Mus.; "West Tehuantepec," Oaxaca; F. Sumichrast collector). 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Alexico, here restricted to Veracruz, Veracruz. 

Range. — Atlantic slopes, from central Veracruz to the Isthmus of 
Tehuantepec. Recorded only from the states of Veracruz and 
Oaxaca. 

CNEMIDOPHORUS GUTTATUS IMMUTABILIS Cope 

Cnemidophorus immutahilis Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 93 
Cnemidophorus guttatus immutahilis, Cope, Trans. Amer. Philos. Soc, vol. 17 

1892, p. 31.— Gadow, Proc. Zool. Soc. London, 1906, pp. 309, 326-327 

figs. 74, 75. — Hartweg and Oliver, Occ. Papers Mus. Zool. Univ. Michigan 

No. 359, 1937, pp. 3-7. 
Cnemidophorus microlepidopus Cope, loc. cit. (U.S.N.M. No. 30187; "West 

Tehuantepec," here restricted to the city of Tehuantepec, Oaxaca; F. 

Sumichrast collector). 
Cnemidophorus guttatus striatus Gadow, Proc. Roy. Soc. London, vol. 72, 1903, 

p. 115 (Brit. Mus. Nat. Hist., "Isthmus of Tehuantepec," here restricted to 

the city of Tehuantepec, Oaxaca; Hans Gadow collector). 

Type.— U.S. ISl.M. No. 30141; F. Sumichrast collector. 

Type locality. — ^"West Tehuantepec," Oaxaca, here restricted to 
the city of Tehuantepec. 

Range. — Pacific slopes from Colima to Chiapas. Recorded only 
from the states of Colima (Manzanillo), Michoacan, Guerrero, Morelos, 
Oaxaca, and Chiapas (Tonala). 

sexlineatus qjroup 

Species. — Six, with a total of 16 forms as at present recognized. 

Range. — Southern California, southeastern Wyoming, Wisconsin, 
and Maryland southward to the Gulf of Mexico, to Guatemala and 
into northwestern Baja California. 

Remarks. — The arrangement presented herewith for this group is 
far from final. We have, however, made an attempt to correlate 
zoogeography with variations pointed out by others, and the tentative 
conclusions we trust will at least be a step forward. The present pic- 
ture does make some geographic sense, as previous arrangements 
have not. 

CNEMIDOPHORUS SACKII SACKU Wiegmann 

Cnemidophorus sackii Wiegmann, Herpetologia Mexicana, 1834, pp. 28-29. — 

Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 249-251.83 
Cnemidophorus sexlineatus sackii, Burt, Proc. Biol. Soc. Washington, vol. 44, 

1931, pp. 73-78. 
Cnemidophorus sackii sackii, Smith, Journ. Washington Acad. Sci., vol. 39, 1949, 

pp. 41-42. 

•» See also discussions in Taylor, Univ. Kansas Sci. Bull., vol. 24, 1938, pp. 520-523; Schmidt and Stuart, 
Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1941, pp. 244-255; and Stuart, Misc. Publ. Mus. Zool. Univ. 
Michigan, No. 69, 1948, pp. 57-59. 



CHECKLIST OF REPTILES OF MEXICO 181 

Cnemidophoriis mexicanus Peters, Monatsb, Akad. Wiss. Berlin, 1869, pp. 62-63 

(Zool. Mus. Berlin, three cotypes; Uhde collector; "Mexico," ^* here restricted 

to Matamoros, Puebla). 
Cnemidophorus costatus Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, pp. 95-96 

(U.S.N. M, No. 31359; Boucard collector; "Mexico," here restricted to 

Cuernavaca, Morelos). 
Cnemidophorus viexicanus balsas Gadow, Proc. Zool. Soc. London, 1906, pp. 363- 

367, fig. 83 (many cotypes, Chicago Nat. Hist. Mus. and Brit. Mus. Nat. 

Hist.; lectotype here designated as Brit. Mus. Nat. Hist. No. 1906.6.1.71; 

Hans Gadow collector; type locality here restricted to Cuernavaca, Morelos). 

Type. — Zool. Mus. Berlin No. 884; F. Deppe collector. 
Type locality. — "Alexico," restricted to Cuernavaca, Morelos. 
Range. — The Rio Balsas Valley. Recorded from numerous locali- 
ties in Morelos, Guerrero, Puebla, and Michoacan. 

CNEMIDOPHOKUS SACKII BOCOURTI Boulenger 

Cnemidophorus bocourti Boulenger, Catalogue of the lizards in the British 

Museum, vol. 2, 1885, p. 367. 
Cnemidophorus communis bocourti, Gadow, Proc. Zool. Soc. London, 1906, pp. 

356-358, fig. 80. 
Cnemidophorus sackii bocourti, Smith, Rev. Soc. Mex. Hist. .Nat., vol. 7, 1946, 

pp. 69-70. 
Cnemidophorus sexlineatus sacHi, Bocourt, Mission scientifique au Mexique . . ., 

fitudes sur les reptiles, livr. 4, 1874, pp. 276-277, pi. 20c, fig. 6. 
Cnemidophorus sackii, Schmidt and Stuart, Publ. Field Mus. Nat. Hist., zool. 

ser., vol. 24, 1941, pp. 244-245. 
Cnemidophorus communis copei, Gadow, Proc. Zool. Soc. London, 1906, pp. 350- 

351 (specimen from Santo Domingo, Oaxaca, only). 
Cnemidophorus motaguae Sackett, Notulae Nat., No. 77, 1941, pp. 1-4 (Acad. 

Nat. Sci. Philadelphia No. 22143; J. T. Sackett collector; Motagua River 

Valley 10 kilometers northeast of Zacapa, Zacapa, Guatemala). 

Type.— Brit. Mus. Nat. Hist., three cotypes; No. 1857.10.28.81 here 
designated lectotype. 

Type locality. — "Mexico" and "California," here restricted to Oax- 
aca, Oaxaca. 

Range. — Uplands from central Oaxaca through central Chiapas 
into central Guatemala; range perhaps discontinuous. Recorded in 
Mexico only from Oaxaca: Oaxaca, Santo Domingo; and Chiapas: 
Piedra Parada. 

CNEMIDOPHORUS SACKII AUSTRALIS Gadow 

Cnemidophorus communis australis Gadow, Proc. Zool. Soc. London, 1906, pp. 

352-356, figs. 62C, D, 64C, D, 65F, 79B, C. 
Cnemidophorus mexicanus typica Gadow, Proc. Zool. Soc. London, 1906, pp. 

360-363, fig. 81A-D, fig. 82. 

M Gadow {op. cit., p. 358) states that Peters's cotypes are immature and therefore not readily placed. The 
original description is of little assistance. We do not follow Gadow's allocation of the name, with a distinct 
southern race, for two reasons: (1) The name can simply be disposed of, if the types are unidentifiable, by 
placing it in synonsrmy; and (2) we believe it quite probable that the types actually came from southwestern 
Puebla in the Balsas Basin, where a number of Peters's species are known to be restricted. This latter area 
is occupied by C. s. sackii, of which Peters's name can reasonably be held a synonym. 



182 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Type. — Numerous cotypes, Brit. Mus. Nat. Hist, and Chicago Nat. 
Hist. Mus.; lectotype here designated as Brit. Mus. Nat. Hist. No. 
1906.7.19.11. 

Type locality. — Not specified, Lagunas or Cuicatlan, Oaxaca; here 
restricted to Cuicatlan, Oaxaca. 

Range. — Foothills about the central Oaxacan highlands, excluding 
the Balsas Basin, in the upper headwaters of the B,io Papaloapam 
and Rio Tehuantepec, and near the Isthmus of Tehuantepec. 

CNEMIDOPHORUS SACKII COMMUNIS Cope 

Cnemidophorus communis Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 95. 
Cnemidophorxis communis occidentalis Gadow, Proc. Zool. Soc. London, 1906, pp. 

339-346, fig. 69, 77A-F, 78B, 79A (many cotypes, Brit. Mus. Nat. Hist. 

and Chicago Nat. Hist. Mus.; lectotype here designated as Brit. Mus. Nat. 

Hist. No. 1892.2.8.33, from IxtMn, Nayarit). 
Cnemidophorus communis copei Gadow, Proc. Zool. Soc. London, 1906, pp. 346- 

352, fig. 78A, C, E (cotypes in Brit. Mus. Nat. Hist, and Chicago Nat. Hist. 

Mus.; lectotype here designated as Brit. Mus. Nat. Hist. No. 1906.7.19.5, 

from Colima, Colima; a cotype from Santo Domingo de Guzmdn we refer to 

C. s. bocourii). 

Type. — Numerous cotypes, all lost. 

Type locality. — Not definitely stated: Colima, Guadalajara, C6r- 
doba, Coban (Guatemala) or San Antonio (Texas) ; here restricted to 
Colima, Colima. 

Range. — Most of the central plateau of Mexico, from Chiliuahua 
and presumably Nuevo Le6n southward to central Puebla ; the Pacific 
coast from the Balsas basin northward to southern Sinaloa and perhaps 
Sonora. Recorded from ? Sonora, Chihuahua, Durango, Coahuila, 
Tamaulipas, San Luis Potosi, Zacatecas, Hidalgo, Queretaro, Guana- 
juato, Aguascalientes, Jalisco, Sinaloa, Nayarit (including Isabel Is- 
land), Colima, Michoacan, Mexico, Distrito Federal, Puebla, and 
Veracruz. 

CNEMIDOPHORUS SACKII SCALARIS Cope 

Cnemidophorus sexlineatus tigris Cope {nee Baird and Girard), Proc. Amer. 

Philos. Soc, vol. 23, 1886, p. 283 (types not designated; Chihuahua, 

Chihuahua). 
Cnemidophorus gularis scalaris Cope, Trans. Amer. Philos. Soc, vol. 17, 1892, 

p. 47. — Smith and Mittleman, Trans. Kansas Acad. Sci., vol. 46, 1943, 

pp. 246-247. 
Cnemidophorus gularis gularis verus Cope, op. cil., p. 45 (no types designated; 

Chihuahua, Chihuahua). 
Cnemidophorus gularis gularis obsoletus Cope, loc. cit. (as above). 

Type.— U.S.N. M. Nos. 8319, 14302, seven cotypes (No. 14302a, 
lectotype) ; John Potts collector. 

Type locality. — Not specifically designated, either "Mexican plateau 
south of Chihuahua," or "City of Chihuahua;" here restricted to 
Chihuahua, Chihuahua. 



CHECKLIST OF REPTILES OF MEXICO 183 

Range. — Central and eastern Chihuahua, western Coahuila and 
northern Durango. Recorded from each state mentioned. 

CNEMIDOPHORUS SACKII STICTOGEAMMUS Burger 

Cnemidophorus sackii stictogrammus, Bukger, Chicago Acad. Sci. Nat. Hist. Misc., 

No. 65, 1950, p. 5. 
Cnemidophorus gularis octolineatus, Smith (nee Baird), Handbook of lizards, 1946, 

pp. 409-412, figs. 126-127, pi. 114 (part). 

Type.— V.S.N. M. (Hensley-Burger No. 768). 

Type locality. — Yank Springs, 6 miles southeast of Ruby, Santa 
Cruz County, Ai'iz. 

Range. — Central Arizona and Mexico southward through Sonora 
and probably northern Sinaloa, and the northern parts of Chihuahua 
and Coahuila. Recorded in Mexico from Sonora and Chihuahua. 

CNEMIDOPHORUS SACKII MARIARUM Gunther 

Cnemidophorus mariarum GiJNTHER, Biologia Centrali-Americana, Reptilia and 
Batrachia, 1885, p. 28. — Stejneger, North Amer. Fauna, No. 14, 1899, pp. 
67-68.— Gadow, Proc. Zool. Soc. London, 1906, pp. 328-330. 

Type.— Brit. Mus. Nat. Hist. Nos. 1881.10.1.81-82, 86-88, five 
cotypes. 

Type locality. — Tres Marias Islands. 

Range. — Restricted to the Tres Marias Islands, on each one of which 
it has been taken. 

CNEMIDOPHORUS SACKU ANGUSTICEPS Cope 

Cnemidophorus angusticeps Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 
95.— Smith, Publ. Field Mus. Nat. Hist., zool. ser., vol. 24, 1939, pp. 24-25. 

Type.— U.S. N.M. Nos. 24876-24878, three cotypes. 

Type locality. — "Yucatan," here restricted to Chichen Itza. 

Range. — The northern and western portions of the Yucatan 
Peninsula. Recorded from Yucatan: Chichen Itza, Progreso, Tunicas; 
Quintana Roo: Coba, Cozumel Island; Campeche: Champot6n. 

CNEMIDOPHORUS SACKII GULARIS Baird and Girard 

Cnemidophorus gularis Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 

1852, p. 128.. 
Cnemidophorus gularis gularis, Cope, Trans. Amer. Philos. Soc, vol. 17, 1892, p. 

334.— Smith, Handbook of lizards, 1946, pp. 406-409, fig. 121, pi. 113. 
Cnemidophorus guttatus Hallowell (nee Wiegmann), Proc. Acad. Nat. Sci., 

Philadelphia, 1854, p. 192 (types lost; Texas, here restricted to Brownsville). 
Cnemidophorus gularis sealous Cope, Amer. Nat., vol. 26, 1892, p. 522 {nomen 

nudum). 
Cnemidophorus gularis sericeus Cope, Trans. Amer. Philos. Soc, vol. 17, 1892, 

p. 48 (U.S.N.M. No. 15650; Wm. Taylor collector; San Diego, Tex.). 



184 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Cnemidophorus gularis meeki Gadow, Proc. Zool. Soc. London, 1906, pp. 332-334 
(numerous cotypes, Brit. Mus. Nat. HLst. and Chicago Nat. Hist. Mus.; 
lectotype here designated as Chicago Nat. Hist. Mus. No. 1294, the larger 
male specimen, from Montemorelos, Nuevo Le6n). 

TV^e.— U.S.N.M. Nos. 3022, 2989, 14 cotypes (lectotype No. 
3022a) ; Colonel Graham collector. 

Type locality. — "Indianola and the Valley of the Rio Grande del 
Norte," restricted to mouth of Devils River, Tex, 

Range. — Oklahoma and most of Texas southward through Tamau- 
lipas and eastern Nuevo Le6n to northern Veracruz. Recorded in 
Mexico from Tamaulipas, San Luis Potosi, Veracruz, and Nuevo Le6n. 

CNEMIDOPHORUS SACED SEMIFASCIATUS Cope 

Cnemidophorus gularis semifasciatus Cope, Trans. Amer. Philos. Soc, vol. 17, 

1892, p. 49. 
Cnemidophorus sackii semifasciatus, Burger, Chicago Acad. Sci. Nat. Hist. Misc., 

No. 65, 1950, pp. 4-5. 
Cnemidophorus septemvittatus Cope, Trans. Amer. Philos. Soc, vol. 17, 1892, p. 40 

("El Dorado County, California," in error; restricted to Marfa, Presidio 

County, Tex.; U.S.N.M. No. 42141). 
Cnemidophorus octolineatus Schmidt and Smith (nee Baird), Publ. Field Mus. 

Nat. Hist., zool. ser., vol. 29, 1944, pp. 85-86. 

Type.— U.S.N.M. No. 9248; Lieutenant Couch collector. 
Type locality. — Agua Nueva, Coahuila. 

Range. — Big Bend area of Texas and adjacent Coahuila. Recorded 
in Mexico only from the state of Coahuila. 

CNEMIDOPHORUS INORNATUS Baird 

Cnemidophorus inornalus Baird, Proc. Acad. Nat, Sci. Philadelphia, 1858, p. 

255. — Schmidt and Owens, Publ. Field Mus. Nat. Hist., zool. ser., vol. 

29, 1944, pp. 106-107; Burger, Chicago Acad. Sci. Nat. Hist. Misc., 1950 

(in press). 
Cnemidophorus octolineatus Baird, Proc Acad. Nat. Sci. Philadelphia, 1858, p. 

255 (Pesqueria Grande, Nuevo Le6n; U.S.N.M. No. 3009; Lieutenant 

Couch collector). 
Cnemidophorus perplexus {nee Baird and Girard), Van Denburgh, Occ Pap. 

California Acad. Sci., No. 10, 1922, pp. 495-497.- — Schmidt and Smith, 

Publ. Field Mus. Nat. Hist., zool. ser., vol. 29, 1944, pp. 86-87. — Smith, 

Handbook of lizards, 1946, pp. 412-414, pi. 15. 
Cnemidophorus arizonae Van Denburgh, Proc. California Acad. Sci., ser. 2, 

vol. 6, 1896, p. 344 (Stanford Univ. No. 2631; W. W. Price, collector; Fair- 
bank, Cochise County, Ariz.). 
Cnemidophorus gularis velox Springer, Copeia, No. 169, 1928, p. 102 (Butler 

Univ. No. 848; Pueblo Bonito, San Juan County, N. Mex., by present 

restriction). 

Type.— V.S.N. M. No. 3032; Lieutenant Couch collector. 

Type locality. — Pesqueria Grande (= Garcia), Nuevo Le6n. 

Range. — Western Texas to extreme southeastern Arizona, south- 
ward into Chihuahua, Coahuila, and Nuevo Le6n. Recorded in 
Mexico only from the states cited. 



CHECKLIST OF REPTILES OF MEXICO 185 

CNEMIDOPHORUS BURTI Taylor 

Cnemidophorus burti Taylor, Univ. Kansas Sci. Bull., vol. 24, 1938, pp. 485-487, 
pi. 42, fig. 2. 

Type.—ERT-RMS No. 13117; E. H. Taylor collector. 
Type locality. — La Posa, 10 miles northwest of Guaymas, SoDora. 
Range. — Southwestern Sonora. Known only from the vicinity 
of Guaymas. 

CNEMIDOPHORUS SEXLINEATUS (Linnaeus) 

Lacerta sexlineaia Linnaeus, Systema naturae, ed. 12, vol. 3, 1766, p. 364. 
Cnemidophorus sezlineatus, Dum^iril and Bibron, Erp^tologie g^n^rale, vol. 5, 

1839, p. 131.— Taylor, Univ. Kansas Sci. Bull., vol. 24, 1939, pp. 520-522.— 

Smith, Handbook of lizards, 1946, pp. 415-418, figs. 13, 122, pi. 116. 
Cnemidophorus sexlineatus sexlineatus, Cope, Ann. Rep. U. S. Nat. Mus., 1898 

(1900), pp. 598, 603-605.— Burt, U. S. Nat. Mus., Bull. 154, 1931, pp. 7(>-97, 

fig. 20. 
1 Cnemidophorus per plexus Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 

1852, p. 128 (U. S. N. M. No. 3060); Wm. Gambell collector; type locality 

unknown). 

Type. — Not kno^vn. 

Type locality. — "Carolina," here restricted to Charleston, S. C. 

Range. — Eastern United States from Wisconsin, southeastern 
Wyoming, and Maryland southward to the Gulf as far west as, per- 
haps, the mouth of the Rio Grande. No definitely reliable records 
are known for Mexico, but we are informed by Bryce Brown that the 
species occurs near Brownsville, Tex., and may be expected across the 
river in Tamaulipas. Records available from Matamoros, Tamauh- 
pas, may or may not belong here. 

CNEMIDOPHORUS GADOVI Burger 

Cnemidophorus gadovii Burger, Chicago Acad. Sci. Nat. Hist. Misc., No. 65. 
1950, p. 2. 

Type.— V.S.N. M. No. 40042. 

Type locality. — HermosUlo, Sonora. 

Range. — Known only from the type locality. 

CNEMIDOPHORUS LABIALIS Stejneger 

Cnemidophorus labialis Stejneger, Proc. U. S. Nat. Mus., vol. 12, 1890, pp. 
643-644.— Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 610-612, fig. 
122.— Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 141-146.— Tevis, Copeia, 
1944, pp. 16-17. 

r^pe.— U.S.N.M. No. 15596; L. Belding collector. 
Type locality. — "Cerros Island," Pacific coast, Baja California. 
Range. — Northwestern Baja California between San QuLntin and 
Punta Eugenia, Cedros Island. 



186 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

HYPERYTHRUS GROUP 

Species. — One, represented by six forms as at present recognized. 
Range. — Extreme southwest California south throughout the penin- 
sula of Baja California and on adjacent islands. 

CNEMIDOPHORUS HYPERYTHRUS BELDINGI (Stejneger) 

Verticaria beldingi Stejneger, Proc. U. S. Nat. Mus., vol. 17, 1894, p. 17. 
Verticaria hyperythra beldingi, Van Denburgh, Occ. Pap. California Acad. Sci., 

No. 10, 1922, pp. 560-563, pi. 55. 
Cnemidophorus hyperythrus beldingi, Grinnell and Camp, Univ. California Publ. 

Zool., vol. 17, 1917, p. 175. 

Type.— U.S. IS^.M. No. 11980; L. Belding collector. 
Type locality. — "Cerros" Island, Baja California. 
Range. — Extreme southwestern California southward on Pacific 
slopes (not Gulf) to Cedros Island. 

CNEMIDOPHORUS HYPERYTHRUS CAERULEUS (Dickerson) 

Verticaria caerulea Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 472. 
Cnemidophorus hyperythrus caeruleus, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
240-242. 

Type. — Amer. Mus. Nat. Hist. No. 5517; C. H. Townsend collector. 
Type locality. — Carmen Island, Baja California. 
Range. — Known only from the type locality. 

CNEMIDOPHORUS HYPERYTHRUS DANHEIMAE Burt 

Verticaria sericea Van Denburgh, Proc. California Acad. Sci., ser. 2, vol. 5, 
1895, pp. 132-133, pi. 12. 

Cnemidophorus hyperythrus danheimae Burt, Proc. Biol. Soc. Washington, vol. 42, 
1929, p. 154 (new name for Verticaria sericea Van Denburgh, 1895, pre- 
occupied by Cnemidophorus gularis sericeus Cope, 1892); U. S. Nat. Mus. 
Bull. 154, 1931, pp. 244-246. 

Type. — California Acad. Sci. No. 435; Walter E. Bryant collector. 
Type locality. — San Jose Island, Gulf of California, Baja California. 
Range. — Known only from the type locality. 

CNEMIDOPHORUS HYPERYTHRUS PICTUS (Van Denburgh and Slevin) 

Verticaria picta Van Denburgh and Slevin. Proc. California Acad. Sci., ser. 4, 

vol. 11, 1921, p. 98. 
Cnemidophorus hyperythrus pictus, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 

242-244. 

Type. — California Acad. Sci. No. 49155; Joseph R. Slevin collector. 
Type locality. — Monserrate Island, Baja California. 
Range. — Known only from the type locality. 



CHECKLIST OF REPTILES OF MEXICO 187 

CNEMIDOPHORUS HYPERYTHRUS SCHMIDTI (Van Denbnrgh and Slevin) 

Verticaria hyperythra schmidt Van Denburgh and Slevin, Proc. California 
Acad. Sci., ser. 4, vol. 11, 1921, p. 397. — Van Denburgh, Occ. Pap. California 
Acad. Sci., No. 10, 1922, pp. 563-566. 

Cnemidophorus hyperythrus schmidti, Linsdale, Univ. California Publ. ZooL, 
vol. 38, 1932, p. 373. 

Tyjpe. — California Acad. Sci. No. 50512; Joseph R. Slevin collector. 

Type locality. — San Marcos Island, Gulf of California, Baja Cali- 
fornia. 

Range. — The central fifth of the peninsula of Baja California from 
about lat. 28° south to 25° 30' N. (the Vizcaino Desert south to 
Comondu), and adjacent islands. 

CNEMIDOPHORUS HYPERYTHRUS HYPERYTHRUS Cope 

Cnemidophorus hyperythrus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 103. 
Verticaria hyperythra hyperythra, Van Denburgh, Occ. Pap. California Acad. 

Sci., No. 10, 1922, pp. 556-559. 
Cnemidophorus hyperythrus hyperythrus, Burt, Proc. Biol. Soc. Washington, 

vol. 42, 1929, p. 154 (part); U. S. Nat. Mus., Bull. 154, 1931, pp. 226-240 

(part). — Linsdale, Univ. California Publ. Zool., vol. 38, 1932, pp. 372-373. 
Verticaria espiritensis Van Denburgh and Slevin, Proc. California Acad. Sci., 

ser. 4, vol. 11, 1921, p. 397 (Calif. Acad. Sci. No. 50511; Espfritu Santo 

Island, Baja California; J. R. Slevin collector). — Van Denburgh, op. cit., 

pp. 566-568. 
Verticaria franciscensis Van Denburgh and Slevin, loc. cit. (Calif. Acad. Sci. 

No. 50513; San Francisco Island, Baja California; J. R. Slevin collector). — 

Van Denburgh, op. cit., pp. 568-570. 

Type.— U.S.N.M. No. 5299; John Xantus collector. 

Type locality. — Cape San Lucas, Baja California. 

Range. — The southern third of the peninsula of Baja California 
and its adjacent islands on both the Gulf and Pacific sides, south 
from about lat. 25°30'. 

TESSELATUS GROUP 

Species. — Six, one of which is represented by eleven subspecies, as 
now recognized. 

Range. — Western United States from Idaho and Oregon south 
through Baja California and western Texas to southern Coahuila, 
Chihuahua, and Sonora. 

CNEMIDOPHORUS BACATUS Van Denburgh and Slevin 

Cnemidophorus bacatus Van Denburgh and Slevin, Proc. California Acad. Sci., 
ser. 4, vol. 11, 1921, p. 97. — Van Denburgh, Occ. Pap. California Acad. Sci., 
No. 10, 1922, pp. 544-546.— Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
211-213. 

Type. — California Acad. Sci. No. 49152; Joseph R. Slevin collector. 
Type locality. — San Pedro Nolasco Island, Sonora. 
Range. — Known only from the type locality. 

861316—50 13 



188 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

CNEMIDOPHORUS CATALINENSIS Van Denburgh and Slevin 

Cnemidophorus catalinensis Van Denburgh and Slevin, Proc. California Acad. 
Sci., ser. 4, vol. 11, p. 396. — Van Denburgh, Occ. Pap. California Acad. Sci., 
No. 10, 1922, pp. 542-544.— Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
213-216, fig. 29. 

TVi?^.— California Acad. Sci. No. 50507; Joseph K. Slevin collector. 
Type locality. — Santa Catalina Island, Baja California. 
Range. — Known only from the type locality. 

CNEMIDOPHORUS CERALBENSIS (Van Denburgh and Slevin) 

Verticaria ceralbensis Van Denburgh and Slevin, Proc. California Acad. Sci. 

ser, 4, vol. 11, 1921, p. 396. — Van Denburgh, Occ. Pap. California Acad* 

Sci., No. 10, 1922, pp. 554-556. 
Cnemidophorus ceralbensis, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 216-218. 

Ty^^e. —California Acad. Sci. No. 50510; Joseph R. Slevin collector. 
Type locality. — Ceralbo Island, Baja California. 
Range. — Known only from the type locality. 

CNEMIDOPHORUS TESSELATUS (Say) 

Ameiva tesselata Sat, in Long's Expedition to the Rockj' Mountains, vol. 2, 1823, 
p. 50. 

Cnemidophorus tesselatus, Smith and Burger, Bull. Chicago Acad. Sci., vol. 8, 
1949, p. 282. 

Cnemidophorus grahamii Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 
1852, p. 128 (U.S.N. M. No. 3036, 2 cotypes; Colonel Graham collector; 
between San Antonio and El Paso, Texas, here restricted to Fort Davis). — 
Strecker, Baylor Univ. Bull. No. 13, 1910, pp. 8-13, pi. 1.— Smith, Hand- 
book of lizards, 1946, pp. 419-421, pi. 117. 

Type. — Lost. 

Type locality. — Arkansas River, near Castle Rock Creek, Colo. 
(= Beaver Creek, Fremont County). 

Range. — Western Texas, eastern New Mexico, and probably southern 
Colorado, southward into adjacent northern Chihuahua and Coahuila. 
No reliable Mexican records are known to us ; Yarrow (U. S. Nat. Mus. 
Bull. 24, 1883, p. 43) records it from ''between Panos and St. Luis, 
Mexico," and Cope (U. S. Nat. Mus. Bull. 32, 1887, p. 45) cites it 
from "between Mexico City and Chihuahua," but neither record 
definitely refers to the species as now understood, and even if they did, 
the locality of collection is very uncertain. 

CNEMIDOPHORUS MAXIMUS Cope 

Cnemidophorus maximus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 104; 
Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 570-571, fig. 104.— Van Den- 
burgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 506-508.— Burt, 
U. S. Nat. Mus. Bull. 154, 1931, pp. 218-221. 

T2/;)e.— U.S.N.M. No. 5297; John Xantus collector. 



CHECKLIST OF REPTILES OF MEXICO 189 

Type locality. — Cape San Lucas, Baja California. 
Range. — Southern Baja California, southward from La Paz Bay, or 
perhaps Magdalena Bay. 

CNEMIDOPHORUS TIGRIS TIGRIS Baird and Girard 

Cnemidophorus tigris Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 

1852, p. 69. 
Cnemidophorus tigris tigris, Camp, Univ. California Publ. Zool., vol. 17, 1916, 

pp. 71-72. — Smith and Burger, Bull. Chicago Acad. Sci., vol. 8, 1949, 

p. 282. 
Cnemidophorus tessellatus tessellatus, Cope (nee Say), U. S. Nat. Mus. Bull. 1, 

1875, p. 46. — Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 

1922, pp. 508-516, pi. 23.— Burt, U. S. Nat. Mus. Bull. 154, 1922, pp. 

146-199 (part). — Smith, Handbook of hzards, 1946, pp. 421-424, pi. 118. 

Type. — U.S.N.M. No. 4103; Capt. Howard Stansbury collector. 

Type locality. — "Valley of the Great Salt Lake," Utah, restricted to 
Salt Lake City, Utah. 

Range. — Southern Idaho and southeastern Oregon south through 
western Utah and southeastern California to extreme northeastern 
Baja California, and extreme northwestern Sonora. Recorded in 
Mexico only from Baja California and Sonora (between Sonoyta and 
Puerto Penasco). 

CNEMIDOPHORUS TIGRIS GRACILIS Baird and Girard 

Cnemidophorus gracilis Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, 
■ 1852, p. 69. 

Cnemidophorus tigris gracilis, Burger, Chicago Acad. Sci. Nat. Hist. Misc., No. 
65, 1950, p. 6. 

Cnemidophorus jnelanostethus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, 
p. 104 (U.S.N.M. No. 3067, 12 cotypes; H. B. Mollhausen collector; "region 
of the Colorado, California," here restricted to Yuma, Ariz.). 

2V2?e.— U.S.N.M. No. 3034; J. L. LeConte collector. 

Type locality. — "Desert of Colorado," restricted to Yuma, Ariz. 

Range. — Extreme eastern California in Colorado River Valley, 
southern Arizona south of the plateau, extreme southwestern New 
Mexico, northern Sonora. Recorded in Mexico only from the state 
of Sonora. 

CNEMIDOPHORUS TIGRIS AETHIOPS Cope 

Cnemidophorus tessellatus aethiops Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 

p. 582. — Smith, Handbook of hzards, 1946, pp. 424-426, pi. 119 (part). 
Cnemidophorus tigris aethiops. Smith and Burger, Bull. Chicago Acad. Sci., 

vol. 8, 1949, p. 282. 
Cnemidophorus disparilis Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 

p. 473 (U.S.N.M. No. 64445; C. H. Townsend collector; Tibur6n Island, 

Sonora) . 
Cnemidophorus estebanensis Dickerson, op. cit., p. 474 (U.S.N.M. No. 64446; 

C. H. Townsend collector; San Esteban Island, Sonora). 
Cnemidophorus punctilineatus Dickerson, op. cit., p. 475 (U.S.N.M. No. 64447; 

C. H. Townsend collector; Tibur6n Island, Sonora). 



190 BULLETrN" 199, tJNITED STATES NATIONAL MUSEUM 

Cnemidophorus melanostethus, Van Denburgh, Occ. Pap. California Acad. 
Sci., No. 10, 1922, pp. 529-533 (part). 

Type. — U.S.N.M. Nos, 64240-5, co types; Jenkins and Evermann 
collectors. 

Type locality. — Hermosillo, Sonora. 

Range. — Southern Sonora and adjacent islands (Tiburdn, San 
Esteban). 

CNIMIDOPHORUS TIGRIS MARMORATUS Baird and Girard 

Cnemidophorus marmoratus Baird and Girard, Proc. Acad. Nat. Sci. Philadel- 
phia, 1852, p. 128. 

Cnemidophorus tigris marmoratus, Burger, Chicago Acad. Sci. Nat. Hist. Misc., 
No. 65, 1950, p. 7. 

Cnemidophorus variolosus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1863, p. 39 
(U.S.N.M. No. 3066; Lieutenant Couch collector; Parras, Coahuila). 

TVpc— U.S.N.M. No. 3024; two cotypes; J. H. Clark collector. 

Type locality. — "Between San Antonio and El Paso," restricted to 
El Paso, Tex. 

Range. — Southwestern New Mexico to southwestern Coahuila. 
Recorded in Mexico only from Chihuahua, Durango (5 km. west of 
Torre6n), and Coahuila. 

CNEMIDOPHORUS TIGRIS CANUS Van Denburgh and Slevln 

Cnemidophorus canus Van Denburgh and Slevin, Proc. California Acad. Sci., 

ser. 4, vol. 11, No. 6, 1921, p. 97. — Van Denburgh, Occ. Pap. California Acad. 

Sci., vol. 10, 1922, pp. 540-542. 
Cnemidophorus tessellatus canus, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 

208-211. 
Cnemidophorus tigris canus, Smith and Burger, Bull. Chicago Acad. Sci., vol. 8, 

1949, p. 282. 

Type. — California Acad. Sci. No. 49153; Joseph R. Slevin collector. 

Type locality. — Sal Si Puedes Island, Gulf of California, Baja 
California. 

Range. — The type locality, and North and South San Lorenzo 
Islands (on the latter, intergrades with C. t. martyris). 

CNEMIDOPHORUS TIGRIS CELERIPES Dickerson 

Cnemidophorus celeripes Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, 

p. 472. — Van Denburgh, Occ. Pap. California Acad. Sci., vol. 10, 1922, pp. 

549-551. 
Cnemidophorus tessellatus celeripes, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 

202-205. 
Cnemidophorus tigris celeripes, Smith and Burger, Bull. Chicago Acad. Sci., 

vol. 8, 1949, p. 282. 

Type. — Amer. Mus. Nat. Hist. No. 5514; C. H. Townsend collector. 
Type locality. — San Jos6 Island, Baja California. 
Range. — Known only from the type locality. 



CHECKLIST OF REPTILES OF MEXICO 191 

CNEMIDOPHORUS TIGRIS MARTYRIS Stejncger 

Cnemidophorus martyris Stejneger, Proc. U. S. Nat. Mus., vol. 14, 1891, p. 407. — 
Van Denburgh, Occ. Pap. California Acad. Sci., vol. 10, 1922, pp. 538-540, 

Cnemidophorus tessellatus martyris, Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 
205-208. 

Cnemidophorus tigris martyris, Smith and Burger, Bull. Chicago Acad. Sci.,. 
vol. 8, 1949, p. 282. 

Type.— U.S.N. M. No. 15620; E. Palmer collector. 
Type locality. — San Pedro Martir Island, Gulf of California, Sonora. 
Range. — The type locality (intergrades with C. t. canus on South 
San Lorenzo Island). 

CNEMIDOPHORUS TIGRIS RUBIDUS Cope 

Cnemidophorus tessellatus ruhidus Cope, Trans. Amer. Philos. Soc, ser. 2, vol. 17, 

1892, p. 36, pi. 12, fig. 1.— Burt, U. S. Nat. Mus. Bull. 154, 1931, pp. 199-202. 
Cnemidophorus ruhidus, Van Denburgh, Occ. Pap. California Acad. Sci. vol. 10, 

1922, pp. 546-549. 
Cnemidophorus tigris ruhidus, Smith and Burger, Bull, Chicago Acad. Sci., vol, 

8, 1949, p. 282. 
Cnemidophorus vandenhurghi Dickersoj«t, Bull. Amer. Mus. Nat. Hist., vol. 41, 

1919, p. 477 (U.S.N.M. No. 64449; C. H. Townsend collector; Carmen 

Island, Baja California). 

Type.— U.S.N.M. Nos. 15149-15155, cotypes; U, S, Fish Commis- 
sion collector. 

Type locality. — Santa Margarita Island, Baja California. 

Range. — The southern third of Baja California, south of San 
Marcos Island, including adjacent islands in the Gulf and off the 
western edge of the peninsula (notably Carmen, Santa Margarita, 
and Magdalena Islands, Baja California). 

CNEMIDOPHORUS TIGRIS MULTISCUTATUS Cope 

Cnemidophorus tessellatus multiscutatus Cope, Trans, Amer, Philos, Soc, vol. 17, 
1892, p. 38. 

Cnemidophorus stejnegeri Van Denburgh, Proc. California Acad. Sci., ser. 2, 
vol. 4, 1894, p. 300 (Stanford Univ. Mus. No. 1861; Ensenada, Baja Cali- 
fornia; J. M. Stovell collector). 

Cnemidophorus tessellatus stejnegeri, Stejneger and Barbour, Check list North 
American Amphibia and Reptilia, ed. 1, 1917, p. 68. — Van Denburgh, 
Occ. Pap. California Acad. Sci., vol. 10, 1922, pp. 519-523, pi. 54.— Smith, 
Handbook of lizards, 1946, pp. 426-428, pi, 120, 

Cnemidophorus bartolomas Dickerson, Bull. Amer. Mus. Nat. Hist., vol. 41, 
1919, p. 476 (U.S.N.M. No. 64448; C, H, Townsend collector, San Barto- 
lom^ Bay, Baja Calfornia), 

Cnemidophorus dickersonae Van Denburgh and Slevin, Proc, California Acad, 
Sci,, ser, 4, vol. 11, 1921, p, 97 (California Acad, Sci. No, 49154; J, R, Slevin 
collector; Isla Partida near Angel de la Guarda Island, Baja California). 

Type.— V.S.N. M. Nos. 15160-3; U, S, Fish Commission donor. 
Type locality. — Cedros Island, Baja California. 



192 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Range. — Southern California west of the Sierra Nevada, south to 
central Baja California, exclusive of the extreme northeastern corner; 
islands in the Gulf of California near the center of the Peninsula 
(Isla Partida, Angel de la Guarda). 

Genus GYMNOPHTHALMUS Merrem 

Gymnophthalmus Merrem, Tentamen systematis Amphibiorum, 1820, p. 74. — 
Stuart, Occ. Pap, Mus. Zool. Univ. Michigan No. 409, 1939, pp. 1-10, pi. 1. 

Epaphelus Cope, Journ. Acad. Nat. Sci. Philadelphia, 1875, p. 115 (type, Epa- 
phelus sumichrasti Cope;. 

Blepharactisis Hallowell, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 484 
(type, Blepharactisis speciosa Hallowell). 

Genotype. — Lacerta guadrilineata Linnaeus [= Gymnophthalmus lin- 
eata (Linnaeus)]. 

Range. — Central Argentina north through South and Central 
America. In Mexico, Oaxaca and possibly Chiapas. 

Species. — Seven. Only one occurs in Mexico. 

GYMNOPHTHALMUS SUMICHRASTI (Cope) 

Epaphelus sumichrasti Cope, Journ. Acad. Nat. Sci. Philadelphia, 1876, p. 115. 

Gymnophthalmus sumichrasti,"^^ BocouRT, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 7, 1881, pp. 471-472, pi. 22H, fig. 2, 2a-2h.— 
BotJLENGER, Catalogue of the lizards in the British Museum, vol. 2, 1885, 
pp. 428-429.— Stuart, Occ. Pap. Mus. Zool. Univ. Michigan, No. 409, 1939, 
pp. 5-6. 

TVi^e.— U.S.N. M. No. 30245-46; Frangois E. Sumichrast collector. 

Type locality. — "Western part of the State of Tehuantepec" ("Near 
Ventose 'B&j" fide Dunn in Stuart, loc. cit.). 

Range. — Southern Oaxaca near the Isthmus of Tehuantepec east- 
ward on Pacific slopes possibly to Honduras. Recorded in Mexico 
only from Oaxaca: Tehuantepec, Bahia Ventosa, Mount Guengola, 
Cacoprieto. A specimen, presumably of this species, was seen near 
San Ricardo, Chiapas, by the junior author. 

Family HELODERMIDAE Gray 

Helodermidae Gray, Proc. Zool. Soc. London, 1837, p. 132. 

Genera. — A single genus, Heloderma Wiegmann, is known. 
Range. — Southern Utah to the Isthmus of Tehuantepec, on Pacific 
slopes. 

Genus HELODERMA Wiegmann 

Trachyderma Wiegmann (nee Latreille), Isis von Oken, vol. 22, 1829, p. 421 (type, 

Trachyderma horridum Wiegmann). 
Heloderma Weigmann, Isis von Oken, vol. 22, 1829, p. 624. 

'» Gymnodactylus sumichrasti Qadow, Proc. Zool. Soc. London, 1905, p. 211, from the Isthmus of Tehuan- 
tepec, appears to be a lapsus for this species. 



CHECKLIST OF REPTILES OF MEXICO 193 

Genotype. — Trachyderma horridum Wiegmann, 

Range. — Southern Utah to the Isthmus of Tehuantepec. 

Species. — Two. 

KEY TO SPECIES OF HELODERMA 

1. Tail equal to or longer than body; head and neck about half of body length; 
predominating color black ; pterygopalatine teeth present. -horridum (p. 193) 
Tail about two-thirds of body length; head and neck about one-third of body 
length; colors pink, yellowish, and black; none or rarely 1 or 2 pterygo- 
palatine teeth suspectum (p. 193) 

HELODERMA SUSPECTUM Cope 

Heloderma suspectum Cope, Proc. Acad. Nat. Sci. Philadelphia, 1869, pp. 5-6; 
Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 47&-483, fig. 87.— Smith, 
Handbook of lizards, 1946, pp. 472-475, pi. 134. 

Type. — U.S.N.M. No. 2971, three cotypes; Major Emory collector. 

Type locality. — Sierra de la Union, "Sonora"(= Arizona). 

Range. — Sonora as far south as Guaymas; in the United States, 
Arizona, southern Nevada, and southwestern Utah. Recorded from 
Sonora: Guadelupe Canon, San Pedro Bay, La Posa about 10 miles 
northwest of Guaymas, Monument 88, San Bernardino, Niggerhead 
Mountain, 10 miles south of Noria, etc. 

HELODERMA HORRIDUM (Wiegmann) 

Trachyderma horridum Wiegmann, Isis von Oken, vol. 22, 1829, p. 421. 
Heloderma horridum, Wiegmann, Isis von Oken, vol.22, 1829, p. 628; Herpeto- 

logia Mexicana, pt. 1, 1834, pp. 24-25, pi. 1. — Gtjnther, Biologia Centrali- 

Americana, Reptilia and Batrachia, 1885, pp. 43-44, pi. 26. — Smith, Univ. 

Kansas Sci. Bull., vol. 22, 1935, pp. 145-146. 
Heloderma Hernandesvi Wiegmann, Herpetologia Mexicana, pt. 1, 1834, p. 25 

(for a theoretical form). 

Type. — Zool. Mus. Berlin; Ferdinand Deppe collector. 

Type locality. — "Vivit in ferventibus terrae Mexicanae regionibus," 
here restricted to Huajintlan, Guerrero. 

Range.- — Coastal areas from Oaxaca to Sonora, extending up 
the Rio Balsas Valley to Morelos.^^ Recorded from Sonora: 
Alamos, Guirocoba; Sinaloa: Presidio near Mazatlan, San Bias; 
Jalisco: Autlan, Barranca de San Crist6bal; Morelos: Canon del 
Lobo near Cuernavaca; Oaxaca: Tapanatepec, Salina Cruz, Jimiltepec, 
Juchitan, Tehuantepec, Quiengola and Mixtequilla Mountains; 
Michoacdn: Apatzingan, Parecuaro, Oropeo, Etucuaro, Puruaran; 
Colima: Paso del Rio, Colima; C^errgro; Huajintlan; CAiapos ; Rancho 
San Bartolo. 



»« A sight record for Zongolica, Veracruz (Martin del Campo, Anal. Inst. Biologfa, vol. 6, 1935, p. 297) 
is unacceptable and may be referable to Xenosawus. Likevrise unacceptable are records for Guanajuato 
and Yucatto. We are unable to find authoritative records for Nayarit, although Martin del Campo 
(loc. cit.) cites the state. 



194 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Family ANGUIDAE Cope 

Angu[i]dae Gray, Ann. Philos., vol. 26, 1825, p. 201. 

Anguidae Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 228. 

Genera. — Eleven, of which nine are restricted to the Americas, 
one (Anguis) is restricted to the Old World, and one (Ophisaurus) 
occurs in both hemispheres. Five occur in Mexico.^^ 

Range. — Southern Canada to Argentina, West Indies, Europe, 
northern Africa, southern Asia. 

KEY TO MEXICAN GENERA OF ANGUIDAE 

1. No lateral fold Celestus (p. 194) 

A lateral fold present 2 

2. Lateral fold weakly developed, containing few or no granules; sides of neck 

with coarse granules; head much widened and depressed; pterygoid teeth 
absent; minimum scales in a transverse row across neck 4-6; arboreal 

species Abronia (p. 196) 

Lateral fold moderately to well developed, containing a moderate or large 
granular area; sides of neck with fine granules; head thicker, not depressed; 
pterygoid teeth various; minimum scales in a transverse row across neck 
no less, often more numerous ; terrestrial species 3 

3. No anterior internasals, but instead a pair of enlarged supranasals, which 

much resemble internasals; no unpaired median internasals; nasal in contact 

with rostral; pterygoid teeth well developed Elgaria (p. 205) 

Anterior internasals, posterior internasals and supranasals '^ all present; 
nasal seldom in contact with rostral; pterygoid teeth various 4 

4. One or more median, unpaired postrostrals; frontals in contact with maxilla; 

pterygoid teeth well developed ; tail extremely elongate _ Qerrhonotus (p. 202) 

No postrostrals; frontals separated from maxilla; pterygoid teeth absent 

or vestigial; tail normal Barisia (p. 198) 

Genus CELESTUS Gray 

Celestus Gray, Ann. Mag. Nat. Hist., vol. 2, 1839, p. 288.»9 

Siderolamprus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 368 (type, S. 
enneagrammus Cope). 

Genotype. — Celestus striatus Gray. 

Range. — West Indies, Central America, and southern Mexico. 

Species. — About 18, two (possibly three) of which occur in Mexico. 



" Ophisaurus ventralis, cited from Jalapa, Veracraz, as early as 1884 (Yarrow, U. S. Nat. Mus. Bull. 24, 
1884, p. 46) , and retained in accounts of the species even up to the present time, has never been authoritatively 
recorded from Mexico and probably does not occur there. It almost certainly does not occur at Jalapa. 
It is possible that the record arose from a simple misunderstanding of Peale and Green's Scinais ventralis 
(= Qerrhonotus liocephalus) which occurs in the same general area. Regardless of the nature of the error, 
we emphatically believe the record is erroneous. 

•' Supranasals ab.sent in Oerrhonotus liocephalus austrinus. which has a postrostral. 

«» Dumi (Notulao Nat., No. 4, 1939, p. 3) expresses the opinion that forms here referred to Celestus, char- 
acterized by absence of sheaths on the claws, are congeneric with Diploglossut (Wiegmann, Herpetologia 
Mexicana, 1834, p. 36; type "Sc.fasciatus Qrny" = Diploglossus fasciatus) , which possesses sheaths. 



CHECKLIST OF REPTILES OF MEXICO 195 

KEY TO MEXICAN SPECIES OF CELESTUS 

1. Median prefrontal in contact with 2 supraoculars, wider than long; no lateral 

prefrontals (fused with median loreal); 2 dorsolateral light stripes in adults 

separated by 4 and 2 half scale rows atitlanensis (p. 195) 

Median prefrontal in contact only with anterior supraocular, as long as wide; 
small lateral prefrontals usually present; adults not with 2 dorsolateral light 
stripes 2 

2. Snout normal, not especially elongate or flattened; lamellae under fourth toe, 

15-18; scales in 33-35 rows; first labial reaching to middle of naris; distance 
between anterior and posterior loreals less than the length of either; no ver- 
tical light bars on sides in young or adults enneagrammus (p. 195) 

Snout flattened, elongate; lamellae under fourth toe, 23-26; scales in 31-33 
rows; first labial reaching to anterior border of naris; distance between an- 
terior and posterior loreals as great or greater than length of either; vertical 
light bars present on sides in young and adults rozellae (p. 195) 

CELESTUS ROZELLAE Smith 

Diploglossus steindachneri, Guntheb {nee Cope), Biologia Central!- Americana, 

Reptilia and Batrachia, 1885, p. 34, pi. 22, fig. A. 
Celestus rozellae Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 372-374. 

TVpe.— U.S.N.M. No. 113526; Rozella Smith collector. 

Type locality. — Palenque, Chiapas. 

Range. — Atlantic slopes, presumably from the Isthmus of Tehuan- 
tepec to British Honduras. Recorded in Mexico only from the type 
locality. 

CELESTUS ATITLANENSIS Smith, new species i 

Diploglossus (Celestus) steindachneri, Bocourt (nee Cope), Mission scientifique 
au Mexique . . . , Etudes sur les reptiles, livr. 6, 1879, pp. 383-384, pi. 22, 
fig. 3. 

Type. — Mus. Hist. Nat. Paris, specimen figured as above. 

Type locality. — Atitlan, Guatemala. 

Range. — Known only from the type specimen from Atitlan, Guate- 
mala, but probably occurring along Pacific slopes from Chiapas to 
Nicaragua. 

CELESTUS ENNEAGRAMMUS (Cope) 

Siderolamprus enneagrammus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1860, 
p. 368. 

Celestus enneagrammus, Cope, U. S. Nat. Mus. Bull. 32, 1887, p. 43; Ann. Rep. 
U. S. Nat. Mus., 1898 (1900), pp. 504-505.— Smith, Proc. U. S. Nat. Mus., 
vol. 92, 1942, pp. 370-372, 374. 

Diploglossus steindachneri Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 179 
(type locality, Orizaba, Veracruz; U.S.N.M. No. 6342). — Boulenger, Cata- 
logue of the lizards in the British Museum, vol. 2, 1885, p. 293. 

* Diagnosis. — Allied to C. enneagrammus and C. rozellae, but with median prefrontal in contact with two 
(instead of one) supraoculars, no lateral prefrontals (presumably fused to median loreals), a very broad 
median prefrontal (broader than long), and a pattern in the single known adult (112 mm. snout to vent) 
consisting of two dorsolateral light stripes separating three broad dark stripes from each other. Similar to 
C. bisittatiis Boulenger but with three loreals instead of two, seven supralabials to a point below middle of 
eye instead of six. — H. M. S, 



196 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Diploglossus chalybaeus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 321 
(type locality, Orizaba, Mexico, 4,000-6,000 feet; U.S.N.M. No. 6603). 

Type. — Mus. Comp. Zool. No. 2848; R. Montes de Oca collector. 

Type locality. — Jalapa, Veracruz. 

Range. — Atlantic foothills of Veracruz and Oaxaca. Reported 
from Veracruz: Orizaba, Jalapa, Tequeyutepec, "one to three miles 
west of La Goya"; Oaxaca: Totontepec, "Tehuantepec," Santa 
Efigenia. 

Genus ABRONIA Gray 

Ahronia Gray, Ann. Mag. Nat. Hist., ser. 1, vol. 1, 1838, p. 389. — Tihen, Amer. 
Midi. Nat., vol. 41, 1949, pp. 587-591. 

Genotype. — Gerrhonotus deppii Wiegmann [=Abronia deppii (Wieg- 
mann).] 

Range. — Hidalgo, south to Guerrero and Chiapas. In Central 
America, Guatemala. 

Species. — Nine species, 10 forms; 6 species and 7 forms in Mexico. 

KEY TO MEXICAN FORMS OF ABRONIA 

1. Postmental unpaired 2 

Postmental paired 3 

2. Scale rows 30-33; parietal separated from supraoculars or very narrowly in 

contact; dorsal markings usually indistinct, those on neck, if present, com- 
bining to form a large single mark ochoterenai (p. 197) 

Scale rows 39; parietals broadly in contact with the supraocular; markings on 
back distinct, those on neck discrete, not forming a large single mark. 

matudai (p. 196) 

3. Suboculars missing or reduced to minute scales; one anterior temporal bordering 

orbit deppii (p. 197) 

Suboculars present, well developed; 2 anterior temporals bordering orbit. _ 4 

4. Area of granular scales on sides of neck very narrow; no granular zone in 

lateral fold oaxacae (p. 197) 

Area of granular scales covering entire sides of neck; a granular zone in lateral 
fold 5 

5. Prominent light markings along the sides; dorsal osteoderms absent over 

posterior part of body; dorsal scales about 31-34 (average about 33); mini- 
mum number of scales in a single nuchal transverse row usually 6. 

taeniata taeniata (p. 197) 

No light markings on sides of body; dorsal osteoderms present over the whole 

body 6 

6. Dorsal scale rows 25 to 29; minimum number of scales in a transverse nuchal 

row, 4 or 5 (75 percent) ; chin and lower labials white with occasionally a few 

scattered dark blotches taeniata graminea (p. 198) 

Dorsal scale rows 30-31; minimum number of scales in a transverse nuchal row, 
6; infralabials darker than chin and as dark as granular area of neck, with 
indistinct lighter bands fuscolabialis (p. 198) 

ABRONIA MATUDAI (Hartweg and Tihen) 

Gerrhonotus matudae Hartweg and Tihen, Occ. Pap. Mus. Zool. Univ. Michigan, 

No. 497, 1946, pp. 3-5. 
Abronia matudai, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 591. 



CHECKLIST OF REPTILES OF MEXICO 197 

Type.— Umv. Michigan Mus. Zool. No. 88331; Eizi Matuda 
collector. 

Type locality. — Volcan de Tacan4, Chiapas, 2,000 meters. 
Range. — Known only from the type locality. 

ABRONIA OCHOTERENAI (Martin del Campo) 

Gerrhonotus vasconcelosii ochoterenai MartIn del Campo, Anal. Inst. Biol. Mexico, 

vol. 10, 1939, pp. 357-359, fig. 3 (cotypes). 
[Gerrhonotus] ochoterenai, Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 368. 
Abronia ochoterenai, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 591. 
^Gerrhonotus fimbriatus,^ MaetIn del Campo, Anal. Inst. Biol. Mexico, vol. 10, 

1939, p. 359. 

Type. — Instituto de Biologia, Mexico, cotypes, male and female; 
Mario del Toro collector. 

Type locality. —Santa Rosa, Comitan, Chiapas. 
Range. — Known only from the type locality. 

abronia DEPPn (Wiegmann) 

Gerrhonotus deppii Wiegmann, Isis von Oken, vol. 21, 1828, p. 379; Herpetologia 

Mexicana, pt. 1, 1834, pp. 31-32, pi. 9, figs. 3, 4. 
Abronia deppii, Gray, Ann. Nat. Hist., vol. 1, 1838, p. 389. — Tihen, Amer. 

Midi. Nat., vol. 41, 1949, p. 590. 
Gerrhonotus (Abronia) Deppei, Bocourt, Mission Scientifique au Mexique . . . , 

Etudes sur les reptiles, livr. 5, 1878, pp. 325-327, pi. 21A, figs. 3, 3a. 

Type. — Zool. Mus. Berlin; F. Deppe collector. 
Type locality. — Mexico. Here restricted to OmUteme, Guerrero. 
Range. — Sierra Madre del Sur. Known only from the vicinity of 
Omilteme, Guerrero. 

abronia OAXACAE (GUnther) 

Gerrhonotus oaxacae GtJNTHER, Biologia Centrali-Americana, Reptilia and 

Batrachia, 1885, p. 36, pi. 24, figs. A, A', A", 
Abronia oaxacae, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 591. 

Type. — Brit. Mus, Nat. Hist., 2 adult males, 1 young; A. Boucard 
collector. 

Type locality. — "Oaxaca," Mexico. 

Range. — Mountains in Oaxaca. Reported definitely only from 
"Luvina, Tehuantepec, Oaxaca." 

abronia TAENIATA TAENIATA (Wiegmann) 

Gerrhonotus taeniatus Wiegmann, Isis von Oken, vol. 21, 1828, p. 379; Herpetologia 

Mexicana, 1834, p. 32, pi. 9, fig. 1, 2. 
Abronia taeniatus, Gray, Ann. Nat. Hist., vol. 1, 1838, p. 390. — Tihen, Amer. 

Midi. Nat., vol. 41, 1949, p. 590. 
Gerrhonotus (Abronia) taeniatus, Bocourt, Mission scientifique au Mexique . . ., 

Etudes sur les reptiles, livr. 5, 1878, pp. 327-329, pi. 21A, figs. 4, 4a, and 5. 



' This arrangement has been suggested verbally by Dr. Tihen. 



198 BULLETIN 19 9, UNITED STATES NATIONAL MUSEUM 

Gerrhonotus deppii digueti Mocquard, Bull. Mus. Hist. Nat. Paris, 1905, p. 79 
(Mus. Hist. Nat. Paris; Sierra de Zacapoaxtla, Puebla; Leon Diguet, collector). 

Type. — Zool. Mus, Berlin; F. Deppe collector. 
Type locality. — Mexico. Here restricted to El Chico, Hidalgo.' 
Range. — Southern Hidalgo, central and northern Puebla. Re- 
ported from Hidalgo: El Chico; Puebla: Zacapoaxtla, Ahuacatlan. 
Intergrades of taeniata and graminea are reported from La Joya, 
Veracruz. 

ABRONIA TAENIATA GRAMINEA (Cope) 

Gerrhonotus gramineus Cope, Proc. Acad. Nat. Sci. Philadelphia, Aug. 1864, p. 

179. — GtJNTHER, Biologia Centrali-Americana, Reptilia and Batrachia, 

1885, p. 36, pi. 24, fig. B. 
Abronia taeniata graminea, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 591. 

Type. — U.S.N.M. (Field No. 50); Frangois E. Sumichrast collector. 

Type locality. — Orizaba, Mexico. 

Range. — Central Veracruz and eastern Puebla. Reported from 
Puebla: Pajaro Verde, Puente Colorado; Veracruz: Orizaba, 
Acultzingo. 

ABRONIA FUSCOLABIALIS (Tihen) 

Gerrhonotus fuscolabialis Tihen, Copeia, 1944, pp. 112-115, figs. 1-3. 
Abronia fuscolabialis, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 591. 

Type.—R. T. Moore coll. No. 400 (California Inst. Tech.). 
Type locality. — Mount Zempoaltepec, Oaxaca. 
Range. — Known only from the type locality. 

Genus BARISIA Gray 

Barisia Gray, Ann. Mag. Nat. Hist., ser. 1, vol. 1, 1838, p. 390. — Tihen, Amer. 

Midi. Nat., vol. 41, 1949, pp. 596-598, figs. 10-12; Univ. Kansas Sci. Bull., 

vol. 23, 1949, pp. 217-256, pis. 1-2, fig. 1. 
Mesaspis Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 96 (type, M. moreletii 

Bocourt) . 

Genotype. — Gerrhonotus imbricatus Wiegmann [=Barisia imbricata 
imbricata (Wiegmann)]. 

Range. — Chihuahua and Coahuila south to Chiapas; Central 
America to Panama. 

Species. — Nine, with 16 forms; 8 species, 12 forms, occur in Mexico. 

KEY TO MEXICAN SPECIES OF BARISIA 

1. Postmental unpaired; prefrontal bones in contact 2 

Postmental paired; prefrontal bones separated by frontonasal 4 

2. Eighteen to 20 longitudinal dorsal scale rows; frontonasal normally present- 3 
Fourteen longitudinal dorsal scale rows; frontonasal normally absent. 

viridiflava (p. 200) 

3. Upper postnasal separated from lower by a loreonasal contact. 

moreleti rafaeli (p. 199) 



• It is known that Deppe collected in this region. 



CHECKLIST OF REPTILES OF MEXICO 199 

Upper and lower postnasals in contact moreleti temporalis (p. 199) 

4. Superciliary series complete ; pterygoid teeth absent 5 

Superciliary series incomplete; pterygoid teeth vestigial 8 

5 . Supranasals unexpanded ; an anterior canthal present 6 

Supranasals expanded; no anterior canthal 7 

6. Scales of neck keeled; anterior loreal in contact with anterior canthal. 

gadovii gadovii (p. 200) 

Scales of neck smooth; anterior loreal separated from anterior canthal by a 

contact of posterior loreal with upper postnasal__gadovii levigata (p. 200) 

7. Frontonasal and postrostral present raodesta (p. 201) 

No frontonasal or postrostral present antauges (p. 200) 

8. One loreal; more than 33 dorsal scale rows 9 

Two loreals; fewer than 33 dorsal scale rows rudicollis (p. 202) 

9. More than 1 superciliary (usually 3) ; transverse dorsal scale rows usually 

fewer than 47 10 

A single (middle) superciliary element; transverse dorsal scale rows usually 
more than 47 levicollis (p. 202) 

10. Fewer than 16 longitudinal dorsal scale rows; contact of anterior superciliary 

with loreal usually smaller than that of first medial supraocular with 

loreal 11 

Sixteen longitudinal dorsal scale rows; contact of anterior superciliary with 
loreal as great or greater than that of first medial supraocular with loreal. 

imbricata irabricata (p. 201) 

11. Lowest primary temporal in contact with penultimate as well as with ante- 

penultimate supralabial; 39-45 transverse dorsal scale rows. 

imbricata ciliaris (p. 202) 

Lowest primary temporal not in contact with penultimate supralabial; 35-39 

transverse dorsal scale rows imbricata planifrons (p. 201) 

BARISIA MORELETI RAFAEU (Hartweg and Tihen) 

Gerrhonotus moreleti rafaeli Hartweg and Tihen, Occ. Pap. Mus. Zool. Univ. 

Michigan, No. 497, 1946, pp. 8-10. 
Barisia moreleti rafaeli, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 223-224. 

Type.— M.M^. Zool. Univ. Michigan No. 88228; Eizi Alatuda 
collector. 

Type locality. — 16 km. south of Siltepec, Chiapas, 2,300 meters 
elevation. 

Range. — High momitains of southern Chiapas. Reported from 
Cerro Paxtal, 1,500 meters; Chiquihuite, Volcan de Tacana, 2,500 
meters; Cerro Male. 

BARISIA MORELETI TEMPORALIS (Hartweg and Tihen) 

Gerrhonotus moreleti temporalis Hahtweg and Tihen, Occ. Pap. Mus. Zool. 

Univ. Michigan, No. 497, 1946, pp. 10-15. 
Barisia moreleti temporalis, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 224-225. 

Type. — Mus. Zool. Univ. Michigan No. 94910; Norman Hartweg 
collector. 

Type locality. — Eleven km. southeast of Ciudad de las Casas, 
Chiapas, 2,300 meters elevation. 

Range. — Known only from the type locality. 



200 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

BARISIA GADOVn GADOVn (Boulenger) 

Gerrhonotus gadovii Boulengkr, Ann. Mag. Nat. Hist., ser. 8, vol. 12, 1913, pp. 

564-565. 
Barisia gadovii gadovii, Tihen, Univ. Kansas Sci. Bull., vol. 33, 1949, pp. 230-231. 

Type. — Brit. Mus. Nat. Hist., two co types; Hans Gadow collector. 
Type locality. — Omilteme, Guerrero. 

Range. — Sierra Madre del Sur, in Guerrero. Known only from 
mountains west of Chilpancingo and the type locality. 

BARISIA GADOVn LEVIGATA Tihen 

Barisia gadovii levigata Tihen, Univ. Kansas Sci. Bull., vol. 33, 1949, pp. 231-233, 
pi. 2, figs. 3-5. 

Type.— U.S.N. M. No. 47212; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — "Valley of Oaxaca," Oaxaca. 

Range. — Mountains of central Oaxaca. Reported from ''moun- 
tains west of Oaxaca city." 

BARISIA VIRIDIFLAVA (Bocourt) 

Gerrhonotus viridiflavus Bocourt, Ann. Sci. Nat., vol. 17, 1873, art. 2 (unpaged). 
Barisia viridiflava, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 227-229, fig. 1. 
Gerrhonotus bocourti Peters, Monatsb. Acad. Wiss. Berlin, 1876, p. 297 (type 

locality, "Mexico," here restricted to Oaxaca, Oaxaca; Zool. Mus. Berl.). 
Gerrhonotus antauges (part), Bocourt, Mission scientifique au Mexique . . ., 

Etudes sur les reptiles, livr. 5, 1878, pp. 34&-349, pi. 21B, figs. 7, 7a. 
Gerrhonotus obscurus GtJNTHER, Biologia Centrali-Americana, Reptilia and 

Batrachia, 1895, p. 40, pi. 25, figs. E, E' (type locality, "Mexico," here 

restricted to Oaxaca, Oaxaca; Brit. Mus. Nat. Hist.). 

Type. — Mus. Hist. Nat. Paris; "un seul exemplaire qui a et4 c6d6 
par M. Boucard, comme provenant des collections de M. F. Sumi- 
chrast." 

Tjpe locality. — Mexico. Restricted by Tihen to the higlilands of 
central Oaxaca, near the city of Oaxaca. 

Range. — Mountains north of Oaxaca (city). Reported from "sum- 
mit of Cerro San Felipe north of Oaxaca" (city). 

BARISIA ANTAUGES Cope 

Barissia antaxiges Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 132. 
Barisia antauges, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 235-236. 

Type.—\J. S. N. M. No. 30221; Francois E. Sumichrast collector. 
Type locality. — Orizaba, Veracruz [=Volcan Citlaltepetl]. 
Range. — Volcan Citlaltepetl (Mount Orizaba), Veracruz; known 
only from the type locality. 



CHECKLIST OF REPTILES OF MEXICO 201 

BARISIA MODESTA (Cope) 

Pterogasterus modestus Cope, Proc. Amer. Philos. Soc, 1877, pp. 96, 97. 
Gerrhonotus modestus, Boulengek, Catalogue of the lizards in the British Museum, 

vol. 2, 1885, p. 276. 
Barisia modestus, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 234-235. 

Type. — U. S. Nat. Mus., three specimens; collector unknown. 

Type locality. — "Probably Guatemala," here restricted to Mount 
Orizaba. 

Range. — Region of Mount Orizaba (Volcan Citlaltepetl), Veracruz 
Caot known from Guatemala). 

BARISIA IMBRICATA IMBRICATA (Wiegmann) 

Gerrhonotusimbricatus Wiegmann, Isis von Oken, vol. 21, 1828, p. 379; Herpetologia 

Mexicana, 1834, pt. 1, p. 34, pi. 10, figs. 2, 5. 
Barissia imbricata, Gray, Catalogue of the lizards in the British Museum, 1845, 

p. 55. 
Gerrhonotus (Barissia) imbricatus, Bocotjrt, Mission scientifique au Mexique . , ., 

fitudes sur les reptiles, livr. 5, 1878, pi, 21 B, figs. 1, la, 2, 2a, and livr. 6, 

1879, pp. 363-365. 
Barisia imbricata imbricata, TiGEt^, Amer. Midi. Nat., vol. 41, 1949, pp. 240-244. 
Gerrhonotus lichenigerus Wagler, Descriptiones et icones amphibiorum, 1833, 

pi. 34, fig. 2 (type locality here restricted to Mexico, D. F.). — Wiegmann, 

Herpetologia Mexicana, 1834, pi. 10, figs. 2-5. 
Gerrhonotus adspersus Wiegmann, Herpetologia Mexicana, 1834, pi. 10 (type 

locality, Mexico by inference, here restricted to San Martin, Mexico; Zool. 

Mus. Berl.; F. Deppe collector). 
Gerrhonotus olivaceus Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255 

(type locality, "near San Diego;" U.S.N. M. No. 3096, two cotypes). 

Type. — Zool. Mus. Berlin, two cotypes; F. Deppe collector. 

Type locality. — Mexico, here restricted to Mexico, D. F. 

Range. — The periphery and southern part of the central Mexican 
Plateau, from Veracruz to Jalisco. Reported from numerous localities 
in the states of Jalisco, Michoac^n, Mexico, Morelos, Puebla, Oaxaca, 
Veracruz, Hidalgo, and Guanajuato, and from Distrito Federal. 

BARISIA IMBRICATA PLANIFRONS (Bocourt) 

Gerrhonotus (Barissia) planifrons Bocotjrt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 5, 1878, pi. 21C, figs. 1, la; livr. 6, 1879, pp. 
361-363 (pi. 21C was published with the name, in livr. 5, 1878; this con- 
stitutes the "type description"). 

Barisia imbricata planifrons, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 246-247, 

Type. — Mus. Hist. Nat. Paris; Adolphe Boucard collector. 
Type locality. — "Oaxaca." 

Range. — Oaxaca, mountainous areas; not known from a definite 
locality. 



202 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

BARISIA IMBRICATA CILIARIS (Smith) 

Gerrhonotus levicolUs ciliaris Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 

365-367. 
Bar isia imhricata ciliaris, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 244-245. 

2Vpe.— U.S.N.M. No. 47496; E. W. Nelson and E. A. Goldman 
collectors. 

Type locality. — Sierra Guadelupe, Coahuila. 

Range. — The northern portion of the plateau from extreme southern 
Coahuila to Guanajuato, east to San Luis Potosi and west to Sinaloa. 
Reported from Coahuila: Sierra Guadelupe; Zacatecas: Sierra Madre; 
Durango: Coyotes, Inde; San Luis Potosi: mountains near San Luis 
Potosi (city), near Xilitla, near Jesiis Maria; Guanajuato: San Felipe, 
Sierra de Santa Rosa; Hidalgo: Durango; Nuevo Leon: Pablillo, 
Galeana; Sinaloa: Escuinapa. 

BARISIA LEVICOLLIS Stejneger 

Barissia levicollis Stejneger, Proc. U. S. Nat. Mus., vol. 13, 1890, pp. 184-185. — 
Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), pp. 535-536, fig. 96. 

[Gerrhonotus] levicollis levicollis, Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 368. 

Barisia levicollis, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 247-248; Univ. 
Kansas Sci. Bull., vol. 33, 1949, pp. 247-248. 

TV^e.— U.S.N.M. No. 9362; "evidently obtained by one of the 
surveying parties of the United States and Mexican Boundary 
Survey," ^c?e Stejneger. 

Type locality. — "Mexican boundary." Probably Chihuahua, fide 
Tihen, loc. cit. 

Range. — Chihuahua and probably adjacent areas in Sonora. 
Reported from Chihuahua: north Chihuahua, Colonia Garcia, Meadow 
Valley, 65 miles east of Batopilas, Samachique. 

BARISIA RUDICOLLIS (Wiesmnnn) 

Gerrhonotus rudicollis Wiegmann, Isis von Oken, vol. 21, 1828, p. 379; Herpeto- 

logia Mexicana, 1834, pp. 33-34, pi. 10, figs. 1, 4. — MartIn del Campo, 

Anal. Inst. Biol., vol. 10, No. 3-4, 1939, p. 360, fig. 4. 
Barissia rudicollis, Gray, Catalogue of the lizards in the collection of the British 

Museum, London, 1845, p. 55. 
Gerrhonotus (Barissia) rudicollis, Bocourt, Mission scientifique au Mexique . . . , 

Etudes sur les reptiles, livr. 5, 1878, pi. 21B, figs. 3, 3a; and livr. 6, 1879, 

pp. 367-369. 
Barisia rudicollis, Tihen, Amer. Midi. Nat., vol. 41, 1949, pp. 238-240, pi. 1. 

Type. — Zool. Mus. Berlin; F. Deppe collector. 

Type locality. — Mexico. Here restricted to Hacienda de le Gavia, 
Mexico (state). 

Range. — Western Mexico and perhaps adjoining parts of Mich- 
oacan. Reported from Mexico: Hacienda de la Gavia. 



CHECKLIST OF REPTILES OF MEXICO 203 

Genus GERRHONOTUS Wiegmann 

Gerrhonotus Wiegmann, Isis von Oken, vol. 21, 1828, No. 3, p. 379. — Tihen, 

Amer. Midi. Nat., vol. 41, 1949, pp. 591-593. 
Pterogasienes Peale and Green, Journ. Acad. Nat. Sci. Philadelphia, vol. 6, 

1830, p. 234 (type, Scincus ventralis Peale and Green). 

Genotype. — Gerrhonotus tessellatus Wiegmann [=Gerrhonotus lioce- 
phalus Wiegmann]. 

Range. — Central Texas southward through the plateaux of Mexico 
to southwestern Chiapas. 

Species. — One. Five forms recognized, all occurring in Mexico. 

KEY TO SPECIES OF GERRHONOTUS 

1 . Three loreals (loreocanthals) on each side 2 

Four or more loreocanthals on each side 3 

2. Supranasals absent; azygous prefrontal much longer than broad; frontal 

widely separated from interparietal; all except lowermost anterior temporal 

touch fifth medial supraocular liocephalus austrinus (p. 204) 

Supranasals present; azygous prefrontal as broad as or broader than long; 
frontal in contact with or very narrowly separated from interparietal; onlj' 
2 uppermost anterior temporals in contact with fifth supraocular. 

liocephalus liocephalus (p. 203) 

3. Dorsal bands obsolete; venter nearly without marking; 52-60 dorsal scales 

(average 55. 6) ; tail to body ratio, 2. 5-2. 6; caudals 157-163; second primary 
temporal touches the fifth medial supraocular. 

liocephalus loweryi (p. 204) 
Dorsal bands distinct ; venter mottled and flecked 4 

4. Dorsal scales 45-54 (average 49); tail-body ratio, 1. 75-2. 1; caudals, 116-137. 

liocephalus infemalis (p. 204) 
Dorsal scales 49-52 (average 51); tail-body ratio 2. 3; caudals (approx.) 140. 

liocephalus ophiurus (p. 204) 

GERRHONOTUS LIOCEPHALUS UOCEPHALUS Wiegmann 

G[errhonotus] liocephalus Wiegmann, Isis von Oken, vol. 21, 1828, p. 381. 
Gerrhonotus liocephalus, Bocourt, Mission scientifique au Mexique . . . , Etudes 

sur les reptiles, livr. 5, 1878, pp. 342-346, pi. 21A, figs. 1, 2, 2a. 
Gerrhonotus liocephalus liocephalus, Cope, Ann. Rep. U, S. Nat. Mus. 1898 (1900), 

pp. 516-517.— Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, p. 369. 
G[errhonotus] tessellatus Wiegmann, Herpetologia Mexicana, pt. 1, 1834, pp. 

32-33, pi. 10, fig. 3 (substitute name for preceding). 
Scincus ventralis Peale and Green, Journ. Acad. Nat. Sci. Philadelphia, 1838, 

pt. 4, p. 233 (type locality, "Mining Districts of Mexico" = "Valley of Mexico" 

fide Cope, here restricted to Magdalena, D. F.; type in Acad. Nat. Sci. 

Philadelphia, two cotypes). 

Type. — Zool. AIus. Berlin; F. Deppe collector. 

Type locality. —Mexico (Oaxaca, fide Bocourt loc. cit., p. 344, here 
restricted to Tlapancingo). 

Range. — Central plateau region, from Guanajuato to Guerrero and 
Oaxaca. Reported from Distrito Federal: Magdalena, Mixiuhca; 
Guerrero: Omilteme; Mexico: Temascaltepec; Oaxaca: Llano Ocotal, 

861316—50 14 



204 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

La Concepci6n, Tlapancingo, Tres Cruces; Puebla: Cacaloapam; 
Morelos: Cuernavaca; Guanajuato: Silao. 

GEKKHONOTUS LIOCEPHALUS LOWERYI Tihen 

Gerrhonotus liocephalus loweryi Tihen, Trans. Kansas Acad. Sci., vol. 51, 1948, 
pp. 302-305. 

Type.— La. State Univ. Mus. No. 480; Marcella Newman collector. 

Type locality. — "Xilitla region" (neighborhood of Xilitla, San Luis 
PotosI). 

Range. — Known onlf from the type locality and possibly also from 
Ciudad del Maiz, San Luis Potosi. 

GERRHONOTUS LIOCEPHALUS OPHIURUS Cope 

Gerrhonotus ophiurus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, pp. 321-322. 
Pterogasterus ophiurus, Cope, Proc. Amer. Philos. Soc, vol. 17, 1877, p. 96. 
Gerrhonotus liocephalus ophiurus, Cope, Ann. Rep. U. S. Nat. Mus., 1898 (1900), 

pp. 516-517. 
Gerrhonotus lemniscatus Bocourt, Nouv. Arch. Mus. Hist. Nat. Paris, vol. 7, 

1872, pp. 105-107 (Mus. Hist. Nat. Paris; Veracruz). 

Type.— U.S.N.M. No. 30206; Francois E. Sumichrast collector. 

Type locality. — Orizaba, Veracruz, Mexico. 

Range. — Eastern foothills of the plateau. Known only from Vera- 
cruz: Orizaba, Jicaltepec, C6rdoba, Cerro Gordo; Puebla: Hueyta- 
malco (Teziutlan). 

GERRHONOTUS LIOCEPHALUS INFERNALIS Baird 

Gerrhonotus infernalis Baird, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 255. 
Gerrhonotus liocephalus infernalis, Cope, Ann. Rept. U. S. Nat. Mus., 1898 (1900), 

pp. 517-519, fig. 9.— Smith, Handbook of lizards, 1946, pp. 463-464, pi. 

131.— Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 593. 

Type.—? U.S.N.M. No. 3090; C. B. R. Kennerly collector. 

Type locality. — Devils Eiver, Tex. 

Range. — San Luis PotosI, Coahuila and probably adjoining states 
of Chihuahua and Nuevo Leon, northward to central Texas across 
the Rio Grande valley from Rio Grande City to mouth of Devil's 
River. Reported from Coahuila: Carmen Mountains, Cerro Encarna- 
ci6n, Sierra Guadelupe south of La Cuchilla, Monclova; San Luis 
Potosi: Alvarez, 

GERRHONOTUS LIOCEPHALUS AUSTRINUS Hartweg and Tihen 

Gerrhonotus liocephalus austrinus Hartweg and Tihen, Occ. Pap. Mus. Zool. 
Univ. Michigan, No. 497, 1946, pp. 6, 7. 

Type. — Univ. Michigan Mus. Zool. No. 94921 ; Eizi Matuda collector. 
Type locality. — 3,200 meters elevation on Cerro Male, Porvenir, 
Chiapas. 
Range. — Known only from the type locality. 



CHECKLIST OF REPTILES OF MEXICO 205 

Genus ELGARIA Gray 

Elgaria Ghat, Ann. Mag. Nat. Hist,, ser. 1, vol. 1, 1838, p. 390. — Tihbn, Amer. 

Midi. Nat., vol. 41, 1949, pp. 593-596, figs. 8-9. 
Tropidolepis Skilton (nee Cuvier), Amer. Journ. Sci., ser. 2, vol. 7, 1849, p. 202 

(type, T. scincicauda Skilton^ . 

Genotype. — Cordylus {Gerrhonotus) multicarinatiLsB\simYi\].e= Elgaria 
multicarinata multicarinata. 

Range. — Chihuahua and Baja California, northward through 
California to British Columbia, Utah, and Montana. 

Species. — Six species and 13 forms. Six forms occur in Mexco. 

KEY TO MEXICAN SPECIES OF ELGARIA 

1. Alternate black and white marks on labials (white spots somewhat ocellate). 

kingii group 4 
No alternate black and white marks on labials multicarinata group 2 

2. Temporal scales all or partly keeled 3 

Temporal scales smooth; granular area of lateral fold white with gray or black 
reticulations; scales on arm smooth; 2 rows of scales keeled on thigh; lateral 
body scales smooth cedrosensis (p. 206) 

3. Large (130 mm. maximum); dorsals all heavily keeled; two upper temporal 

rows sharply keeled, others weakly keeled or nearly smooth; three rows of 
keeled scales on arm; dark transverse bands indented with white markings 
5 scale rows above lateral fold and on back; ventral longitudinal lines faint 

or obsolete multicarinata webbii (p. 207) 

Smaller (114 mm. maximum); dorsals less strongly keeled; two upper temp- 
orals strongly keeled; 1 to 3 keeled rows on arm; dark ventral longitudinal 
lines; wide brown band on temporal region from eye to ear. 

multicarinata nana (p. 207) 

4. Granular area of lateral fold white crossed by black bands usually continuous 

with dark bodv bands; eight dorsal scale rows keeled moderately, lateral rows 
faintly keeled; 4 internasals of about same size; arm scales smooth; thigh 

scales with 2 rows keeled paucicarinatus (p. 206) 

Granular area of lateral fold dull gray; heavy broad dorsal bands on neck, body, 
tail, the latter almost encircling tail; usually 6 or 8 weakly keeled dorsal 
scale rows; scales on arm and thigh smooth 5 

5. Dorsal longitudinal scale rows usually 16; scales from occiput to posterior 

part of thigh in 51 to 56 transverse rows (average, 52.9); transverse row of 
four anterior temporals, uppermost always in contact with uppermost 
secondary temporal; granular area in fold usually black, kingii kingii (p. 205) 
Dorsal longitudinal scale rows usually 14; scales from occiput to posterior part 
of thigh 55 to 60 (average 57.3) ; four anterior temporals, uppermost sep- 
arated from uppermost secondary temporal (60 percent) ; granular area in 
fold usually gray kingii nobilis (p. 206) 

ELGARIA KINGII KINGH Gray 

Elgaria kingii Geay, Ann. Mag. Nat. Hist., vol. 1, 1838, p. 390. 

Elgaria kingii kingii, Tihen, Trans. Kansas Acad. Sci., vol. 51, 1948, pp. 299-300. 

Gerrhonotus kingii, Bocourt, Nouv. Arch. Mus. Hist. Nat. Paris, vol. 7, p. 106; 
Mission scientifique au Mexique . . ., fitudes sur les reptiles, livr. 5, 1878, 
pp. 339-342, pi. 21C, figs. 2, 2a (name attributed to Bell, his manuscript 
published by Gray). 



206 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

Gerrhonotus multifasciatus Dum^ril and Bocourt, Erp(5tologie g^n^rale, vol. 5, 
1839, p. 401 (type locality, "Mexico", here restricted to Mojarachic, Chi- 
huahua; Mus. Hist. Nat. Paris). 

Type. — Brit. Mus. Nat. Hist., adult (in bad state); from the T. 
Bell collection, collector unknown. 

Type locality. — Mexico [so stated by Boulenger, Catalogue of 
Lizards . . ., vol. 2, 1885, p. 2751. Tiben suggests "presumably 
Chihuahua." Here restricted to Mojarachic. 

Range. — Eastern Sonora and western Chihuahua, probably south- 
ward into Durango and Sinaloa. Reported from Chihuahua: Madera, 
Mojarachic, Chiricahui, Colonia Garcia (possibly intergrades with 
v/ith nobilis) ; Sonora: (no specific locality) , 

ELGARIA KINGn NOBILIS Baird and Girard 

Elgaria nobilis Baird and Girard, Proc. Acad. Nat. Sci. Philadelphia, vol. 6, 

1852, p. 129. 
Gerrhonotus nobilis, Baird, United States and Mexican boundary survej', vol. 2, 

Reptiles, 1859, p. 11, pi. 25, fig. 108. 
Elgaria kingii nobilis, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 596. 
Gerrhonotus kingii, Smith, Handbook of lizards, 1946, pp. 452-454, pi. 127. 

Type. — Possibly in Acad. Nat. Sci. Philadelphia. 

Type locality .—Fort Webster Copper Mines of the Gila [Santa 
Rita del Cobre], N. Mex. 

Range. — Southwestern New Mexico and southeastern Arizona. 
While no records are available from Mexico, specimens have been 
taken within a few hundred yards of the border, and the form must 
occur in Mexico (Sonora). 

elgaria PAUCICARINATUS (Fitch) 

Gerrhonotus paucicarinatus Fitch, Copeia, 1934, No. 4, 1935, pp. 172-173. 
Elgaria paucicarinata, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 595. 

Type.—Mus. Vert. Zool. Univ. California No. 11768, male; C. C. 
Lamb collector. 

Type locality. — Todos Santos, Baja California. 

Range. — Southern end of Baja California in the vicinity of the type 
locality. 

ELGARIA CEDROSENSIS (Fitch) 

Gerrhonotus cedrosensis Fitch, Copeia, 1934, No. 1, pp. 6-7. 

Elgaria cedrosensis, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 595. 

Type. — California Acad. Sci. No. 56187; J. R. Slevin collector. 
Type locality. — "Canon on southeast side of Cedros Island," Baja 
California. 

Range. — Cedros Island, Baja California. 



CHECKLIST OF REPTILES OF MEXICO 207 

ELGARIA MULTICARINATA NANA (Fitch) 

Oerrhonotus scincicauda nanus Fitch, Copeia, 1934, p. 7. 

Gerrhonotus multi-carinatus nanus, Fitch, Amer. Midi. Nat., vol. 20, 1938, pp. 

397-399, fig. 2. 
Elgaria multicarinata nana, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 596. 

Type.—Mus. Vert. Zool. Univ. California No. 5402; A. B. Howell 
collector. 

Type locality. — South Island, Los Coronados Islands, Baja 
California. 

Range. — Los Coronados Islands, Baja California. 

ELGARIA MULTICARINATA WEBBII (Baird) 

Gerrhonotus webhii Baird, Proc. Acad. Nat, Sci. Philadelphia, 1858, p. 225. 
Gerrhonotus scincicauda webhii, Grinnell and Camp, Univ. California Publ. ZooL, 

vol. 17, 1917, p. 168. 
Gerrhonotus multi-carinatus webhii, Fitch, Amer. Midi. Nat., vol. 20, 1938, pp. 

395-397.— Smith, Handbook of lizards, 1946, pp. 460-462, pi. 130. 
Elgaria multicarinatus webhii, Tihen, Amer. Midi. Nat., vol. 41, 1949, p. 596. 
Gerrhonotus scincicauda ignavus Van Denburgh, Proc. California Acad. Sci.. ser. 

3, vol. 4, 1905, p. 19, pi. 7, figs. 1-2 (type locality, San Martin Island, Baja 

California). 

Type.—V.S.'NM. No. 3078; T. H. Webb collector. 

Type locality. — "San Diego to El Paso," "probably from near 
vicinity of San Diego, San Diego Co. California [?]" (Fitch, loc. cit.). 
Here restricted to San Diego, Calif. 

Range. — Southern California and extreme northern Baja California. 
Reported in Baja California from San Pedro Martir Mountains, 

Family XENOSAURIDAE Cope 

Xenosauridae Cope, Proc. Acad. Nat. Sci. Philadelphia, vol. 18, 1866 (1867), 
p. 322. 

Genera. — A single genus, Xenosaurus Peters, is known. 
Range. — San Luis PotosI to Guatemala. 

Genus XENOSAURUS Peters 

Xenosaurus Peters, Monatsb. Acad. Wiss. Berlin, 1861, p. 453. 

Genotype. — Xenosaurus fasciatus FeteTS= Xenosaurus grandis 
(Gray). 
Range. — San Luis Potosi to Guatemala. 
Species. — Three. 

KEY TO SPECIES OF XENOSAURUS 

1. Arm with widely spaced tubercles on its dorsal surface; a row of supraoculars 

each 1}^ to 2 times as long as wide; cream, black spotted on venter 2 

Arm regularly covered with tubercles on dorsal surface; supraoculars not 
forming a series of more or less regular enlarged scales; venter uniformly gray 
or gray-white without spots or flecks newmanonira (p. 208) 



208 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

2. Scales bordering posterior gular fold small, rather widely separated from each 

other, considerably smaller than median chest scales grandis (p. 208) 

Scales bordering posterior gular fold larger, in contact with each other or nar- 
rowly separated, about as large as median chest scales -.rackhami (p. 208) 

XENOSAURUS NEWMANORUM Taylor 

Xenosaurus newmanorum Taylor, Univ. Kansas Sci. Bull., vol. 33, 1949, pp. 
183-187. 

Type. — Louisiana State Univ. No. 85; Marcella Newman collector. 
Type locality. — "Xilitla Region" (near Xilitla, San Luis PotosI). 
Range. — Known only from the type locality. 

XENOSAURUS GRANDIS (Gray) 

Cubina grandis Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 18, 1856, p. 270. 
Xenosaurus grandis, Cope, Proc. Acad. Nat. Sci. Philadelphia, 1866, p. 322. — 

BocouRT, Mission scientifique au Mexique . . . , fitudes sur les reptiles, 

livr. 5, 1878, pp. 303-306, pi. 20F, figs, la-d. If, Ig.— Cope, Ann. Rep. U. S. 

Nat. Mus., 1898 (1900), pp. 538-539.— Barrows and Smith, Univ. Kansas 

Sci. BuU., vol. 31, 1947, pp. 227-281, pis. 11-16. 
Xenosaurus fasciatus Peters, Monatsb. Acad. Wiss. Berlin, 1861, p. 453 (type 

locality, Huanusco [=Huatusco], Veracruz; Berl. Mus.). 

Type.— Brit. Mus. Nat. Hist.; A. Sall^ collector. 

Type locality. — C6rdoba, Veracruz (probably from nearby moun- 
tains) . 

Range. — Foothills of central Veracruz southward to the Isthmus. 
Reported from Veracruz: Huanusco [=Huatusco], C6rdoba, Orizaba; 
Oaxaca: "Oaxaca," Tehuantepec* 

XENOSAURUS RACKHAMI Stuart 

Xenosaurus rackhami Stuart, Proc. Biol. Soc. Washington, vol. 54, 1941, pp. 
47-48. — Smith, Journ. Washington Acad. Sci., vol. 39, 1949, p. 43. 

Type.—M.ns. Zool. Univ. Michigan No. 89072. 

Type locality. — Finca Volcan, 49 km. east of Coban, Alta Verapaz, 
Guatemala, 4,000 feet. 

Range. — The highlands of central eastern Chiapas and adjacent 
Guatemala. Recorded in Mexico only from Chiapas: Santa Rosa 
(near Comitan) . 

Family ANNIELLIDAE Boulenger 

Aniellidae Cope, Proc. Acad. Nat. Sci. Philadelphia, 1864, p. 230. 
Anniellidae BotrLENGER, Catalogue of the lizards in the British Museum, vol. 2, 
1885, p. 299. 

Genera. — Only a single genus, Anniella Gray, is known. 

Range. — Southwestern California and northwestern Baja California. 



< Fide Cope, U. S. Nat. Mus. Bull. 32, 1887, p. 39. A sight record cited by Martin del Campo, Folleto 
Div. Cient. Inst. Biol., No. 2, 1934, p. 7, of "Heloderma horridum" from Sierra de Zangolica, Veracruz, may 
belong with this species; it is very unlikely that Heloderma actually occurs on the Atlantic coast. 



CHECKLIST OF REPTILES OF MEXICO 209 

Genus ANNIELLA Gray 

Anniella Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 440. 

Genotype. — Anniella pulchra Gray. 

Range. — Southern California and northern Baja California. 

Species. — Two, one with two subspecies. 

KEY TO MEXICAN SPECIES OF ANNIELLA 

1. Rostral sharply pointed in profile; fourth supralabial largest. 

geronimensis (p. 209) 
Rostral rounded in profile ; second supralabial largest. 

pulchra pulchra (p. 209) 

ANNIELLA GERONIMENSIS Shaw 

Anniella geronimensis Shaw, Trans. San Diego Soc. Nat. Hist., vol. 9, 1940, 
pp. 225-228, 2 figs. 

Type.—L. M. Klauber No. 7543. 

Type locality. — Isla San Ger6nimo, Baja California. 

Range. — Known only from the type locality. 

ANNIELLA PULCHRA PULCHRA Gray 

Anniella pulchra Gray, Ann. Mag. Nat. Hist., ser. 2, vol. 10, 1852, p. 440. — 

Van Denburgh, Occ. Pap. California Acad. Sci., No. 10, 1922, pp. 465-467, 

pi. 42. 
Anniella pulchra pulchra, Grinnell and Camp, Univ. California Publ. Zool., 

vol. 17, 1917, p. 170.— Miller, Copeia, 1943, p. 2. 
Anniella texana Boulenger, Ann. Mag. Nat. Hist., ser. 5, vol. 20, 1887, p. 50 

(Brit. Mus. Nat. Hist.; El Paso, Tex., in error, here restricted to San Diego, 

California) . 

Type. — Brit. Mus. Nat. Hist.; Sir John Richardson collector. 

Type locality. — California, here restricted to San Diego. 

Range. — Pacific slopes of central and southern California, and 
extreme northern Baja California, west of the desert and the Sierra 
San Pedro Martir. Recorded in Baja California: San Salado Canon, 
San Jos6, San Quintin, Los Coronados Islands. 

Subclass Archosauria 

Archosauria Romer, Vertebrate paleontology, 1945, p. 597. 
Orders. — A single living order, the Loricata, exists. 

Order LORICATA 

Loricata Merrem, Tentamen systematis amphibiorum, 1820, p. 34. 

Families. — Three families exist at the present time, two of which 
occm' in Mexico. 



210 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

KEY TO LIVING FAMILIES OF LORICATA 

1. Dentary teeth 25-26, none received by sockets in upper jaw; maxillary bones 

broadly in contact with each other on dorsal surface of skull; mandibular 
symphysis extraordinarily long, only 2 or 3 teeth at rear of mandible not 

opposite the symphysis Qavialidae * 

Dentary teeth 20 or fewer, all or nearly all received by sockets in upper jaw; 
maxillary bones never in contact with each other on dorsal surface of skull; 
mandibular symphysis shorter, 4 or more teeth at rear of mandible not 
opposite the symphysis 2 

2. Fourth mandibular tooth fitting into a pit in upper jaw, and 17 to 22 dentary 

teeth Alligatoridae (p. 211) 

Fourth mandibular tooth fitting into a notch in upper jaw or, if into a pit, 
dentary teeth 16 or fewer .Crccodylidae (p. 210) 

Family CROCODYLIDAE Gray 

Crocodilidae Gray, Ann. Philos., ser. 2, vol. 10, 1825, p. 195. — Mertens, Sencken- 
bergiana, vol. 26, 1943, pp. 252-312. 

Genera. — Four genera and 17 species and subspecies are recognized 
at the present time; one genus occurs in the Americas. 
Range. — World wide in tropical lands. 

Genus CROCODYLUS Laurenti 

Crocodylus Laurenti, Specimen medicum exhibens synopsin reptilium, 1768, p. 53. 
Molinia Gray, Catalogue of the tortoises, crocodiles and amphisbaenians in the 

collection of the British Museum, 1844, pp. 60, 272 (type, Crocodylus ameri- 

canus Laurenti =? Crocodylus acutus Cuvier). 

Genotype. — Crocodylus niloticus Laurenti. 

Species. — Twelve species, two with two subspecies, are recognized; 
only four species, one with two subspecies, occur in the New World, 
and two in Mexico. 

Range. — Northern Australia, New Guinea, East Indies, Siam, 
Ceylon, tropical Africa and India, Madagascar; southern Florida, 
Greater Antilles, northern Mexico south to Ecuador and Venezuela. 

KEY TO MEXICAN SPECIES OF CROCODYLUS 

1, Premaxillo-maxillary suture transverse moreletii (p. 211) 

Premaxillo-maxillary suture extending posteriorly in an arc to level of posterior 
border of seventh tooth acutus (p. 210) 

CROCODYLUS ACUTUS ACUTUS Cuvier' 

Crocodilus acutus Cuvier, Ann. Mus. Hist. Nat. Paris, vol. 10, 1807, p. 55, pi. 

1, fig. 3, pi. 2, fig. 3. 
Crocodylus acutus acutus, Muller and Hellmich, Ibero-Amerik. Stud., vol. 

13, 1940, pp. 128-130. 

' Not in Mexico; restricted to India. 

• Allocation of the Mexican and, for that matter, Central American specimens is at present very uncertain. 
South American (Colombia, Ecuador) specimens are referrable to the race C. a. lewyanus, and Santo Do 
mlngo specimens to C. a. acutus, but no study has been made of material from elsewhere. 



CHECKLIST OF REPTILES OF MEXICO 211 

f Crocodilus americanus Laurenti, Specimen medicum exhibens synopsin rep- 
tilium, 1768, p. 54 (type, pi. 106 in Seba's "Locupletissimi Rerum Naturalium 
Thesauri Occurata Descriptio," vol. 1, 1734; type locality, America, here 
restricted to Veracruz, Veracruz; unidentifiable). — Bocourt, Mission scien- 
tifique au Mexique . • . , Etudes sur les reptiles, livr. 1, 1870, pp. 
30-31, pi. 8, fig. 1, pi. 9, fig. 1. 

Crocodilus biscutatus Ctjvier, Ann. Mus. Hist. Nat. Paris, vol. 10, 1807, p. 53, 
pi. 2, fig. 6 (type presumably in Mus. Hist. Nat. Paris; type locality, 
"Senegal", in error; here restricted to Tampico, Tamaulipas). 

Crocodilus mexicanus Bocourt, Nouv. Arch. Mus. Hist. Nat. Paris, 1869, pp. 
20-21 (type presumably in Mus. Hist. Nat. Paris; type locality, Tampico, 
Tamaulipas) ; Mission scientifique au Mexique . . . , fitudes sur les reptiles, 
livr. 1, 1870, pp. 34-35, pi. 8, fig. 3. 

Crocodilus pacificus Dum^ril and Bocourt, Mission scientifique au Mexique . . ., 
Etudes sur les reptiles, livr. 1, 1870, pp. 31-33, pi. 9, fig. 5 (type presumably 
in Mus. Hist. Nat. Paris; type locality, Rio Nagualate, Guatemala). 

Type. — Mus. Hist. Nat. Paris, presumably. 

Type locality. — Santo Domingo (Republic). 

Range. — Greater Antilles, Central America north to Tamaulipas 
and Sinaloa. Reported from various localities in the states of 
Quintana Roo (Isla de Mujeres), Campeche, Cohma, Guerrero, 
Michoaclin, Oaxaca, Tabasco, Tamauhpas, Veracruz, and Maria 
Magdalena Island, Nayarit. 

CROCODYLUS MORELETH Dum&ll and Dumeril 

Crocodilus moreletii Dum6ril and Dumeril, Catalogue m^thodique de la collection 
des reptiles, 1851, p. 28. — Bocourt, Mission scientifique au Mexique . . ., 
fitudes sur les reptiles, livr. 1, 1870, pp. 37-38, pi. 9, fig. 2.— Schmidt, Publ. 
Field Mus. Nat. Hist., zool. ser., vol. 12, 1924, pp. 79-84, pi. 6, figs. 2, 7, 8. 

Alligator lacordairei Preudhomme de Borre, Bull. Acad. Roy. Sci. Lett. Beaux- 
Arts Belgique, ser. 2, vol. 28, 1869, p. 110, pi. (type presumably in Brussels; 
M. Levy collector; type locality, Belize, British Honduras). 

Type. — Mus. Hist. Nat. Paris, presumably. 

Type locality. — Lake Pet^n, Guatemala. 

Range. — Atlantic slopes from Guatemala to Tamaulipas. Reported 
in Mexico from the states of Campeche, Chiapas (Palenque), Cohma, 
Tabasco, Tamaulipas, Veracruz; records from Colima (by Duges) 
and the Pacific side of Chiapas (Tapachula, by Sumichrast) are to be 
considered erroneous until corroborated. 

Family ALLIGATORIDAE Gray 

Alligatoridae Gray, Catalogue of tortoises, crocodiles and amphisbaenians in the 
British Museum, 1844, p. 56. — Mertens, Senckenbergiana, vol. 26, 1943, 
pp. 252-312. 

Genera. — Four, of which only one occurs in Mexico.' 



' Alligator missif:sippiensis is recorded from "Mexico" by Garman (Bull. Kssex Inst., vol. 16, 1884, p. 11) 
and Boettger (Kat. Kept. Senckenb. Mus., pt. 1, 1893, p. 19), but we regard the records as erroneous 
until verlfled; 



212 BULLETIN 19 9, XWITED STATES NATIONAL MUSEUM 

Genus CAIMAN Spix 

Caiman Spix, Animalia nova sine species lacertarum . . . Brasiliam . . ., 1825, p. 3. 
Perosuchus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1868, p. 203 (genotype, 
Perosuchus fuscus Cope) . 

Genotype. — Caiman Jissipes Spix (=CrocodiIus latirostris Daudin). 
Species. — Two species, one with three subspecies, are recognized; 
one occurs in Mexico. 

Range. — The Isthmus of Tehuantepec to Argentina. 

CAIMAN CPOCODILUS FUSCUS (Cope) 

Perosuchus fuscus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1868, p. 203. 
Caiman crocodilus fuscus, Mertens, Senckenbergiana, vol. 26, 1943, p. 275. 
All[igator] (Jacare) Cfiiaparius Bocourt, Ann. Sci. Nat., ser. 6, vol. 3, No. 12, 

1876, pp. 1-2 (type probably in Mus. Hist. Nat. Paris; type locality, Tonald, 

Chiapas) . 
All[igator] Ohiapasius Bocourt, Journ. Zool., vol. 5, 1876, pp. 400-401 (equivalent 

to preceding). 

Type. — Unknown. 

Type locality. — Rio Magdalena, Colombia. 

Range. — Oaxaca and perhaps Michoacan to Colombia. Reported 
from the states of Oaxaca: Tapanatepec, Agua Fria; and Chiapas: 
Tonala, Belen, Colonia Soconusco. A record from the Balsas 
Valley in Michoacan (by Gadow) may be correct, but one from 
Yucatan (Tozzer) is, we believe, certainly incorrect. 

SPECIES INQUIRENDAE 

A few species whose natural ranges remain unknown have been 
included in their appropriate systematic place in the preceding pages. 
A few others, described from "Mexico," have since been found to occur 
elsewhere. Among these are the following: 

HERPETOCHALCIS HETEROPUS Boetlger 

Herpetochalcis heteropus Boettger, Bericht. Offenb. Ver. Nat., vols. 22-23, 1883, 
pp. 150-151. 

The type locality was stated to be perhaps California, or Mexico, 
or Central America. The name is apparently a synonym of Chalcides 
heteropus Lichtenstein, Nomencl. Mus. Zool. Berol., 1856, p. 17 
(=Bachia heteropa) described from La Guaira. Venezuela, and re- 
stricted in range to that country. 

LEIOSAURUS BELLI DmnfirU and Bibron 

Leiosaurus belli 'DvM^mii and Bibron, Erp^tologie g^n^rale, vol. 4, 1837, p. 242. 

The type locality was stated to be "Mexico," but the species (and 
genus) has since been discovered to be restricted to Argentina. 



CHECKLIST OF REPTILES OF MEXICO 213 

ANEUPORUS OCCIPITAUS Bocoort 

AneuTporus occipitalis Bocourt, Mission scientifique au Mexique . . ., Etudes sur 
les reptiles, livr. 4, 1874, pp. 215-217, pi. 18, fig. 1. 

This has since been renamed Tropidurus hocourti by Boulenger, since 
Bocourt's name is suppressed as a homonym of Tropidurus occipitalis 
(Peters). The species occurs in Peru. 

SPHAERODACTYLUS ANTHRACINUS Cope 

Sphaerodactylus anthracinus Cope, Proc. Acad. Nat. Sci. Philadelphia, 1861, p. 
500 (type locality, "Mexico" [Jalapa]). 

Barbour (Mem. Mus. Comp. Zool. Harvard, vol. 47, 1921, pp. 
258-259) associates this name with a species from Andros Island, but 
Boulenger (Catalogue of the lizards in the British Museum, vol. 1, 
1885, p. 225) records a specimen from "San Domingo," and Peters 
(Monatsb. Akad. Wiss. Berlin, 1869, p. 874) records one from the 
state of Puebla (possibly in the vicinity of Izucar de Matamoros). 
The final word has not yet been said. For a discussion see Taylor, 
Univ. Kansas Sci. Bull., vol. 31, 1947, pp. 300-301. 

Three other species recorded from "Mexico," but with a type 
locality elsewhere, are known to be restricted to extralimital areas: 

BACHIA DORBIGNYI (Dum6rU and Bibron) 

Bachia dorbignyi (Dum^ril and Bibron), Mttller, Reisen in den Vereinigten 
Staaten, Canada und Mexico, vol. 3, pt. 3, 1865, p. 604. 

This species, still recognized by this name, occurs in Bolivia and 
Chile. 

ECPHYMOTES OBTUSIROSTRIS (Wiegmann) 

Ecphymotes ohtusirostris (Wiegmann), Mtjller, op. cit., p. 600. 

This name is a synonym of Anisolepis undulatus (Wiegmann), 
which species occurs in Brasil. 

TROPIDURUS PTYCHOPLEURUS Lichtensteln 

Tropidurus ptychopleurus Lichtensteln, Mxjller, op. cit., p. 602. 

This name is a synonym of Liolaemus tenuis (Dum^ril and Bibron), 
which species occurs in Chile. 

One other name, supposedly based upon a Mexican specimen, has 
been impossible to place: 



214 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 

DASIA MICROCEPHALUS (Hallowell) 

Euprepis microcephalus Hallowell, Proc. Acad. Nat. Sci. Philadelphia, vol. 8, 
1856, p. 155; Trans. Amer. Philos. Soc, ser. 2, vol. 11, 1857, pp. 79-80. 

Diploglossus microcephalus, Boulenger, Catalogue of the lizards in the British 
Museum, vol. 3, 1887, p. 504, 

[Dasia] microcephalus, Smith, Proc. U. S. Nat. Mus., vol. 92, 1942, pp. 369-370 

Type. — Acad. Nat. Sci. Philadelphia; W. H. Keating collector. 

Type locality. — Mexico. 

Remarks. — The first words in Hallowell's original description are 
"Syn. Scincus ventralis, Peale and Green." Then follows the descrip- 
tion. We believe the citation of Peale and Green's name as a synonym 
(an earlier one at that) of Hallowell's own name could be construed 
to make his name unavailable, that is, actually a synonym of Scincus 
ventralis and thus a part of the synonymy of Gerrliunotus I. liocephalus. 
The fact remains that the animal described by Hallowell, and pre- 
sumably the type of his name, is far different from Gerrhonotus. We 
recommend association of the name with the species represented by 
the animal described and not assignment to the synonomy of Scincus 
ventralis. 

The type is not in good condition (portions of the head mutilated), 
but appears to belong to a non-American genus, probably Dasia.^ 
It may possibly be one of the original series of three specimens of 
Peale and Green's Scincus ventralis, since only two of them are now 
present in the series labeled as cotypes, and thus would be explained 
Hallowell's citation of Scincus ventralis as a synonym of his species. 
It is also possible that a confusion of specimens occurred, the original 
Gerrhonotus being exchanged for the present type of microcephalus, 
which Hallowell erroneously thought was one of Peale and Green's 
cotypes. One of these alternatives must be true: Either a peculiar 
skink, unloiown except by the type of microcephalus, occurs in Mexico, 
or else some shift of specimens occurred in the Philadelphia Academy 
collections between 1830 and 1856. We favor the latter alternative. 

Still another species is represented by a specimen questionably 
secured in Mexico, whUe the range of the species is suspected of being 
entirely extralimital. 

DIPLOGLOSSUS MONOTROPIS (Kuh!) 

Diploglossus monotropis (Kuhl), Beitrage zur Zoologie und vergleichende Ana- 
tomic, 1820, p. 128. 

The species is known to occur from Costa Rica to Ecuador. The 
U. S. National Museum has one specimen questionably from "Colima." 
Occurrence in Mexico is highly questionable. 

> We are indebted to Dr. E. R. Dunn for this information. 



CHECKLIST OF REPTILES OF MEXICO 215 

STATE LISTS 

The accompanying lists have been constructed in the same way as 
those for the snakes (U. S. Nat. Mus. Bull. 187, 1945, pp. 202-203) 
and amphibians (U. S. Nat. Mus. Bull. 194, 1948, pp. 100-101). 
Table 2 shows for each state and each of the indicated groups, first, 
the number of forms recorded; second, the number of genera repre- 
sented (these two numbers separated by a diagonal line) ; and third, 
the relative position of the state (so far as number of forms is con- 
cerned) as compared with other states (this number in parentheses). 
The lists of snakes and amphibians have been brought more or less 
up to date by inclusion of the additional forms defined and recorded 
by various authors such as Blair, Bogert, Burger, Conant, Klauber, 
Smith, Tanner, and Taylor (see Univ. Kansas Sci. Bull., vol. 33, 1950, 
pp. 313-380, for references). 



216 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



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CHECKLIST OF REPTILES OF MEXICO 



217 



AGUASCALIENTES 



Kinostemon integrum 
Cnemidophorus sackii communis 



Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 



BAJA CALIFORNIA 



Amyda emoryi 
Caretta caretta gigas 
Clemmj^s marmorata pallida 
Dermochelys coriacea 
Eretmochelys imbricata 
Gopherus agassizii 
Pseudemys scripta nebulosa 
Bipes biporus 
AnnieUa geronimensis 
Anniella pulchra pulchra 
Callisaurus draconoides draconoides 
Callisaurus draconoides carmenensis 
Callisaurus draconoides crinitus 
Callisaurus draconoides gabbii 
Callisaurus draconoides splendidus 
Cnemidophorus catalinensis 
Cnemidophorus ceralbensis 
Cnemidophorus hyperythrus beldingi 
Cnemidophorus hyperythrus caeruleus 
Cnemidophorus hyperythrus danheimae 
Cnemidophorus hyperythrus hyperythrus 
Cnemidophorus hyperythrus pictus 
Cnemidophorus hyperythrus schmidti 
Cnemidophorus labialis 
Cnemidophorus maximus 
Cnemidophorus tigris canus 
Cnemidophorus tigris celeripes 
Cnemidophorus tigris multiscutatus 
Cnemidophorus tigris rubidus 
Cnemidophorus tigris tigris 
Coleonyx variegatus variegatus 
Coleonyx variegatus abbotti 
Coleonyx variegatus peninsularis 
Coleonyx variegatus slevini 
Crotaphytus collaris baileyi 
Crotaphytus insularis 
Ctenosaura hemilopha 
Dipsosaurus carmenensis 
Dipsosaurus catalinensis 
Dipsosaurus dorsalis dorsalis 
Dipsosaurus dorsalis lucasensis 
Elgaria cedrosensis 
Elgaria multicarinata nana 
Elgaria multicarinata webbii 
Elgaria paucicarinata 
Eumeces gilberti rubricaudatus 
Eumeces lagunensis 



Eumeces skiltonianus skiltonianus 

Gambelia wislizenii wislizenii 

Petrosaurus repens 

Petrosaurus thalassinus 

Phrynosoma cerroense 

Phrynosoma coronatum coronatum 

Phrynosoma coronatum blainvillii 

Phrynosoma coronatum frontale 

Phrynosoma coronatum jamesi 

Phrynosoma m'callii 

Phrynosoma plat}^rhinos platyrhinos 

Phrynosoma solare 

?Phyllodactylus tuberculosus 

Phyllodactylus unctus 

Sator angustus 

Sator grandaevus 

Sauromalus ater 

Sauromalus australis 

Sauromalus hispidus 

Sauromalus klaubeii 

Sauromalus slevini 

Sceloporus graciosus vandenburgianus 

Sceloporus magister lineatulus 

Sceloporus magister magister 

Sceloporus magister monserratenais 

Sceloporus magister rufidorsum 

Sceloporus magister zosteromus 

Sceloporus occidentalis biseriatus 

Sceloporus orcutti licki 

Sceloporus orcutti orcutti 

Streptosaurus mearnsi 

Streptosaurus slevini 

Uma notata notata 

Urosaurus graciosus 

Urosaurus microscutatus 

Urosaurus nigricaudus 

Urosaurus ornatus symmetricus 

Uta concinna 

Uta mannophorus 

Uta martinensis 

Uta squamata 

Uta stansburiana elegans 

Uta stansburiana hesperis 

Uta stansburiana stejnegeri 

Uta stellata 

Xantusia gilberti 

Xantusia henshawi 

Xantusia vigilis 



218 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



CAMPECHE 



Chelydra serpentina 

Claudius angustatus 

Dermatemys mawii 

Eretnaochelys imbricata 

Geoemyda areolata 

Kinosternon acutum 

Kinosternon cruentatum cruentatum 

Kinosternon leucostomum 

Pseudemys scripta ornata 

Ameiva undulata gaigeae 

Ameiva undulata stuarti 

Anolis kidderi 

Anolis lemurinus bourgeaei 

Anolis limifrons rodriguezii 

Anolis sagrei mayensis 

Anolis tropidonotus 

Anolis ustus 

Basiliscus vittatus 

Cnemidophorus deppii oligoporus 



Cnemidophorus sackii angusticeps 
Coleonyx elegans elegans 
Corythophanes cristatus 
Ctenosaura similis similis 
Enyaliosaurus erythromelas 
Eumeces schwartzei 
Hemidactylus turcicus turcicus 
Iguana iguana rhinolopha 
Laemanctus serratus 
Scincella cherriei ixbaac 
Mabuj-a mabouya mabouya 
Sceloporus chrysostictus 
Sceloporus lundelli lundelli 
3celoporus serrifer serrifer 
Sceloporus serrifer plioporus 
Sceloporus teapensis 
Sphaerodactylus glaucus glaucus 
Crocodylus acutus acutus 
Crocodylus moreletii 



CHIAPAS 



Geoemyda pulcherrima incisa 

Kinosternon abaxillare 

Kinosternon cruentatum cruentatum 

Kinosternon leucostomum 

Lepidochelys olivacea 

Pseudemys scripta ornata 

Abronia matudai 

Abronia ochoterenai 

Ameiva festiva edwardsi 

Ameiva undulata hartwegi 

Ameiva undulata parva 

Ameiva undulata stuarti 

Ameiva undulata thomasi 

Anolis biporcatus 

Anolis humilis uniformis 

Anolis limifrons rodriguezii 

Anolis pentaprion 

Anolis senceus 

Barisia moreleti rafaeli 

Barisia moreleti temporalis 

Basiliscus vittatus 

Celestus rozellae 

Cnemidophorus deppii deppii 

Cnemidophorus guttatus immutabilis 

Cnemidophorus sackii bocourti 

Coleonyx elegans elegans 

Corythophanes cristatus 

Corjrthophanes hernandezii 

Corythophanes percarinatus 



Ctenosaura similis similis 

Eumeces sumichrasti 

Gerrhonotus liocephalus austrinus 

Gonatodes fuscus 

Heloderma horridum 

Iguana iguana rhinolopha 

Scincella assata assata 

Scincella assata taylori 

Scincella cherriei cherriei 

Lepidophyma flavimaculata fiavimacu- 

lata 
Lepidophyma smithii smithii 
?L€pidophyma smithii tehuanae 
Mabuya mabouya mabouya 
Phrynosoma asio 
Phyllodactylus magnus 
Sceloporus carinatus 
Sceloporus malachiticus acanthinus 
Sceloporus malachiticus taeniocnemis 
Sceloporus melanorhinus stuarti 
Sceloporus prezygus 
Sceloporus siniferus siniferus 
Sceloporus squamosus 
Sceloporus teapensis 
Sceloporus variabilis variabilis 
Urosaurus bicarinatus anonymorphus 
Xenosaurus rackhami 
Caiman crocodilus fuscus 
Crocodylus moreletii 



CHECKLIST OF REPTILES OF MEXICO 



219 



CHIHUAHUA 



Chrysemjs picta bellii 
?Gopherus berlandieri 
Kinosternon hirtipes 
Kinosternon sonoriense 
Terrapene ornata 
Anolls nebuloides 
Barisia levicollis 
Cnemidophorus inornatus 
Cnemidophorus sackii communis 
Cnemidophorus sackii stictogrammus 
Cnemidophorus sackii scalaris 
Cnemidophorus tigris marmoratus 
Crotaphj'tus collaris baileyi 
Ctenosaura hemilopha 
Elgaria kingii kingii 
Eumeces callicephalus 
Eumeces multivirgatus 
Eumeces obsoletus 
Eumeces parviauriculatus 
Gambelia wisHzenii wislizenii 
Holbrookia bunkeri 



Holbrookia maculata approximau3 
Holbrookia texana 
Phryuosoma cornutum 
Phrynosoma douglassii hernandesi 
Phrj'nosoma modestum 
Phrynosoma orbiculare orbiculare 
Phyllodactylus homolepidurus 
Sceloporus clarkii clarkii 
Sceloporus gramraicus disparilis 
Sceloporus jarrovii jarrovii 
Sceloporus magister magister 
Sceloporus neLsoni 
■'^"eloporus poinsettii 
Sceloporus scalaris slevini 
Sceloporus undulatus consobrinus 
Sceloporus undulatus virgatus 
Urosaurus ornatus caeruleus 
Urosaurus ornatus linearis 
Urosaurus ornatus schmidti 
Urosaurus unicus 
Uta stansburiana ste^negeri 



COAHUILA 



Amyda emoryi 
Gopherus berlandieri 
Kinosternon flavescens flavescens 
Kinosternon flavescens stejnegeri 
Pseudemys scripta elegans 
Pseudemys scripta gaigeae 
Terrapene coahuila 
Barisia imbricata ciliaris 
Cnemidophorus inornatus 
Cnemidophorus sackii communis 
Cnemidophorus sackii semifasciatus 
Cnemidophorus sackii scalaris 
Cnemidophorus tigris marmoratus 
Coleonyx brevis 
Crotaphytus collaris baileyi 
Crotaphytus reticulatus 
Eumeces obsoletus 
Gambelia wislizenii wislizenii 
Gerrhonotus liocephalus infernalis 
Holbrookia maculata approximans 
Holbrookia maculata dickersonae 



Holbrookia maculata lacerata 
Holbrookia texana 
Scincella laterale 
Phrynosoma cornutum 
Phrynosoma modestum 
Sceloporus cautus 
Sceloporus couchii 
Sceloporus goldmani 
Sceloporus grammicus disparilis 
Sceloporus jarrovii oberon 
Sceloporus magister magister 
Sceloporus merriami annulatus 
Sceloporus olivaceus 
Sceloporus ornatus caeruleus 
Sceloporus ornatus ornatus 
Sceloporus parvus parvus 
Sceloporus poinsettii 
Sceloporus scalaris slevini 
Sceloporus undulatus consobrinus 
Sceloporus variabilis marmoratus 
Uma exsul 
Uta stansburiana stejnegeri 



861316—50 15 



220 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



COLIMA 



Geoemyda rubida 

Geoemyda pulcherriiua pulcherrima 

Kinosternon hirtipes 

Kinosternon iiitegruni 

Lepidochelys olivacea 

Ameiva undulata sinistra 

Anolis nebuloides 

Anolis nebulosus 

Anolis schmidti 

Basiliscus vittatiis 

Cnemidophorus deppii lineatissimus 

Cnemidophorus giiltatus immutabilis 

Cnemidophorus sackii communis 

Coleonyx elegans nemoralis 

Ctenosaura pectinata 

Eumeces colimensis 

Eumeces parvulus 

Heloderma hoiridum 



Iguana iguana rhinolopha 
?Laemanctus longipes 
Scincella assata tayloii 
Mabuya mabouya mabouya 
Phrynosoma asio 
Phrynosoma orbiculare dugesii 
Phyllodactylus lanei 
Sceloporus dugesii dugesii 
Sceloporus grammicus microlepidotus 
Sceloporus horridus oligoporus 
Sceloporus melanorhinus calligaster 
Sceloporus pyrocephalus 
Sceloporus utiformis 
Urosaurus auriculatus 
Urosaurus bicarinatiis tuberculatus 
Urosaurus clarionensis 
Crocodylus acutus acutus 
Crocodylus moreletii 



DISTRITO FEDERAL 



Kinosternon hirtipes 

Barisia imbricata imbricata 

Cnemidophorus sackii communis 

Eumeces copei 

Gerrhonotus liocephalus liocephalus 

Phrvnosoma orbiculare orbiculare 



Sceloporus aeneus aeneua 
Sceloporus grammicus microlepidotus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 



DURANGO 



Kinosternon flavescens stejnegeri 
Kinosternon sonoriense 
Pseudemys scripta gaigeae 
Anolis nebulosus 
Barisia imbricata ciliaris 
Cnemidophorus sackii communis 
Cnemidophorus sackii scalaris 
Cnemidophorus tigris marmoratus 
Coleonyx brevis 
Coleonyx fasciatus 
Crotaphytus collaris baileyi 
Ctenosaura pectinata 
Eumeces brevirostris 
Eumeces callicephalus 
Holbrookia maculata approximans 
Holbrookia maculata dickersonae 
Holbrookia texana 



Phrynosoma cornutum 
Phrynosoma douglassii brachycercum 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
? Phyllodactylus lanei 
Sceloporus grammicus disparilis 
Sceloporus horridus oligoporus 
Sceloporus jarrovii jarrovii 
Sceloporus lineolateralis 
Sceloporus maculosus 
Sceloporus magister magister 
Sceloporus poinsettii 
Sceloporus scalaris scalaris 
Sceloporus scalaris slevini 
Sceloporus spinosus spinosus 
Sceloporus undulatus consobrinus 
Uta stansburiana stejnegeri 



CHECKLIST OF REPTILES OF MEXICO 



221 



GUANAJUATO 



Kinosternon hirtipes 

Kinosternon integrum 

Barisia imbricata ciliaris 

Barisia imbricata imbricata 

Cnemidophorus sackii communis 

Eumeces callicephalus 

Eumeces dugesii 

Gerrhonotus liocephalus liocephalus 

Holbrookia maculata dickersonae 

Laemanctus serratus 

Phrynosoma boucardi 



Phrynosoma orbiculare orbiculare 
Sceloporus aeneus aeneus 
Sceloporus dugesii intermedius 
Sceloporus grammicus disparilis 
Sceloporus grammicus microlepidotus 
Sceloporus jarrovii minor 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 
Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 



GUERRERO 



Chelonia mydas 

Geoemyda pulcherrima pulcherrima 

Geoemyda rubida 

Kinosternon integrum 

Lepidochelys olivacea 

Abronia deppii 

Ameiva undulata dextra 

Anolis dunni 

Anolis gadovii 

Anolis liogaster 

Anolis megapholidotus 

Anolis nebuloides 

Anolis nebulosus 

Anolis taylori 

Barisia gadovii gadovii 

Basiliscus vittatus 

Bipes canaliculatus 

Bipes tridactylus 

Cnemidophorus deppii deppii 

Cnemidophorus deppii lineatissimus 

Cnemidophorus guttatus immutabilis 

Cnemidophorus sackii sackii 

Coleonyx elegans nemoralis 

Ctenosaura pectinata 

Eumeces brevirostris 

Eumeces ochoterenai 

Gerrhonotus liocephalus liocephalus 

Heloderma horridum 



Hemidactylus frenatus 
Iguana iguana rhinolopha 
Scincella assata taylori 
Mabuya mabouya mabouya 
Phrynosoma asio 
Phrynosoma taurus 
Phyllodactylus bordai 
PhyUodactylus delcampi 
Phyllodactylus lanei 
Phyllodactylus magnatuberculatus 
Phyllodactylus magnus 
?Sceloporus asper 
Sceloporus formosus scitulus 
Sceloporus gadoviae 
Sceloporus grammicus grammicus 
Sceloporus horridus horridus 
Sceloporus horridus oligoporus 
Sceloporus melanorhinus caUigaster 
Sceloporus mucronatus omiltemanus 
Sceloporus ochoterenai 
Sceloporus pyrocephalus 
Sceloporus siniferus siniferus 
Sceloporus stejnegeri 
Sceloporus utiformis 
Urosaurus bicarinatus bicarinatus 
Urosaurus bicarinatus anonymorphus 
Crocodylus acutus acutus 



861316—50- 



-16 



222 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



HIDALGO 



Kinosternon hirtipes 
Abronia taeniata taeniata 
Barisia imbricata imbricata 
Barisia imbricata ciliaris 
Cnemidophorus sackii communis 
Eumeces lynxe furcirostris 
Eumeces lynxe lynxe 
Gaigeia gaigeae 
Gaigeia sylvatica 
Laemanctus serratus 
Scincella gemmingeri gemmingeri 
Scincella gemmingeri forbesorum 



Phrynosoma boucardi 
Phrynosoma orbiculare orbiculare 
Sceloporus aeneus bicanthalis 
Sceloporus grammicus disparilis 
Sceloporus grammicus microlepidotus 
Sceloporus jarrovii immucronatus 
Sceloporus mucronatus mucronatus 
Sceloporus parvus scutulatus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 



JALISCO 



Kinosternon integrum 

Pseudemys scripta ornata 

Ameiva undulata sinistra 

Anclis nebulosus 

Baiisia imbricata imbricata 

Basiliscus vittatus 

Cnemidophorus deppii lineatissimus 

Cnemidophorus sackii communis 

Ctenosaura pectinata 

Eumeces brevirostris 

Eumeces callicephalus 

Eumeces dugesii 

Heloderma horridum 

Holbrookia maculata approximans 

Iguana iguana rhinolopha 

Scincella assata taylori 

Phrynosoma orbiculare orbiculare 

?Phyllodactylus lanei 

?Sceloporus aeneus aeneus 



Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Sceloporus 
Urosaurus 
Urosaurus 



asper 

buUeri 

clarkii boulengerl 

dugesii dugesii 

grammicus microlepidotus 

heterolepis 

horridus albiventris 

horridus oligoporus 

melanorhinus caliigaster 

nelsoni 

pyrocephalus 

scalaris scalaris 

scalaris unicanthalis 

spinosus spinosus 

torquatus melanogaster 

utiformis 

bicarinatus tubereulatus 



MEXICO 



Ejnosternon hirtipes 
Barisia imbricata imbricata 
Barisia rudicollis 
Cnemidophorus sackii communis 
Eumeces copei 
Eumeces indubitus 
Gerrhonotus liocephalus liocephalus 
Phrynosoma orbiculare orbiculare 
Sceloporus aeneus aeneus 



Sceloporus aeneus bicanthalis 
Sceloporus grammicus microlepidotus 
Sceloporus jarrovii sugillatus 
Sceloporus jarrovii minor 
Sceloporus mucronatus mucronatus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 



CHECKLIST OF REPTILES OF MEXICO 



223 



MICHOACAN 



Geoemyda rubida 

Kinosternon hirtipes 

Kinosternon integrum 

Ameiva undulata sinistra 

Anolis nebulosis 

Barisia imbricata imbricata 

Baslliscus vittatus 

Cnemidophorus deppii lineatissimus 

Cnemidophorus guttatiis immutabilis 

Cnemidophorus sackii communis 

Cnemidophorus sackii sackii 

?Coleonyx elegans nemoralis 

Ctenosaura pectinata 

Enyaliosaurus clarki 

Eumeces altamirani 

Eumeces brevirostris 

Eumeces callicephalus 

Eumeces copei 

Eumeces dugesii 

Eumeces indubitus 

Heloderma horridum 



Iguana iguana rhinolopha 

Mabuya mabouya mabouya 

Phrynosoma asio 

Phrynosoma orbiculare orbiculare 

Phyllodactylus lanei 

Sceloporus aeneus aeneus 

Sceloporus asper 

Sceloporus dugesii intermedins 

Sceloporus gadoviae 

Sceloporus grammicus microlepidotus 

Sceloporus horridus oligoporus 

Sceloporus melanorhinus calligaster 

Sceloporus pyrocephalus 

Sceloporus scalaris scalaris 

Sceloporus spinosus spinosus 

Sceloporus torquatus torquatus 

Sceloporus torquatus melanogaster 

Sceloporus utiformis 

Urosaurus bicarinatus bicarinatus 

Urosaurus gadovi 

Crocodylus acutus acutus 



MORELOS 



Kinosternon integrum 

Ameiva undulata sinistra 

Anolis nebulosus 

Barisia imbricata imbricata 

Cnemidophorus guttatus immutabilis 

Cnemidophorus sackii sackii 

Ctenosaura pectinata 

?Enyaliosaurus clarki 

Eumeces copei 

Eumeces indubitus 

Gerrhonotus liocephalus liocephalus 



Heloderma horridum 
Phrynosoma orbiculare orbiculare 
Sceloporus aeneus aeneus 
Sceloporus gadoviae 
Sceloporus grammicus microlepidotua 
Sceloporus horridus horridus 
Sceloporus ochoterenai 
Sceloporus siniferus siniferus 
Sceloporus torquatus torquatus 
Urosaurus bicarinatus bicarinatus 



NAYARIT 



Chelonia mydas 

Geoemyda pulcherrima pulcherrima 

Kinosternon hirtipes 

Kinosternon integrum 

Terrapene nelsoni 

Anolis nebulosus 

Cnemidophorus deppii lineatissimus 

Cnemidophorus sackii communis 

Cnemidophorus sackii mariarum 

Ctenosaura pectinata 

Eumeces callicephalus 

Eumeces parvulus 



Iguana iguana rhinolopha 
Peropus mutilatus 
Phyllodactylus lanei 
Sceloporus asper 
Sceloporus clarkii boulengeri 
Sceloporus dugesii dugesii 
Sceloporus horridus albiventris 
Sceloporus melanorhinus calligaster 
Sceloporus nelsoni 
Sceloporus utiformis 
Urosaurus ornatus schottii 
Crocodylus acutus acutus 



224 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



NUEVO LE6N 



Amyda emoryi 
Gopherus berlandieri 
Pseudemys floridana texana 
Pseudemys scripta elegans 
Barisia imbricata ciliaris 
Cnemidophorus inornatus 
Cnemidophorus sackii gularis 
Coleonyx brevis 
Crotaphytus coUaris baileyi 
Crotaphytus reticulatus 
Eumeces brevilineatus 
Eumeces dicei 
Eumeces obsoletus 
Holbrookia texana 
Scincella caudaequinae 



Phrynosoma cornutum 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
Sceloporus couchii 
Sceloporus cyanogenys 
Sceloporus grammicus disparilis 
Sceloporus jarrovii minor 
Sceloporus olivaceus 
Sceloporus parvus parvus 
Sceloporus poinsettii 
Sceloporus scalaris slevini 
Sceloporus torquatus binocularis 
Sceloporus undulatus consobrinus 
Sceloporus variabilis marmoratus 



OAXACA 



Chelonia mydas 

Dermatemys mawii 

Eretmochelys imbricata 

Geoemyda pulcherrima incisa 

Geoemyda rubida 

Kinosternon cruentatum cruentatum 

Kinosternon integrum 

Lepidochelys olivacea 

Pseudemys scripta ornata 

Pseudemys umbra 

Staurotypus salvinii 

Abronia fuscolabialis 

Abronia oaxacae 

Ameiva undulata amphigramma 

Ameiva undulata dextra 

Ameiva undulata parva 

Ameiva undulata stuarti 

Ameiva undulata undulata 

Anolis heliactin 

Anolis limifrons rodriguezii 

Anolis milleri 

Anolis nebuloides 

Anolis nebulosus 

Anolis sericeus 

Anolis tropidonotus 

Barisia gadovii levigata 

Barisia imbricata imbricata 

Barisia imbricata planifrons 

Barisia viridiflava 

Basiliscus vittatus 

Celestus enneagrammus 

Cnemidophorus deppii deppii 

Cnemidophorus guttatus guttatus 

Cnemidophorus guttatus immutabilis 



Cnemidophorus sackii australis 
Cnemidophorus sackii bocourti 
Coleonyx elegans elegans 
Coleonyx elegans nemoralis 
Corythophanes hernandezii 
Ctenosaura acanthura 
Ctenosaura pectinata 
Ctenosaura similis similis 
Enyaliosaurus quinquecarinatus 
Eumeces brevirostris 
Gaigeia dontomasi 
Gaigeia radula 

Gerrhonotus liocephalus liocephalus 
Gymnophthalmus sumichrasti 
Heloderma horridum 
Iguana iguana rhinolopha 
Laemanctus deborrei 
Laemanctus longipes 
Laemanctus serratus 
Scincella assata taylori 
Scincella cherriei stuarti 
Scincella gemmingeri gemmingeri 
Scincella silvicola 
Lepidophyma smithii tehuanae 
Mabuya mabouya mabouya 
Phrynosoma asio 
Phrynosoma braconnieri 
Phyllodactylus magnus 
Phyllodactylus muralis 
Sceloporus aeneus bicanthalis 
Sceloporus edwardtaylori 
Sceloporus formosus formosus 
Sceloporus gadoviae 
Sceloporus grammicus grammicus 



CHECKLIST OF REPTILES OF MEXICO 



225 



Sceloporus grammicus microlepidotus 
Sceloporus horridus horridus 
Sceloporus jalapae 
Sceloporus malachiticus salvini 
Sceloporus melanorhinus melanorhinus 
Sceloporus mucronatus omiltemanus 
Sceloporus siniferus cupreus 
Sceloporus siniferus siniferus 
Sceloporus spinosus caeruleopunctatus 
Sceloporus teapensis 



Sceloporus variabilis smithi 
Sceloporus variabilis variabilis 
Sphaerodactylus glaucus glaucus 
Sphaerodactylus glaucus torquatus 
Urosaurus bicarinatus anonj^morphus 
Urosaurus bicarinatus nelsoni 
Xenosaurus grandis 
Caiman crocodilus fuscus 
Crocodylus acutus acutus 



PUEBLA 



Kinosternon integrum 

Kinosternon leucostomum 

Abronia taeniata taeniata 

Abronia taeniata graminea 

Ameiva undulata amphigramma 

Ameiva undulata sinistra 

?Anolis cumingii 

Anolis nebuloides 

Anolis nebulosus 

Barisia imbricata imbricata 

Cnemidophorus deppii lineatissimus 

Cnemidophorus sackii sackii 

Cnemidophorus sackii communis 

Ctenosaura pectinata 

Eumeces brevirostris 

Eumeces copei 

Eumeces lynxe furcirostris 

Eumeces lynxe lynxe 

Gerrhonotus liocephalus liocephalus 

Gerrhonotus liocephalus ophiurus 



Iguana iguana rhinolopha 
Phrynosoma braconnieri 
Phrynosoma orbiculare orbiculare 
Phrynosoma orbiculare cortezii 
Phrynosoma taurus 
Sceloporus aeneus aeneus 
Sceloporus aeneus bicanthalis 
Sceloporus formosus formosus 
Sceloporus gadoviae 
Sceloporus grammicus microlepidotus 
Sceloporus jalapae 
Sceloporus megalepidurus 
Sceloporus mucronatus aureolus 
Sceloporus mucronatus mucronatus 
Sceloporus pictus 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 
Urosaurus bicarinatus bicarinatus 



QUERETARO 



Cnemidophorus sackii communis 
Eumeces callicephalus 
Lepidophyma smithii occulor 
Phrynosoma orbiculare orbiculare 



Sceloporus jarrovii immucronatus 
Sceloporus jarrovii minor 
Sceloporus variabilis variabilis 



QUINTANA R00« 



Chelonia mydas 
Eretmochelys imbricata 
Geoemyda areolata 
Kinosternon creaseri 
Kinosternon cruentatum censors 
Lepidochelys kempii 
Pseudemys scripta ornata 



Terrapene mexicana yucatana 
Ameiva undulata gaigeae 
Anolis cozumelae 
Anolis limifrons rodriguezii 
Anolis tropidonotus 
Aristelliger georgeensis 
Basiliscus vittatus 



» Morfln (Informe rendido por la comisiou geogrSflco-exploradora de Quintana Roo al C. Secretario de 
Fomento, 1918, pp. 1-57, pis. 1-10) records, in a previously overlooked paper, a number of snakes from 
Quintana Roo. They are Boa imperator (= Constrictor constrictor imperator) , Bothrops atrix (sic, = B. alrox 
asptr), Crotalus basiliscus (=C. d. durissus), and Flaps fulvius (=Micrurus affinis maj/ensis), all from 
"between Payo Obispo and Champoton." 



226 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



Cnemidophorus deppii cozumelus 
Cnemidophorus sackii angusticeps 
Coleonyx elegans elegans 
Corythophanes hernandezii 
Ctenosaura similis similis 



Iguana iguana rhinolopha 
Mabuya mabouya mabouya 
Sceloporus chrysostictus 
Sceloporus cozumelae 
Crocodylus acutus acutus 



SAN LUIS POTOSI 



Kinosternon hirtipes 
Kinosternon integrum 
Terrapene mexicana mexicana 
Ameiva undulata podarga 
Anolis petersi 
Anolis sericeus 
Anelytropsis papillosus 
Barisia imbricata ciliaris 
Corythophanes hernandezii 
Cnemidophorus sackii communis 
Cnemidophorus sackii gularis 
Crotaphytus collaris baileyi 
Ctenosaura acanthura 
Eumeces lynxe lynxe 
Eumeces tetragrammus 
Gerrhonotus liocephalus infernalis 
Gerrhonotus hocephalus loweryi 
Holbrookia maculata approximans 
Holbrookia maculata dickersonae 
Holbrookia texana 



Laemanctus serratus 
Scincella caudaequinae 
Lepidophyma smithii occulor 
Phrynosoma cornutum 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
Sceloporus cautus 
Sceloporus goldmani 
Sceloporus grammicus disparilis 
Sceloporus jarrovii minor 
Sceloporus jarrovii immucronatus 
Sceloporus olivaceus 
Sceloporus parvus parvus 
Sceloporus parvus scutulatus 
Sceloporus serrifer plioporus 
Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 
Sceloporus variabiUs variabiUs 
Xenosaurus newmanorum 



SINALOA 



Caretta caretta gigas 
Geoemyda pulcherrima pulcherrima 
ffinosternon hirtipes 
Kinosternon integrum 
Pseudemys scripta ornata 
Anolis nebulosus 
Anolis utowanae 
Barisia imbricata ciliaris 
Callisaurus draconoides bogerti 
Callisaurus draconoides brevipes 
Cnemidophorus sackii communis 
Coleonyx fasciatus 
Ctenosaura hemilopha 
Ctenosaura pectinata 
Dipsosaurus dorsalis sonoriensis 
Eumeces humilis 



Eumeces parvulus 
Heloderma horridum 
Holbrookia maculata elegans 
Holbrookia maculata thermophila 
Iguana iguana rhinolopha 
Peropus mutilatus 
Phrynosoma solare 
Phyllodactylus lanei 
Sceloporus clarkii boulengeri 
Sceloporus horridus albiventris 
Sceloporus nelsoni 
Sceloporus utiformis 
?Sphaerodactylus glaucus torquatus 
Urosaurus bicarinatus tuberculatus 
Urosaurus ornatus schottii 



SONORA 



Chelonia mj'das 
Dermochelys coriacea 
Eretmochelys imbricata 
Geoemyda pulcherrima pulcherrima 
Gopherus agassizii 



Kinosternon flavescens stejnegeri 
Kinosternon integrum 
Kinosternon sonoriense 
Lepidochelys olivacea 
Pseudemys scripta hiltoni 



CHECKLIST OF REPTILES OF MEXICO 



227 



Terrapene klauberi 
Terrapene ornata 
Anolis nebuloides 
Callisaurus draconoides brevipes 
Callisaurus draconoides gabbii 
Callisaurus draconoides inusitatus 
Callisaurus draconoides ventralis 
Cnemidophorus bacatus 
Cnemidophorus burti 
Cnemidophorus gadovi 
Cnemidophorus sackii communis 
Cnemidophorus sackii stictogrammus 
Cnemidophorus tigris aethiops 
Cnemidophorus tigris gracilis 
Cnemidophorus tigris martyris 
Cnemidophorus tigris tigris 
Coleonyx variegatus variegatus 
Coleonyx variegatus sonoriensis 
Crotaphytus dickersonae 
Crotaphytus collaris baileyi 
Ctenosaura hemilopha 
Dipsosaurus dorsalis dorsalis 
Dipsosaurus dorsalis sonoriensis 
Elgaria kingii kingii 
Eumeces callicephalus 
Eumeces parviauriculatus 
Gambelia wislizenii wislizenii 
Heloderma horridum 
Heloderma suspectum 



Holbrookia maculata approximans 
Holbrookia maculata thermophila 
Holbrookia texana 
Phrynosoma cornutum 
Phrynosoma ditmarsi 
Phrynosoma douglassii hernandesi 
Phrynosoma m'callii 
Phrynosoma modestum 
Phrynosoma platyrhinos goodei 
Phrynosoma solare 
Phyllodactylus homolepidurus 
Sauromalus obesus townsendi 
Sauromalus varius 
Sceloporus clarkii clarkii 
Sceloporus clarkii boulengeri 
Sceloporus jarrovii jarrovii 
Sceloporus magister magister 
Sceloporus nelsoni 
Sceloporus scalaris slevini 
Sceloporus undulatus virgatus 
Uma notata cowlesi 
Urosaurus bicarinatus tuberculatus 
Urosaurus ornatus linearis 
Urosaurus ornatus schottii 
Urosaurus ornatvis symmetricus 
Uta nolascensis 
Uta palmeri 

Uta stansburiana stejnegeri 
Uta taylori 



TABASCO 



Claudius angustatus 

Dermatemys mawii 

Geoemyda areolata 

Kinosternon acutum 

Kinosternon cruentatum cruentatum 

Kinosternon leucostomum 

Pseudemys scripta ornata 

Staurotypus triporcatus 

Ameiva undulata amphigramma 

Ameiva undulata stuarti 

Anolis capito 

Anolis humilis uniformis 

Anolis laeviventris 

Anolis lemurinus bourgeaei 

Anolis sagrei mayensis 

Anolis sericeus 

Anolis tropidonotus 



Basiliscus vittatus 
Coleonyx elegans elegans 
Ctenosaura similis similis 
Eumeces schwartzei 
Hemidactylus turcicus turcicus 
Iguana iguana rhinolopha 
Laemanctus deborrei 
Scincella cherriei cherriei 
Lepidophyma flavimaculata flavimacu- 

lata 
Mabuya mabouya mabouya 
Sceloporus serrifer plioporus 
Sceloporus teapensis 
Sphaerodactylus glaucus glaucus 
Crocodylus acutus acutus 
Crocodvius moreletii 



228 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



TAMAULIPAS 



Amyda emoryi 
Gopherus berlandieri 
Pseudemys scripta cataspila 
Pseudemys scripta elegans 
Terrapene mexicana mexicana 
Ameiva undulata podarga 
Anolis sericeus 
Basiliscus vittatus 
Cnemidophorus sackii communis 
Cnemidophorus sackii gularis 
Coleonyx brevis 
Crotaphytus reticulatus 
Ctenosaura acanthura 
Eumeces brevilineatus 
Eumeces dicei 



Phrynosoma orbiculare orbiculare 
Sceloporus grammicus microlepidotus 



Eumeces obsoletus 
Eumeces tetragrammus 
Hemidactylus turcicus turcicus 
Holbrookia texana 
Laemanctus serratus 
Phrynosoma cornutum 
Sceloporus cyanogenys 
Sceloporus grammicus disparilis 
Sceloporus olivaceus 
Sceloporus serrifer piioporus 
Sceloporus spinosus spinosus 
Sceleporus variabilis marmoratus 
Sceloporus variabilis variabilis 
Crocodylus acutus acutus 
Crocodylus moreletii 



TLAXCALA 

Sceloporus megalepidurus 



VERACRUZ 



Chelonia mydas 
Chelydra serpentina 
Claudius angustatus 
Dermatemys mawii 
Geomyda areolata 
Kinosternon acutum 
Kinosternon herrerai 
Kinosternon integrum 
Kinosternon leucostomum 
Pseudemys scripta cataspila 
Staurotypus triporcatus 
Terrapene mexicana mexicana 
Abronia taeniata graminea 
Abronia taeniata taeniata 
Ameiva undulata amphigramma 
Anelytropsis papillosus 
Anolis barkeri 
Anolis cymbops 
Anolis laeviventris 
Anolis lemurinus bourgeaei 
Anolis petersii 
Anolis sericeus 
Anolis tropidonotus 
Barisia antauges 
Barisia imbricata imbricata 
Barisia modesta 
Basiliscus vittatus 
Celestus enneagrammus 
Cnemidophorus deppii oligoporus 



Cnemidophorus guttatus guttatus 

Cnemidophorus sackii communis 

Cnemidophorus sackii gularis 

Coleonyx elegans elegans 

Corythophanes cristatus 

Corythophanes hernandezii 

Ctenosaura acanthura 

Ctenosaura similis similis 

Eumeces brevirostris 

Eumeces lynxe furcirostris 

Eumeces lynxe lynxe 

Eumeces sumichrasti 
Eumeces tetragrammus 

Gerrhonotus liocephalus ophiurus 

Hemidactylus mabouia 

Hemidactylus turcicus turcicus 

Holbrookia propinqua piperata 

Iguana iguana rhinolopha 

Laemanctus longipes 

Laemanctus serratus 

Scincella cherriei stuarti 

Scincella gemmingeri gemmingeri 

Scincella silvicola 

Lepidophyma flavimaculata flavimacu- 

lata 
Mabuya mabouya mabouya 
Phrynosoma orbiculare cortezii 
Sceloporus aeneus bicanthalis 
Sceloporus formosus formosus 



CHECKLIST OF REPTILES OF MEXICO 



229 



Sceloporus grammicus microlepidotus 
Sceloporus jalapae 
Sceloporus jarrovii immucronatus 
Sceloporus malachiticus salvini 
Sceloporus megalepidurus 
Sceloporus mucronatus mucronatus 
Sceloporus mucronatus aureolus 
Sceloporus pictus 
Sceloporus serrifer plioporus 



Sceloporus spinosus spinosus 
Sceloporus teapensis 
Sceloporus torquatus torquatus 
Sceloporus variabilis variabilis 
Sphaerodactylus glaucus glaucus 
Xenosaurus grandis 
Crocodylus acutus acutus 
Crocodylus moreletii 



YUCATAN 



Caretta caretta caretta 
Chelonia mydas 
Dermatemys mawii 
Geoemyda areolata 
Kinosternon creaseri 
Kinosternon cruentatum consors 
Pseudemys scripta ornata 
Terrapene mexicana yucatana 
Ameiva undulata gaigeae 
Anolis beckeri 
Anolis kidderi 
Anolis lemurinus bourgeaei 
Anolis limifrons rodriguezii 
Anolis sagrei mayensis 
Anolis tropidonotus 
Anolis ustus 
Basiliscus vittatus 
Cnemidophorus sackii angusticeps 



Coleonyx elegans elegans 
Corythophanes cristatus 
Corythophanes hernandezii 
Ctenosaura similis sirailLs 
Enyaliosaurus defensor 
Eumeces schwartzei 
Hemidactylus turcicus turcicus 
Laemanctus alticoronatus 
Laemanctus serratus 
Scincella cherriei ixbaac 
Mabuya mabouya mabouya 
Sceloporus chrysostictus 
Sceloporus cozumelae 
Sceloporus lundelli gaigeae 
Sceloporus serrifer serrifer 
Sphaerodactylus glaucus glaucus 
Thecadactylus rapicaudus 



ZACATECAS 



Barisia imbricata ciliaris 
Cnemidophorus sackii communis 
Eumeces callicephalus 
Holbrookia macula ta dickersonae 
Phrynosoma modestum 
Phrynosoma orbiculare orbiculare 
Sceloporus cautus 



Sceloporus grammicus disparilis 
Sceloporus jarrovii minor 
Sceloporus scalaris scalaris 
Sceloporus spinosus spinosus 
Sceloporus torquatus melanogaster 
Sceloporus undulatus consobrinus 



INDEX 



abaxillare, Kinosternon, 22, 218. 
abbotti, Coleonyx variegatus, 42, 44, 

217. 
abnormis, Dermatemys, 19. 
Abronia, 194, 196. 

deppii, 196, 197, 221. 

fuscolabialis, 196, 198, 224. 

matudai, 196, 218. 

oaxacae, 196, 197, 224. 

ochoterenai, 196, 197, 218. 

taeniata graminea, 196, 198, 225, 
228. 

taeniata taeniata, 196, 197, 222, 
225, 228. 

taeniatus, 197. 
acanthinus, Sceloporus, 108. 

Sceloporus malachiticus, 107, 108, 
218. 
acanthura, Ctenosaura, 73, 74, 224, 226, 
228. 

Lacerta, 74. 
acuta, Kinosternon scorpioides, 23. 
acutirostris, Anolis, 64. 

Testudo, 20. 
acutum, Kinosternon, 22, 23, 227, 228. 
acutus, Crocodylus, 210. 

Crocodvlus acutus, 210, 218, 220, 
221, 223, 225, 226, 227, 228, 229. 
adspersus, Gerrhonotus, 201. 
aeneus, Sceloporus, 137. 

Sceloporus aeneus, 136, 137, 220, 
221, 222, 223, 225. 
aethiops, Cnemidophorus tessellatus, 
189. 

Cnemidophorus tigris, 177, 189, 
227. 
Agama collaris, 91. 

cornuta, 99. 

(Phrynosoma) coronata, 102. 

cristata, 68, 69. 

orbiculaire, 97. 

torquata, 126. 

undulata, 105. 
agassizii, Chelonia, 17. 

Gopherus, 28, 217, 226. 

Testudo, 28. 

Xerobates, 28. 
agilis, Scincus, 156. 
Agkistrodon, 3. 
Akleistops, 151. 

guatemalensis, 151, 152. 
albiventris, Sceloporus horridus, 110, 

116, 222, 226. 
albogularis, Gonatodes, 45. 

Gymnodactylus, 45. 



alfaronis, Cnemidophorus, 178. 
alliacea, Mabuia, 156. 
Alligator chiapasius, 212. 

lacordairei, 211. 

mississippiensis, 211. 
Alligatoridae, 210, 211. 
alpha, Sceloporus grammicus, 120. 
altamirani, Eumeces, 160, 161, 163, 223. 
alticoronatus, Laemanctus, 70, 229. 
amblygrammus, Eumeces skiltonianus, 

167. 
Ambystomidae, 2. 
Ameiva, 170. 

ameiva, 170. 

edwardsii, 174. 

festiva edwardsii, 171, 174, 218. 

tesselata, 188. 

undulata amphigramma, 171, 172, 

224, 225, 227, 228. 

undulata dextra, 171, 173, 221, 224. 
undulata gaigeae, 171, 172, 218, 

225, 228. 

undulata hartwegi, 171, 218. 

undulata parva, 171, 173, 218, 224. 

undulata podarga, 171, 172, 226, 
228. 

undulata sinistra, 171, 174, 220, 
222, 223, 225. 

undulata stuarti, 171, 173, 218, 224, 
227. 

undulata thomasi, 171, 173, 218. 

undulata undulata, 171, 174, 224. 
ameiva, Ameiva, 170. 
americana, Lacerta, 170. 
americanus, Basiliscus, 71. 

Crocodylus, 210, 211. 
Amoebopsis, 154. 
amphigramma, Ameiva undulata, 171, 

172, 224, 225, 227, 228. 
Amphisbaenia, 2, 12, 37. 
Amyda, 18. 

cartilaginea, 18. 

emoryi, 18, 217, 219, 224, 228. 

javanica, 18. 

spinifera, 18. 
Anapsida, 12. 
Ancistrodon, 3. 
Anelytropsidae, 39, 170. 
Anelytropsis, 170. 

papillosus, 170, 226, 228. 
Aneuporus occipitalis, 213. 
Anguidae, 39, 40, 194. 
Anguis, 194. 

angustatus, Claudius, 26, 218, 227, 228. 
angusticeps, Cnemidophorus, 183. 

231 



232 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



angusticeps, Cnemidophorus sackii, 178, 

183, 218, 226, 229. 
angustus, Sator, 139, 217. 
Aniellidae, 208. 

annectens, Ctenosaura (Cachryx), 77. 
Anniella, 208, 209. 

geronimensis, 209, 217. 

pulchra, 209. 

pulchra pulchra, 209, 217. 

texana, 209. 
Anniellidae, 39, 208. 
annulata, Geoclemmys, 29. 
annulatus, Sceloporus merriami, 132, 

133, 219. 
Anolis, 4, 54, 55. 

acutirostris, 64. 

aureolus, 64. 

baccatus, 57, 62. 

barkeri, 56, 58, 228. 

beckeri, 57, 62, 229. 

biporcatus, 58, 65, 67, 218. 

boulengerianus, 65. 

bourgeaei, 66, 67. 

bullaris, 56. 

capito, 58, 65, 227. 

carneus, 65. 

carolinensis, 55, 56. 

cobanensis, 63. 

copei, 65. 

cozumelae, 56, 59, 225. 

cumingii, 58, 67, 225. 

cymbops, 57, 58, 62, 228. 

damulus, 57, 61. 

dunni, 56, 61, 221. 

fuscoauratus, 55. 

gadovii, 56, 61, 221. 

guentherii, 64. 

guntherii, 57, 63. 

heliactin, 58, 67, 224. 

humilis uniformis, 56, 60, 218, 227. 

impetigosus, 56, 60. 

intermedius, 63. 

jacobi, 68. 

kidderi, 58, 66, 218, 229. 

laeviventris, 9, 57, 58, 62, 63, 227, 228. 

lemurinus bourgeaei, 58, 66, 218, 
227, 228, 229. 

limifrons rodriguezii, 56, 57, 64, 
218, 224, 225, 229. 

liogaster, 57, 62, 221. 

megapholidotus, 56, 59, 221. 

metallicus, 56, 59. 

miUeri, 56, 57, 64, 224. 

nannodes, 63. 

nebuloides, 58, 66, 219, 220, 221, 
224, 225, 227. 

nebulosus, 58, 65, 220, 221, 222, 
223, 224, 225, 226. 

panamensis, 61. 

pentaprion 55, 57, 61, 218. 

(Coccoessus) pentaprion, 61. 

petersii, 58, 65, 226, 228. 

petersii bivittatus, 65. 

rodriguezii, 64. 

rubigenosus, 64. 

ruthveni, 60. 



Anolis sagraei, 55. 

sagrei mayensis, 56, 59, 218, 227, 
229. 

sallaei, 67. 

schiedii, 56, 57, 58, 63, 64. 

schmidti, 56, 58, 60, 220. 

sericeus, 58, 67, 218, 224, 226, 227. 
228. 

stuarti, 63. 

sulcifrons, 61. 

taylori, 57, 61, 221. 

tropidonotus, 56, 59, 60, 218, 224, 
225, 227, 228, 229. 

uniformis, 59, 60. 

ustus, 58, 66, 218, 229. 

ustus veraepacis, 67. 

utowanae, 57, 62, 226. 

viridis, 55. 

wiegmanni, 63. 
anonymorpha, Uta, 146. 

Uta bicarinata, 146. 
anonymorphus, Urosaurus bicarinatus, 

141, 146, 218, 221, 225. 
Anota, 95. 

calidiarum, 101. 

goodei, 102. 

m'callii, 95, 99. 

naodesta, 101. 

platyrhina, 101. 
antauges', Barisia, 199, 200, 228. 

Gerrhonotus, 200. 
anthracinus, Sphaerodactylus, 52, 213. 
approximans, Holbrookia, 83. 

Holbrookia maculata, 82, 83, 219, 

220, 222, 226, 227. 
Archosauria, 12, 209. 
areolata, Emvs, 30. 

Geoemvda, 29, 30, 218, 225, 227, 
228, 229. 
Aristelliger, 40, 41, 51. 

georgeensis, 51, 225. 

irregularis, 51. 

lar, 51. 

praesignis, 51. 
arizonae, Cnemidophorus, 184. 
armata. Iguana (Ctenosaura), 74. 
articulata, Cyclura, 74. 
asio, Batrachosoma, 102. 

Phrynosoma, 95, 102, 218, 220, 

221, 223, 224. 

asper, Sceloporus, 107, 109, 221, 222, 

223. 
Aspidonectes emoryi, 18. 
Aspidoscelis, 174. 

assata, Scincella assata, 157, 160, 218. 
assatum, Leiolopisma assatum, 160. 

Lygosoma assatum, 160. 
assatus, Lampropholis, 160. 
ater, Sauromalus, 79, 80, 217. 
Atheca, 12, 13. 
atitlanensis, Celestus, 195. 
atrix, Bothrops, 225. 
aureolus, Anolis, 64. 

Sceloporus mucronatus, 123, 124, 
225, 229. 
auriculata, Uta, 146. 



INDEX 



233 



auriculatus, Urosaurus, 140, 146, 220. 
australis, Cnemidophorus communis, 
181. 
Cnemidophorus sackii, 178, 181, 224. 
Sauromalus, 79, 80, 217. 
austrinus, Gerrhonotus liocephalus, 203, 
204, 218. 

bacatus, Cnemidophorus, 176, 187, 227. 
baccatus, Anolis, 57, 62. 
Bachia dorbignyi, 213. 

heteropa, 212. 
baileyi, Crotaphytus, 92. 

Crotaphvtus collaris, 92, 217, 219, 
220, 224, 226, 227. 
bakeri, Ctenosaura, 73. 
balsas, Cnemidophorus mexicanus, 181. 
Barisia, 194, 198. 

antauges, 199, 200, 228. 

gadovii gadovii, 199, 200, 221. 

gadovii levigata, 199, 200, 224. 

imbricata ciliaris, 199, 202, 219, 
220, 221, 222, 224, 226, 229. 

imbricata imbricata, 198, 199, 201, 

220, 221, 222, 223, 224, 225, 228. 
imbricata planifrons, 199, 201, 224. 
levicoUis, 199, 202, 219. 
modesta, 199, 201, 228. 
moreleti rafaeli, 198, 199, 218. 
moreleti temporalis, 199, 218. 
rudicollis, 199, 202, 222. 
viridiflava, 198, 200, 224. 

barkeri, Anolis, 56, 58, 228. 
bartolomas, Cnemidophorus, 191. 
Basiliscus, 54, 71. 

americanus, 71. 

basiliscus, 71. 

(Cristasaura) nuchalis, 72. 

vittatus, 71, 218, 220, 221, 222, 223, 
224, 225, 227, 228, 229. 
basiliscus, Basiliscus, 71. 

Crotalus, 225. 
Batrachosoma, 94. 

asio, 102. 
beckeri, Anolis, 57, 62, 229. 
beldingi, Cnemidophorus hyperythrus, 
175, 186, 217. 

Verticaria, 186. 

Verticaria hyperythra, 186. 
belli. Leiosaurus, 212. 
bellii, Chrysemys picta, 34, 219. 

Emys, 34. " 

Iguana (Ctenosaura), 74. 

Plestiodon, 163. 
berardii, Emys, 19. 
berendtianum, Cinosternum, 23. 
berlandieri, Gopherus, 28, 219, 224. 

Testudo, 28. 

Xerobates, 28. 
bicanthalis, Sceloporus aeneus, 136, 137, 

222, 224, 225, 228. 
bicarinata, Uta, 146. 
bi-carinata, Uta bi-carinata, 146. 
bicarinatus, Phymatolepis, 140. 

Urosaurus bicarinatus, 141, 146, 

221, 223, 225. 



bi-carinatus, Phymatolepis, 146. 

Bimanus, 38. 

binocularis, Sceloporus, 126. 

Sceloporus ferrariperezi, 126. 

Sceloporus torquatus, 122, 126, 224. 
Bipedidae, 37. 
Bipes, 37, 38. 

biporus, 38, 217. 

canaliculatus, 38, 221. 

tridactylus, 38, 221. 
biporcata, Dactyloa, 55, 65. 
biporcatus, Anolis, 58, 65, 67, 218. 

Staurotypus, 27. 
biporus, Bipes, 38, 217. 

Euchirotes, 38. 
bi-seriatus, Sceloporus, 118. 
biseriatus, Sceloporus occidentalis, 117, 

118, 217. 
bischoffi, Holbrookia, 84. 
biscutatus, Crocodilus, 211. 
bissa, Caretta, 16. 
bivittata, Daconura, 71, 72. 
bivittatus, Anolis petersii, 65. 
blainvillii, Phrynosoma, 103. 

Phrynosoma coronatum, 97, 103, 217. 
Blepharactisis, 192. 

speciosa, 192. 
Boa imperator, 225. 
bocourti, Cnemidophorous, 181. 

Cnemidophorus communis, 181. 

Cnemidophorus sackii, 178, 181, 
218, 224. 

Gerrhonotus, 200. 

Tropidurus, 213. 
bocourtii, Eumeces, 166. 
bogerti, Callisaurus draconoides, 86, 
89, 226. 

Coleonvx variegatus, 42, 44. 
bordai, Phyllodactylus, 46, 47, 221. 
Bothrops atrix, 225. 
boucardi, Phrynosoma, 96, 98, 222. 
boucardii, Phrynosoma, 98. 

Tapaya, 98. 
boulengeri, Sceloporus, 113. 

Sceloporus clarkii. 111, 113, 222, 
223, 226, 227. 
boulengerianus, Anolis, 65. 
bourgeaei, Anolis, 66, 67. 

Anolis lemurinus, 58, 66, 218, 227, 
228, 229. 
brachycercum, Phrynosoma douglassii, 

96, 100, 220. 
Brachydactylus, 41. 

mitratus, 41. 
brachylopha, Ctenosaura teres, 76. 
braconnieri, Goniodactylus, 45. 

Phrynosoma, 95, 100, 224, 225. 
brevigulare, Cinosternum, 25. 
brevilineatus, Eumeces, 162, 164,224,228. 
brevipes, Callisaurus draconoides, 86, 

88, 226, 227. 
brevirostris, Ctenosaura, 75. 

Eumeces, 162, 168, 220, 221, 222, 
223, 224, 225, 228. 

Mabouia, 168. 
brevis, Coleonyx, 42, 43, 219, 220, 224, 228. 



234 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



bufoniurn, Phrynosoma, 99. 
buUaris, Anolis, 56. 

Dactyloa, 55. 
bulleri, Sceloporus, 122, 125, 222. 
bunkerL Holbrookia, 82, 84, 219. 
burti, (Jnemidopliorus, 177, 185, 227. 

Cachryx, 76, 77. 

defensor, 77. 

erythromelas, 77. 
caerulea, Uta, 143. 

Verticaria, 186. 
caeruleopunctatus, Sceloporus spinosus, 

110, 116, 225. 
caeruleus, Cnemidophorus hyperythrus, 
175, 186, 217. 

Sceloporus ornatus, 122, 127, 219. 

Urosaurus ornatus, 141, 143, 219. 
Caiman, 212. 

crocodilus fuscus, 212, 218. 

fissipes, 212. 
calidiarum, Anota, 101. 

Phrynosoma, 101. 
callicephalus, Eumeces, 161, 164, 219, 

220, 221, 222, 223, 225, 227, 229. 
calligaster, Sceloporus melanorhinus, 

110, 113, 220, 221, 222, 223. 
callirostris, Emys, 32. 
Callisaurus, 55, 85. 

carmenensis, 87. 

crinitus, 87. 

draconoides, 85, 86. 

draconoides bogerti, 86, 89, 226. 

draconoides brevipes, 86, 88, 226, 
227. 

draconoides carmenensis, 86, 87, 
217. 

draconoides crinitus, 86, 87, 217. 

draconoides draconoides, 86, 217. 

draconoides gabbii, 86, 87, 217, 227. 

draconoides inusitatus, 86, 88, 227. 

draconoides splendidus, 86, 87, 217. 

draconoides ventralis, 86, 88, 227. 

inusitatus, 88. 

rhodostictus, 87. 

splendidus, 87. 

ventralis, 88. 

ventralis gabbii, 87. 

ventralis inusitatus, 88. 

ventralis ventralis, 88. 
canaliculatus, Bipes, 38, 221. 

Chirotes, 39. 
canus, Cnemidophorus, 190. 

Cnemidophorus tessellatus, 190. 

Cnemidophorus tigris, 176, 190,217. 
Caouana, 14, 15. 
caouana, Testudo, 15, 16. 
capito, Anolis, 58, 65, 227. 
Caretta, 14, 15. 

bissa, 16. 

caretta, 16. 

caretta caretta, 15, 16, 229. 

caretta gigas, 15, 16, 217, 226. 

gigas, 16. 



Caretta nasuta, 15. 

remivaga, 15. 
caretta, Caretta, 16. 

Caretta caretta, 15, 16, 229. 

Testudo, 16. 
Carettochelyidae, 18. 
Carettoidea, 12, 14. 
carinatus, Sceloporus, 134, 135, 218. 
carmenensis, Callisaurus, 87. 

Callisaurus draconoides, 86, 87, 217. 

Dipsosaurus, 78, 217. 
carneus, Anolis, 65. 
Carolina, Cistudo, 34. 

Terrapene, 34. 
carolinensis, Anolis, 55, 56. 
cartilaginea, Amyda, 18. 
caspica, Clemmys caspica, 33. 
catalinensis, Cnemidophorus, 176, 188, 
217. 

Dipsosaurus, 78, 217. 
cataspila, Emys, 32. 

Pseudemys scripta, 31, 32, 228. 
caudaequinae, Leiolopisma, 158. 

Scincella, 157, 158, 224, 226. 
Caudata, 2. 

cautus, Sceloporus, 117, 219, 226, 229. 
cedrosensis, Elgaria, 205, 206, 217. 

Gerrhonotus, 206. 
celeripes, Cnemidophorus, 190. 

Cnemidophorus tessellatus, 190. 

Cnemidophorus tigris, 176, 190, 217. 
Celestus, 194. 

atitlanensis, 195. 

enneagrammus, 195, 224, 228. 

rozellae, 195, 218. 

striatus, 194. 
cephalo, Testudo, 16. 
ceralbensis, Verticaria, 188. 

Cnemidophorus, 175, 188, 217. 
cerroense, Phrynosoma, 97, 103, 217. 
Chalcides heteropus, 212. 

sulcata, 39. 
Chalinocnemis, 51. 
chalybaeus, Diploglossus, 196. 
Chamaeleo mexicanus, 68. 
Chamaeleopsis, 68. 

hernandezii, 68. 
chamaeleopsis, Corythophanes, 68. 
Chamaesaura propus, 39. 
Chehdridae, 20. 
Cheliurus, 20. 
Chelona, 17. 
Chelone, 17. 
Chelonia, 14, 17. 

agassizii, 17. 

dussumieri, 15. 

lachrymata, 16. 

maculosa, 17. 

mydas, 17, 221, 223, 224, 225, 226, 
228, 229. 

olivacea, 14, 15. 

pseudo-caretta, 16. 

pseudo-mydas, 16. 

virgata, 17. 



INDEX 



235 



Chelonias, 17. 
Cheloniidae, 14. 
Chelonura, 20. 
Chelopus rubida, 30. 
Chelydra, 20. 

rossignonii, 20. 

serpentina, 20, 21, 218, 228. 
Chelydridae 19, 20. 
cherriei, Leiolopisma, 158. 

Leiolopisma cherriei, 157. 

Lygosoma assatum, 157. 

Lygosoma cherriei, 157. 

Mocoa, 157. 

Scincella cherriei, 157, 218, 227. 
chiapasius, Alligator, 212. 
chiricahuae, Urosaurus ornatus, 142, 
143. 

Uta ornata, 143. 
Chirotes canaliculatus, 39. 
Chloremys, 19. 
Chrysemys, 29, 33. 

nebulosa, 32. 

picta bellii, 34, 219. 

scripta elegans, 32. 
chrvsostictus, Sceloporus, 105, 133, 218, 

226, 229. 
ciliaris, Barisia imbricata, 199, 202, 219, 
220, 221, 222, 224, 226, 229. 

Gerrhonotus levicoUis, 202. 
cinereus, Sphaerodactylus, 52. 
Cinosternidae, 21. 
Cinosternon, 3. 

cruentatum, 23. 

effeldtii, 23. 

hirtipes, 25. 

leucostomum, 25. 

punctatum, 26. 

shawianum, 23. 
Cinosternum berendtianum, 23. 

brevigulare, 25. 

cobanum, 25. 

henrici, 25. 

hirtipes, 25. 

integrum, 25. 

rostellum, 25. 

scorpioides integrum, 25. 

sonoriense, 26. 

triliratum, 23. 
Cistudo Carolina, 34. 

mexicana, 34. 

(Onychotria) mexicana, 34. 

ornata, 36. 

yucatana. 35. 
clarionensis, Urosaurus, 141, 142, 220. 

Uta, 142. 
clarki, Ctenosaura, 76. 

Enyaliosaurus, 76, 223. 
clarkii, Sceloporus, 113. 

Sceloporus clarkii, 111, 113, 219, 
227. 
Claudius, 21, 26. 

angustatus, 26, 218, 227, 228. 

megalocephalus, 26. 

pictus, 27. 

severus, 27. 
clausa, Testudo, 34. 



Clemmys, 29, 37. 

caspica caspica, 33. 
guttata, 37. 

marmorata pallida, 37, 217. 
Cnemaspis, 45. 

Cnemidophorus, 1, 4, 170, 174. 
alfaronis, 178. 
angusticeps, 183. 
arizonae, 184. 
bacatus, 176, 187, 227, 
bartolomas, 191. 
bocourti, 181. 
burti, 177, 185, 227. 
canus, 190. 

catalinensis, 176, 188, 217. 
celeripes, 190. 
ceralbensis, 175, 188, 217. 
communis, 182. 
communis australis, 181. 
communis bocourti, 181. 
communis copei, 181, 182. 
communis occidentalis, 182, 
costatus, 181. 
decemlineatus, 178. 
deppei cozumela, 179. 
deppii, 178. 

deppii cozumelus, 176, 179, 226. 
deppii deppii, 176, 178, 218, 221, 

224. 
deppii lineatissimus 176, 179, 220, 

221, 222, 223, 225. 
deppii oligoporus, 176, 179, 218, 

228. 
dickersonae, 191. 
disparilis, 189. 
estebanensis, 189. 
gadovi, 178, 185, 227. 
gracilis, 189. 
grahamii, 188. 
gularis, 183. 
gularis gularis, 183. 
gularis gularis obsoletus, 182. 
gularis gularis verus, 182. 
gularis meeki, 184. 
gularis octolineatus, 183. 
gularis scalaris, 182. 
gularis sealous, 183. 
gularis semifasciatus, 184. 
gularis sericeus, 183. 
gularis velox, 184. 
guttatus, 183. 
guttatus guttatus, 175, 179, 224, 

228. 
guttatus immutabilis, 175, 180, 218, 

220, 221, 223, 224. 
guttatus striatus, 180. 
hyperythrus, 174. 
hyperythrus beldingi, 175, 186, 217. 
hyperythrus caeruleus, 175, 186, 

217. 
hyperythrus danheimae, 175, 186, 

217. 
hyperythrus hyperythrus, 175, 187, 

217. 
hyperythrus pictus, 175, 186, 217. 



236 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



Cnemidophorus hyperythrus schmidti, 

175, 187, 217. 
immutabilis, 180. 
inornatus, 177, 184, 219, 224. 
labialis, 178, 185, 217. 
lativittis, 178. 
lineatissimus, 179. 
mariarum, 183. 
marmoratus, 190. 
martyris, 191. 
maximus, 176, 188, 217. 
melanostethus, 189, 190. 
mexicanus, 181. 
mexicanus balsas, 181. 
mexicanus typica, 181. 
microlepidopus, 180. 
motaguae, 181. 
octolineatus, 184. 
perplexus, 184, 185. 
punctilineatus, 189. 
rubidus, 191. 
sackii, 181. 
sackii angusticeps, 178, 183, 218, 

226, 229. 
sackii australis, 178, 181, 224. 
sackii bocourti, 178, 181, 218, 224, 
sackii communis, 178, 182, 217, 219, 

220, 221, 222, 223, 225, 226, 227, 

228, 229. 
sackii gularis, 178, 183, 224, 226, 

228. 
sackii mariarum, 178, 183, 223. 
sackii sackii, 178, 180, 221, 223, 225. 
sackii scalaris, 178, 182, 219, 220. 
sackii semifasciatus, 178, 184, 219. 
sackii stictogrammus, 178, 183, 219, 

227. 
septemvittatus, 184. 
sexlineatus, 177, 185. 
sexlineatus sackii, 180, 181. 
sexlineatus tigris, 182. 
stejnegeri, 191. 
tesselatus, 3, 176, 188. 
tessellatus, 3. 
tessellatus aethiops, 189. 
tessellatus canus, 190. 
tessellatus celeripes, 190. 
tessellatus martyris, 191. 
tessellatus multiscutatus, 191. 
tessellatus rubidus, 191. 
tessellatus stejnegeri, 191. 
tessellatus tessellatus, 189. 
tigris aethiops, 177, 189, 227. 
tigris canus, 176, 190, 217. 
tigris celeripes, 176, 190, 217. 
tigris gracilis, 177, 189, 227. 
tigris marmoratus, 177, 190, 219, 

220. 
tigris martyris, 176, 191, 227. 
tigris multiscutatus, 177, 191, 217. 
tigris rubidus, 176, 191. 
tigris tigris, 177, 189, 217, 227. 
undulatus, 174. 
unicolor, 180. 
Cnemidophorus vandenburghi, 191. 
variolosus, 190. 



coahuila, Terrapene, 34, 36, 219. 
cobanensis, Anolis, 63. 
cobanum, Cinosternum, 25. 
Coccoessus, 55, 61. 
cochranae, Sceloporus, 134. 
Coleonyx, 41, 42. 

brevis, 42, 43, 219, 220, 224, 228. 

elegans, 41, 42, 43. 

elegans elegans, 42, 218, 224, 226, 
227, 228, 229. 

elegans nemoralis, 42, 43, 220, 221, 
223, 224. 

fasciatus, 42, 43, 220, 226, 

variegatus, 44. 

variegatus abbotti, 42, 44, 217. 

variegatus bogerti, 42, 44. 

variegatus peninsularis, 42, 44, 217. 

variegatus slevini, 42, 45, 217. 

variegatus sonoriensis, 42, 45, 227. 

variegatus variegatus, 42, 44, 217, 
227. 
coleonyx, Gymnodactylus, 42. 
colimensis, Eumeces, 162, 169, 220. 
collaris, Agama, 91. 

Crotaphytus, 92. 

Crotaphytus collaris, 92. 
Colpochelys, 15. 
communis, Cnemidophorus, 182. 

Cnemidophorus sackii, 178, 182, 
217, 219, 220. 221, 222, 223, 
225, 226, 227, 228, 229. 
completa, Ctenosaura, 73. 
concentrica, Testudo, 36. 
concinna, Testudo, 31. 

Uta, 148, 150, 217. 
consobrinus, Sceloporus, 118. 

Sceloporus undulatis, 117, 118, 219, 
220. 224, 229. 
consors, Kinosternon cruentatum, 22, 

24, 225, 229. 
conspicuosa, Ctenosaura, 75. 
Constrictor constrictor imperator, 225. 

durissus durissus, 225. 
copei. Anolis, 65. 

Cnemidophorus communis, 181, 182. 

Eumeces, 162, 166, 220, 222, 223, 
225. 
copeii, Crotaphytus, 94. 
Cophosaurus, 81. 

texana, 81. 

texanus, 85. 
Cordylus (Gerrhonotus) multicarinatus, 

205. 
coriacea, Dermochelys, 13, 217, 226. 

Testudo, 13. 
cornuta, Agama, 99. 

Tapaya, 99. 
cornutum, Phrvnosoma, 94, 95, 99, 219, 

220, 224, 226, 227. 
coronata, Agama (Phrynosoma), 102. 
coronatum, Phrynosoma, 94, 96. 

Phrvnosoma coronatum, 97, 102, 
2i7. 
cortezii, Phrvnosoma orbiculare, 96, 98, 
225, 228. 

Tapaya, 98. 



INDEX 



237 



Corythaeolus, 71. 
Corythophanes, 3, 54, 68. 

chamaeleopsis, 68. 

cristatus, 68, 69, 218, 228, 229. 

hernandezii, 68, 218, 224, 226, 228, 
229. 

mexicanus, 68. 

percarinatus, 68, 69, 218. 
Corytophanes, 3, 68, 69. 
costatus, Cnemidophorus, 181. 
couchii, Sceloporus, 105, 129, 132, 219, 

224. 
cowlesi, Uma notata, 89, 90, 227. 
cozumela, Cnemidophorus deppei, 179. 
cozumelae, Anolis, 56, 59, 225. 

Sceloporus, 129, 130, 226, 229. 
cozumelus, Cnemidophorus deppii, 176, 

179, 226. 
creaseri, Kinosternon, 22, 23, 225, 229. 
Cricosaura, 151. 
crinitus, Callisaurus, 87. 

Callisaurus draconoides, 86, 87, 217. 
Cristasaura, 71, 72. 

mitrella, 71. 
cristata, Agama, 68, 69. 
cristatus, Corythophanes, 68, 69, 218, 

228, 229. 
Crocodilidae, 210. 
Crocodilus biscutatus, 211. 

latirostris, 212. 

mexicanus, 211. 

pacificus, 211. 
Crocodylidae, 210. 
Crocodylus, 210. 

acutus, 210. 

acutus acutus, 210, 218, 220, 221, 

223, 225, 226, 227, 228, 229. 
americanus, 210, 211. 

moreletii, 210, 211, 218, 220, 227, 
228, 229. 

niloticus, 210. 
Crotalus basiliscus, 225. 

polystictus, 6. 
Crotaphytus, 55, 91. 

baileyi, 92. 

coll&ris 92. 

collaris' baileyi, 92, 217, 219, 220, 

224, 226, 227. 
collaris collaris, 92. 
collaris dickersonae, 93. 
copeii, 94. 

dickersonae, 92, 93, 227. 
dorsalis, 77, 78. 
fasciatus, 94. 
fasciolatus, 94. 
gambelii, 94. 
insularis, 92, 93, 217. 
reticulatus, 92, 93, 219, 224, 228. 
wislizenii, 94. 

cruentatum, Cinosternon, 23. 

Kinosternon cruentatum, 22, 23, 
218, 224, 227. 
Cryptoblepharus, 155. 
Cryptodira, 12, 18, 19. 
Ctenocercus, 55. 



Ctenosaura, 54, 73, 74. 

acanthura, 73, 74, 224, 226, 228. 

(Cachryx) annectens, 77. 

bakeri, 73. 

brevirostris, 75. 

clarki, 76. 

completa, 73. 

conspicuosa, 75. 

cycluroides, 73, 74. 

defensor, 77. 

erythromelas, 76, 77. 

hemilopha, 73, 75, 217, 219, 226, 
227. 

insulana, 75. 

interrupta, 75. 

multispinis, 74. 

D3,rkori 7^ 

pectina'ta, 73, 75, 220, 221, 222, 
223, 224, 225, 226. 

similis, 73, 218. 

similis similis, 218, 224, 226, 227, 
228, 229. 

teres brachylopha, 75. 
Cubina grandis, 208. 
cumingii, Anolis, 58, 67, 225. 
cupreus, Sceloporus, 134. 

Sceloporus siniferus, 134, 225. 
cyanogenys, Sceloporus, 123, 125, 224, 
228. 

Sceloporus torquatus, 125. 
Cyclura articulata, 74. 

denticulata, 74. 

palearis, 76. 

pectinata, 75. 

quinquecarinata, 76. 

semicristata, 74. 

shawii, 74. 

(Ctenosaura) similis, 73. 

teres, 74. 
cycluroides, Ctenosaura, 73, 74. 
cymbops, Anolis, 56, 57, 58, 62, 228. 

Daconura bivittata, 71, 72. 
Dactyloa, 55. 

biporcata, 55, 65. 

bullaris, 55. 

(Anolis) laeviventris, 62. 

nebulosa, 55, 65. 

schiedii, 63. 
Dactyloperus, 51. 
damulus, Anolis, 57, 61. 
danheimae, Cnemidophorus hyperyth- 

rus, 175, 186, 217. 
Dasia microcephalus, 214. 
deborrei, Laemanctus, 70, 71, 224, 227. 
decemlineatus, Cnemidophorus, 178. 
defensor, Cachryx, 77. 

Ctenosaura, 77. 

Enyaliosaurus, 76, 77, 229. 
delcampi, Phyllodactylus, 46, 47, 221. 
delicatissima, Iguana, 72. 
delicatissimus, Sceloporus, 131. 



861316—50- 



238 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



denticulata, Cyclura, 74. 
deppii, Abronia, 196, 197, 221. 

Cnemidophorus, 178. 

Cnemidophorus deppii, 176, 178, 

218, 221, 224. 
Gerrhonotus, 196, 197. 

Dermatemyidae, 19. 
Dermatemys, 19. 

abnormis, 19. 

mawii, 19, 218, 224, 227, 228, 229. 
Dermochelyidae, 13. 
Dermochelys, 13. 

coriacea, 13, 217, 226. 

schlegelii, 13. 
Devisia, 20. 
dextra, Ameiva undulata, 171, 173, 221, 

224. 
dicei, Eumeces, 162, 169, 224, 228. 
dickersonae, Cnemidophorus, 191. 

Crotaphytus, 92, 93, 227. 

Crotaphytus collaris, 93. 

Holbrookia, 83. 

Holbrookia maculata, 82, 83, 219, 
220, 221, 226, 229. 
digueti, Gerrhonotus deppii, 198. 

Sceloporus, 114. 
Diplodactylus, 47. 

unctus, 47. 
Diploglossus, 39. 

chalybaeus, 196. 

microcephalus, 214. 

monotropis, 214. 

steindachneri, 195. 
Dipsosaurus, 54, 77. 

carmenensis, 78, 217. 

catalinensis, 78, 217. 

dorsalis, 78. 

dorsalis dorsalis, 77, 78, 217, 227. 

dorsalis lucasensis, 78, 217. 

dorsalis sonoriensis, 77, 79, 226, 227. 
Discodactylus, 46. 
dispar, Sceloporus, 120. 
disparilis, Cnemidophorus, 189. 

Sceloporus, 120. 

Sceloporus grammicus, 119, 120, 

219, 220, 221, 222, 224, 226, 
228, 229. 

Sceloporus microlepidotus, 120. 

ditmarsi, Phrynosoma, 95, 104, 227. 
Doliosaurus, 95. 

m'calli, 99. 

modestus, 101. 

platyrhinos, 101. 
dontomasi, Gaigeia, 153, 224. 
dorbignyi, Bachia, 213. 
dorsalis, Crotaphytus, 77, 78. 

Dipsosaurus, 78. 

Dipsosaurus dorsalis, 77, 78, 217, 
227. 
douglassii, Phrynosoma, 96, 100. 
draconoides, Callisaurus, 85, 86. 

Callisaurus draconoides 86, 217. 



dugesii, Eumeces, 161, 169, 221, 222. 
223. 

Phrynosoma orbiculare, 97, 98, 220. 

Sceloporus, 127. 

Sceloporus dugesii, 122, 127, 220, 
222, 223. 

Tapaya, 98. 
dunni, Anolis, 56, 61, 221. 
durissus. Constrictor durissus, 225. 
dussumieri, Chelonia, 15. 

Ecphymotes obtusirostris, 213. 
edwardsii, Ameiva, 174. 

Ameiva festiva, 171, 174, 218. 
edwardtaylori, Sceloporus, 110, 112, 224. 
effeldtii, Cinosternon, 23. 
Elaps fulvius, 225. 
elegans, Chrysemvs scripta, 32. 

Coleonyx, 41"", 42, 43. 

Coleonvx elegans, 42, 218, 224, 226, 
227, 228, 229. 

Emys, 32. 

Holbrookia, 84. 

Holbrookia elegans, 84. 

Holbrookia maculata, 82, 84, 226. 

Pseudemys, 32. 

Pseudemys scripta, 31, 32, 219, 228. 

Pseudemys troostii, 32. 

Uta, 148, 149. 

Uta stansburiana, 148, 150, 217. 
Elgaria, 194, 205. 

cedrosensis, 205, 206, 217. 

kingii, 205. 

kingii kingii, 205, 219, 227. 

kingii nobilis, 205, 206. 

multicarinata multicarinata, 205. 

multicarinata nana, 205, 207, 217. 

multicarinata webbii, 205, 207, 217. 

nobilis, 206. 

paucicarinatus, 205, 206, 217, 
emoryi, Amyda, 18, 219, 224, 228. 

Aspidonectes, 18. 
Emyidae, 19, 28, 29. 
Emys areolata, 30. 

bellii, 34. 

berardii, 19. 

callirostris, 32. 

cataspila, 32. 

elegans, 32. 

grayi, 33. 

incisa, 30. 

ornata, 31. 

pulcherrima, 30. 

umbra, 33. 

venusta, 31. 

Emysaurus, 20. 

rossignonii, 20. 

enneagrammus, Celestus, 195, 224, 228. 
Siderolamprus, 194. 



INDEX 



239 



Enyaliosaurus, 54, 75. 

clarki, 76, 223. 

defensor, 76, 77, 229. 

erythromelas, 76, 77, 218. 

quinquecarinatus, 76, 224. 
Epaphelus, 192. 

sumichrasti, 192. 
Eremonia, 15, 
Eretmochelys, 14, 16. 

imbricata, 16, 217, 218, 224, 225, 
226. 

squamata, 17. 
erythromelas, Cachryx, 77. 

Ctenosaura, 76, 77. 

Enyaliosaurus, 76, 77, 218. 
espiritensis, Verticaria, 187. 
estebanensis, Cnemidophorus, 189. 
Eublepharis fasciatus, 43. 

variegatus, 44. 
Euclielonia, 17. 
Euchirotes, 38. 

biporus, 38. 

Eumeces, 155, 160. 

altamirani, 160, 161, 163, 223. 
bocourtii, 166. 

brevilineatus, 162, 164, 224, 228. 
brevirostris, 162, 168, 220, 221, 222, 

223, 224, 225, 228. 
callicephalus, 161, 164, 219, 220, 

221, 222, 223, 225, 227, 229. 
colimensis, 162, 169, 220. 
copei, 162, 166, 220, 222, 223, 225. 
dicei, 162, 169, 224, 228. 
dugesii, 161, 169, 221, 222, 223. 
gilberti rubricaudatus, 161, 167, 

217. 
humilis, 162, 166, 226. 
indubitus, 162, 168, 222, 223. 
lagunensis, 161, 167, 2l7. 
Ivnxe furcirostris, 161, 163, 222, 
' 225, 228. 
lynxe lynxe, 161, 163, 222, 225, 226, 

228. 
multivirgatus, 161, 165, 219. 
obsoletus, 161, 165, 219, 224, 228. 
ochoterenai, 162, 169, 221. 
parviauriculatus, 162, 166, 219, 227. 
parvulus, 162, 166, 220, 223, 226. 
pavimentatus, 160. 
quadrilineatus, 167. 
rovirosae, 164. 
schmidti, 164. 

schwartzei, 160, 162, 218, 227, 229. 
skiltonianus amblygrammus, 167. 
skiltonianus lagunensis, 168. 
skiltonianus skiltonianus, 161, 167, 

217. 
sumichrasti, 161, 164, 218, 228. 
tetragrammus, 161, 165, 226, 228. 

Euphryne, 79. 

obesus, 79, 81. 
Euprepes lynxe, 163. 
Euprepis microcephalus, 214. 



exsul, Hemidactylus, 51. 

Uma, 89, 219. 
fantasticus, Sphaerodactylus, 52. 
fasciatus, Coleonyx, 42, 43, 220, 226. 

Crotaphytus, 94. 

Eublepharis, 43. 

Xenosaurus, 207, 208. 
fasciolatus, Crotaphytus, 94. 
ferrariperezi, Sceloporus, 126. 
fimbriatus, Gerrhonotus, 197. 
fissipes. Caiman, 212. 
flavescens, Kinosternon flavescens, 22, 
24, 219. 

Platythyra, 21, 24. 
flavilenta, Holbrookia maculata, 83. 
flavimaculata, Lepidophyma flavima- 

culata, 152, 218, 227, 228. 
flavimaculatus, Lepidophyma, 151, 152. 
flavomaculatum, Lepidophyma flavo- 

maculatum, 152. 
fioridanus, Sceloporus spinosus, 117. 
forbesorum, Leiolopisma, 159. 

Leiolopisma gemmingeri, 159. 

Scincella gemmingeri, 157, 159, 222. 
formosus, Sceloporus, 107, 108. 

Sceloporus formosus, 107, 108, 224, 
225, 228. 

Tropidolepis, 108. 
franciscensis, Verticaria, 187. 
frenatus, Hemidactylus, 40, 50, 221. 
frontale, Phrynosoma blainvillii, 103. 

Phrynosoma coronatum, 97, 103, 
217. 
frontalis, Phrynosoma, 103. 
fulvius, Elaps, 225. 
fulvus, Sceloporus, 135. 
furcirostris, Eumeces lynxe, 161, 163, 

222, 225, 228. 
fuscoauratus, Anolis, 55. 
fuscolabialis, Abronia, 196, 198, 224. 

Gerrhonotus, 198. 
fuscus. Caiman crocodilus, 212, 218. 

Gonatodes, 45, 218. 

Gonatodes albogularis, 45. 

Gymnodactylus, 45. 

Perosuchus, 212. 

Stenodactylus, 45. 

gabbii, Callisaurus draconoides, 86, 87, 

217, 227. 
Callisaurus ventralis, 87. 

gadovi, Cnemidophorus, 178, 185, 227. 

Urosaurus, 140, 145, 222, 223. 

Uta, 145. 
gadoviae, Sceloporus, 138, 221, 223, 224, 

225. 
gadovii, Anolis, 56, 61, 221. 

Barisia gadovii, 199, 200, 221. 

Gerrhonotus, 200. 
gaigeae, Ameiva undulata, 171, 172, 

218, 225, 228. 
Gaigeia, 153, 154, 222. 
Lepidophyma, 153, 154. 
Pseudemys scripta, 31, 33. 
Sceloporus lundelli. 111, 112, 229. 



240 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



Gaigeia, 151, 153. 

dontomasi, 153, 224. 

gaigeae, 153, 154,222. 

radula, 153, 224. 

sylvatica, 153, 154, 222. 
Gambelia, 55, 93. 

wislizenii, 93, 94. 

wislizenii wislizenii, 94, 217, 219, 
227. 
gambelii, Crotaphytua, 94. 
Gastrotropis, 5.5. 
Gavialidae, 2 JO. 
Gecko mabouia, 50. 
Geckonidae, 40. 
Gehyra, 51. 
Gekko rapicauda, 49. 

tuberculosus, 50. 
Gekkonidae, 39, 40, 41. 
gemmingeri, Leiolopisma, 159. 

Leiolopisma gemmingeri, 159. 

Lygosoma (Mocoa), 159. 

Oligosoma, 159. 

Scincella gemmingeri, 157, 159, 222, 
224, 228. 
Geoclemmys annulata, 29. 
Geoemyda, 29. 

areolata, 29, 30, 218, 225, 227, 228, 
229. 

pulcherrima incisa, 29, 30, 218, 224. 

pulcherrima pulcherrima, 29, 30, 
220, 221, 223, 226. 

rubida, 29, 30, 220, 221, 223, 224. 
georgeensis, Aristelliger, 51, 225. 

Idiodactylus, 51. 
geronimensis, Anniella, 209, 217. 
Gerrhonotus, 194, 203. 

adspersus, 201. 

antauges, 200. 

bocourti, 200. 

cedrosensis, 206. 

deppii, 196, 197. 

deppii digueti, 198. 

fimbriatus, 197. 

fuscolabialis, 198. 

gadovii, 200. 

gramineus, 198. 

imbricatus, 198, 201. 

infernalis, 204. 

kingii, 205, 206. 

lemniscatus, 204. 

levicoUis ciliaris, 202. 

levicoUis levicoUis, 202. 

lichenigerus, 201. 

liocephalus, 203. 

liocephalus austrinus, 203, 204, 218. 

liocephalus infernalis, 203, 204, 219, 
226. 

liocephalus liocephalus, 203, 214, 
220, 221, 222, 223, 224, 225. 

liocephalus loweryi, 203, 204, 226. 

liocephalus ophiurus, 203, 204, 225, 
228. 

matudae, 196. 

microcephalus, 214. 

modestus, 201. 

moreleti rafaeli, 199. 



Gerrhonotus moreleti temporalis, 199. 

multi-carinatus nanus, 207. 

multifasciatus, 206. 

nobilis, 206. 

oaxacae, 197. 

obscurus, 200. 

olivaceus, 201. 

ophiurus, 204. 

paucicarinatus, 206. 

(Barissia) planifrons, 201. 

rudicollis, 202. 

scincicauda ignavus, 207. 

scincicauda nanus, 207. 

scincicauda webbii, 207. 

taeniatus, 197. 

tessellatus, 203. 

vasconcelosii ochoterenai, 197. 

viridiflavus, 200. 

webbii, 207. 
gigas, Caretta, 16. 

Caretta caretta, 15, 16, 217, 226. 
gilberti, Xantusia, 154, 155, 217. 
glaucus, Sphaerodactylus, 53. 

Sphaerodact.ylus glaucus, 52, 53. 
218, 225, 227, 229. 
goldmani, Sceloporus, 136, 137, 219, 226. 

Terrapene, 35. 
Gonatodes, 40, 41, 45. 

albogularis, 45. 

albogularis fuscus, 45. 

fuscus, 45, 218. 
Goniodactylus braconnieri, 45. 
goodei, Anota, 102. 

Phrynosoma, 102. 

Phrynosoma platyrhinos, 96, 101^ 
2i7. 
Gopherus. 28. 

agassizii, 28, 217, 226. 

berlandieri, 28, 219, 224. 
gracilis, Cnemidophorus, 189. 

Cnemidophorus tigris, 177, 189,. 
227. 
graciosa, Uta, 144. 
graciosus, Sceloporus, 106. 

Urosaurus, 140, 141, 144, 217. 

Urosaurus ornatus, 145. 
grahamii, Cnemidophorus, 188. 
graminea, Abronia taeniata, 196, 198, 

225, 228. 
gramineus, Gerrhonotus, 198. 
grammicus, Sceloporus, 105, 119. 

Sceloporus grammicus, 119, 221, 224. 
grandaevus, Sator, 139, 140, 217. 
grandis, Cubina, 208. 

Xenosaurus, 207, 208, 225, 229. 
grayi, Emys, 33. 
grayii, Poriodogaster, 151. 
guatemalensis, Akleistops, 151, 152. 
guentheri, Sceloporus, 108. 
guentherii, Anolis, 64. 
gularis, Cnemidophorus, 183. 

Cnemidophorus gularis, 183. 

Cnemidophorus sackii, 178, 183,, 
224, 226, 228. 

Uta, 144. 
guntherii, Anolis, 57, 63. 



INDEX 



241 



guttata, Clemmys, 37. 
guttatus, Cnemidophorus, 183. 

Cnemidophorus guttatus, 175, 179, 
224, 228. 
guttulatus, Lamprosaurus, 160, 165. 
Gymnodactylus albogularis, 45. 

coleonyx, 42. 

fuscus, 45. 

scapularis, 42. 
Gymnophiona, 2. 
Gymnophthalmus, 39, 170, 192. 

lineata, 192. 

sumichrasti, 192, 224. 

harlanii, Phrynosoma, 99. 
hartwegi, Ameiva undulata, 171, 218. 
heliactin, Anolis, 58, 67, 224. 
Heloderma, 192. 

hernandesii, 193. 

horridum, 193, 208, 218, 220, 221, 
222, 223, 224, 226, 227. 

suspectum, 193, 227. 
Helodermidae, 40, 192. 
Hemichirotes, 38. 

tridactylus, 38. 
Hemidactylus, 40, 41, 49. 

exsul, 51. 

frenatus, 40, 50, 221. 

javanicus, 49. 

mabouia, 40, 50, 228. 

mabuya, 49. 

mutilatus, 51, 52. 

navarri, 51, 52. 

turcicus, 40, 50. 

turcicus turcicus, 50, 218, 227, 228, 
229. 
hemilopha, Ctenosaura, 73, 75, 217, 219, 

226, 227. 
henrici, Cinosternum, 25. 
henshawi, Xantusia, 154, 155, 217. 
hernandesi, Phrynosoma, 100. 

Phrynosoma douglassii, 96, 100, 219, 
227. 

Phrynosoma orbiculare, 96, 97, 98. 

Tapaya, 100. 
hernandesii, Heloderma, 193. 

Iguana, 72. 
hernandezii, Chamaeleopsis, 68. 

Corvthophanes, 68, 218, 224, 226, 
228, 229. 

Tapaya, 100. 
Herpetochalcis heteropus, 212. 
herrerai, Kinosternon, 22, 24, 228. 
hesperis, Uta stansburiana, 148, 149, 217. 
heterolepis, Sceloporus, 119, 121, 222. 
heteropa, Bachia, 212. 
heteropus, Chalcides, 212. 

Herpetochalcis, 212. 
heterurus, Sceloporus, 120. 
hiltoni, Pseudemys scripta, 31, 32, 226. 
hirtipes, Cinosternon, 25. 

Cinosternum, 25. 

Kinosternon, 22, 25, 219, 220, 221, 
222, 223, 226. 
hispidus, Sauromalus, 79, 80, 217. 



Holbrookia, 54, 81. 
approximans, 83. 
bischoffi, 84. 
bunkeri, 82, 84, 219. 
dickersonae, 83. 
elegans, 84. 
elegans elegans, 84. 
elegans thermophila, 84. 
lacerata, 83. 
maculata, 82. 
maculata approximans, 82, 83, 219, 

220. 222, 226, 227. 
maculata dickersonae, 82, 83, 219, 

220, 221, 226, 229. 
maculata elegans, 82, 84, 226. 
maculata flavilenta, 83. 
maculata lacerata, 82, 83, 219. 
maculata pulchra, 82, 84. 
maculata thermophila, 82, 84, 226, 

227. 
propinqua, 85. 

propinqua piperata, 82, 85, 228. 
propinqua propinqua, 82, 83, 85. 
pulchra, 84. 
texana, 82, 85, 219, 220, 224, 226, 

227, 228. 
thermophila, 84. 
homolepidurus, Phyllodactylus, 46, 48, 

219, 227. 
Phyllodactylus tuberculosus, 48. 

Homalosaurus, 86. 

ventralis, 86, 88. 
horridum, Heloderma, 193, 208, 218, 

220, 221, 222, 223, 224, 226, 227. 
Trachyderma, 192, 193. 

horridus, Sceloporus horridus, 110, 116, 

221, 223, 225. 
humeralis, Sceloporus, 134. 
humilis, Eumeces, 162, 166, 226. 
hyperythra, Verticaria hyperythra, 187. 
hyperythrus, Cnemidophorus, 174. 

Cnemidophorus hyperythrus, 175, 
187, 217. 

Idiodactylus, 51. 

georgeensis, 51. 
ignavus, Gerrhonotus scincicauda, 207. 
Iguana, 54, 72. 

(Ctenosaura) armata, 74. 

(Ctenosaura) bellii, 74. 

delicatissima, 72. 

hernandesii, 72. 

iguana iguana, 72. 

iguana rhinolopha, 72, 218, 220, 

221, 222, 223, 224, 225, 226, 227. 
228. 

(Ctenosaura) lanceolata, 74. 

H[ypsilophus] rhinolophus, 72, 

(Ctenosaura) similis, 73. 

tuberculata, 72. 
iguana, Iguana iguana, 72. 

Lacerta, 72. 
Iguanidae, 40, 53. 



242 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



imbricata, Barisia imbricata, 198, 199, 
201, 220, 221, 222, 223, 224, 225, 
228 
Eretmochelys, 16, 217, 218, 224, 

225, 226. 
Testudo, 16. 
imbricatus, Gerrhonotus, 198, 201. 
immucronatus, Sceloporus jarrovii, 123, 

128, 222, 225, 226, 229. 
immutabilis, Cnemidophorus, 180. 

Cnemidophorus giittatus, 175, 180, 

218, 220, 221, 223, 224. 
imperator. Boa, 225. 

Constrictor constrictor, 225. 
impetigosus, Anolis, 56, 60. 
incerta, Scincella, 156. 
incisa, Emys, 30. 

Geoemyda pulcherrima, 29, 30, 218, 
224. 
indubitus, Eumeces, 162, 168, 222, 223. 
infernalis, Gerrhonotus, 204. 

Gerrhonotus liocephalus, 203, 204, 

219, 226. 

inornatus, Cnemidophorus, 177, 184, 
219, 224. 

Sphaerodactylus, 53. 
insulana, Ctenosaura, 75. 
insularis, Crotaphytus, 92, 93, 217. 
integrum, Cinosternum, 25. 

Cinosternum scorpioides, 25. 

Kinosternon, 22, 25, 217, 220, 221, 
222, 223, 224, 225, 226, 228. 

Kinosternum, 21, 25. 
intermedius, Anolis, 63. 

Sceloporus, 127. 

Sceloporus dugesii, 122, 127, 221, 
223. 

Tropidolepis, 127. 
interrupta, Ctenosaura, 75. 
inusitatus, Callisaurus, 88. 

Callisaurus draconoides, 86, 88, 227. 

Callisaurus ventralis, 88. 
irregularis, Aristelliger, 51. 

Phymatolepis (Uta), 146. 

Urosaurus, 140, 146. 

Uta, 146. 
ixbaac, Leiolopisma cherriei, 158. 

Lygosoma assatum, 158. 

Scincella cherriei, 157, 158, 218,229. 

jacobi, Anolis, 68. 

jalapae, Sceloporus, 136, 225, 229. 

jamesi, Phrynosoma, 102. 

Phrynosoma coronatum, 97, 102, 
217. 
jarrovii, Sceloporus, 127. 

Sceloporus jarrovii, 122, 123, 128, 
219, 220, 227. 
javanica, Amyda, 18. 
javanicus, Hemidactylus, 49. 

kempii, Lepidochelys, 14, 15, 225. 

Thalassochelys (Colpochelys), 15. 
kidderi Anolis, 58, 66, 218, 229. 
kingii, Elgaria, 205. 

Elgaria kingii, 205, 219. 

Gerrhonotus, 205, 206. 



Kinosternidae, 19, 21. 
Kinosternon, 3, 21. 
abaxillare, 22, 218. 
acutum, 22, 23, 218, 227, 228. 
creaseri, 22, 23, 225, 229. 
cruentatum censors, 22, 24, 225, 

229. 
cruentatum cruentatum, 22, 23, 

218, 224, 227, 
flavescens flavescens, 22, 24, 219. 
flavescens stejnegeri, 22, 24, 219, 

220, 226. 
herrerai, 22, 24, 228. 
hirtipes, 22, 25, 219, 220, 221, 222, 

223, 226. 
integrum, 22, 25, 217, 220, 221, 

222, 223, 224, 225, 226, 228. 
leucostomum, 22, 25, 218, 225, 227, 

228. 
longicaudatum, 21. 
scorpioides, 21. 
scorpioides acuta, 23. 
sonoriense, 22, 26, 219, 220, 226. 
Kinosternum integrum, 21. 
mexicanum, 23. 
sonoriense, 26. 
klauberi, Terrapene, 34, 35, 227. 
Sauromalus, 79, 80, 217. 

labialis, Cnemidophorus, 178, 185, 217. 
lacerata, Holbrookia, 83. 

Holbrookia maculata, 82, 83, 219. 
Lacerta acanthura, 74. 

americana, 170. 

iguana, 72. 

lumbricoides, 39. 

mexicana, 39. 

orbicularis, 94, 95, 97. 

quadrilineata, 192. 

sexlineata, 174, 185. 

turcica, 50. 
Lacertus mabouya, 155, 156. 

orbicularis, 97. 
lachrymata, Chelonia, 16. 
lacordairei. Alligator, 211. 
Laemanctus, 54, 69 

alticoronatus, 70, 229. 

deborrei, 70, 71, 224, 227. 

longipes, 69, 70, 71, 220, 224, 228. 

serratus, 70, 218, 221, 222, 224, 
226, 228, 229. 
laeviventris, Anolis, 57, 58, 62, 63, 227, 
228. 

Dactyloa (Anolis), 62. 
lagunensis, Eumeces, 161, 167, 217. 

Eumeces skiltonianus, 168. 

Plestiodon, 168. 

Plestiodon skiltonianus, 168. 
Lampropholis assatus, 160. 
Lamprosaurus, 160. 

guttulatus, 160, 165. 
lanceolata. Iguana (Ctenosaura), 74. 
lanei, Phyllodactylus, 47, 48, 220, 221, 
222, 223, 226. 



INDEX 



243 



lar, Aristelliger, 51. 
laterale, Leiolopisma, 159. 

Lygosoma, 159. 

Scincella, 157, 159, 219. 
lateralis, Lysoptychus, 105, 132. 

Sceloporus, 132. 

Scincus, 159. 

Uta, 144. 

Uta ornata, 144. 

Uta (Phymatolepis), 144. 
latirostris, Crocodilus, 212. 
lativittis, Cnemidophorus, 178. 
Leiolopisma, 156. 

assatum assatum, 160. 

assatum taylori, 160. 

caudaequinae, 158. 

cherriei, 158. 

cherriei cherriei, 157. 

cherriei ixbaac, 158. 

cherriei stuarti, 158. 

forbesorum, 159. 

gemmingeri, 159. 

gemmingeri forbesorum, 159. 

gemmingeri gemmingeri, 159. 

laterale, 159. 

silvicolum, 158. 

telfairii, 156. 
Leiosaurus belli, 212. 
lemniscatus, Gerrhonotus, 204. 
Lepidochelys, 14. 

kempii, 14, 15, 225. 

olivacea, 14, 15, 218, 220, 221, 
224, 226. 
Lepidophyma, 151. 

flavimaculata flavimaculata, 152, 

218, 227, 228. 
flavimaculata obscura, 151. 
flavimaculatus, 151, 152. 
flavomaculatum flavomaculatum, 

152. 

gaigeae, 153, 154. 

smithii, 152. 

smithii occulor, 152, 153, 225, 226. 

smithii smithii, 152, 218. 

smithii tehuanae, 152, 218, 224. 

sylvatica, 154. 
Lepidosauria, 12, 37. 
leucostoma, Swanka, 25. 
leucostomum, Cinosternon, 25. 

Kinosternon, 22, 25, 218, 225, 227, 
228. 
levicollis, Barisia, 199, 202, 219. 

Gerrhonotus levicoUis, 202. 
levigata, Barisia gadovii, 199, 200, 224. 
lichenigerus, Gerrhonotus, 201. 
licki, Sceloporus, 114. 

Sceloporus orcutti, 111, 114, 217. 
Limnochelone, 19. 

micrura, 19. 
linearis, Urosaurus ornatus, 142, 143, 

219, 227. 
Uta ornata, 143. 

lineata, Gymnophthalmus, 192. 
lineatissimus, Cnemidophorus, 179. 

Cnemidophorus deppii, 176, 179, 

220, 221, 222, 223, 225. 



lineatulus, Sceloporus, 115. 

Sceloporus magister, 110, 115, 217. 
lineolateralis, Sceloporus, 122, 126, 220. 
lineolatus, Sphaerodactylus, 52. 
liocephalus, Gerrhonotus, 203. 

Gerrhonotus liocephalus, 203, 214, 

220, 221, 222, 223, 224, 225. 
liogaster, Anolis, 57, 62, 221. 
Liolaemus tenuis, 213. 
littoralis, Malaclemmys, 36. 

Malaclemmys terrapin, 36. 

Malaclemys terrapin, 36. 
longicaudatum, Kinosternon, 21. 
longicaudatus, Tapaya orbicularis, 97. 
longipes, Laemanctus, 69, 70, 71, 220, 

224, 228. 
Loricata, 2, 12, 209. 
loweryi, Gerrhonotus liocephalus, 203, 

204, 226. 
lucasensis, Dipsosaurus dorsalis, 78, 

217. 
lumbricoides, Lacerta, 39. 
lunaei, Sceloporus, 107. 
lundelli, Sceloporus lundelli, 111, 112, 

218. 
Lygosoma assatum assatum, 160. 

assatum cherriei, 157. 

assatum ixbaac, 158. 

cherriei cherriei, 157. 

cherriei stuarti, 158. 

(Mocoa) gemmingeri, 159. 

laterale, 159. 
lynxe, Eumeces lynxe, 161, 163, 222, 
225, 226, 228. 

Euprepes, 163. 
Lysoptychus, 105. 

lateralis, 105, 132. 

mabouia. Gecko, 50. 

Hemidactylus, 40, 50, 228. 
Mabouia brevirostris, 168. 
mabouya, Lacertus, 155, 156. 
Mabuia alliacea, 156. 
Mabuya, 155. 

mabouya mabouya, 156, 218, 220, 

221, 222, 224, 226, 227, 228, 229. 
mabuya, Hemidactylus, 49. 
maccalli, Anota, 99. 

Macrochelys, 20. 
maculata, Holbrookia, 82. 

Swanka, 23, 25. 
maculosa, Chelonia, 17. 
maculosus, Sceloporus, 105, 133, 220. 
magister, Sceloporus magister, 111, 112, 

114, 217, 219, 220, 227. 
magnatuberculatus, Phyllodactylus, 47, 

48, 221. 
magnus, Phyllodactylus, 47, 48, 218, 

221, 224. 
malachiticus, Sceloporus malachiticus, 

107. 
Malaclemmys littoralis, 36. 

terrapin littoralis, 36. 
Malaclemys, 29, 36. 

terrapin littoralis, 36. 
mannophorus, Uta, 148, 149, 217. 



244 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



mariarum, Cnemidophorus, 183. 

Cnemidophorus sackii, 178, 183, 
223. 
marmoratus, Cnemidophorus, 190. 

Cnemidophorus tigris, 177, 190, 
219, 220. 

Sceloporus, 131. 

Sceloporus variabilis, 130, 131, 219, 
224, 228. 

Staurotypus, 27. 
martinensis, Uta, 148, 149, 217. 
martyris, Cnemidophorus, 191. 

Cnemidophorus tessellatus, 191. 

Cnemidophorus tigris, 176, 191, 227. 
matudae, Gerrhonotus, 196. 
matudai, Abronia, 196, 218. 
mawii, Dermatemys, 19, 218, 224, 227, 

228, 229. 
maximus, Cnemidophorus, 176, 188, 

217. 
mayensis, Anolis sagrei, 56, 59, 218, 227, 
229. 

Micrurus affinis, 225. 
m'calli, Doliosaurus, 99. 
ni'callii, Anota, 95, 99. 

Phrynosoma, 96, 99, 217, 227. 
mearnsi, Streptosaurus, 91, 217. 

Uta, 91. 
meeki, Cnemidophorus gularis, 184. 
Megadactylus, 85. 
megalepidurus, Sceloporus, 106, 121, 

225, 228, 229. 
megalocephalus, Claudius, 26. 
megapholidotus, Anolis, 56, 59, 221. 
Megemys, 17. 
melanogaster, Sceloporus, 126. 

Sceloporus ferrariperezi, 126. 

Sceloporus torquatus, 122, 126, 217, 
221, 222, 223, 226, 229. 
melanorhinus, Sceloporus, 112, 113. 

Sceloporus melanorhinus, 110, 112, 
225. 
melanostethus, Cnemidophorus, 189, 190. 
mentalis, Phyllodactylus, 46. 
merriami, Sceloporus, 105, 132. 

Sceloporus merriami, 132. 
Mesaspis, 198. 

moreletii, 198. 
metallicus, Anolis, 56, 59. 
mexicana, Cistudo, 34. 

Cistudo (Onychotria), 34. 

Lacerta, 39. 

Onychotria, 34. 

Rhinoclemmys, 30. 

Terrapene, 35. 

Terrapene mexicana, 34, 35, 226, 
228. 
mexicanum, Kinosternum, 23. 
mexicanus, Chamaeleo, 68. 

Cnemidophorus, 181. 

Corythophanes, 68. 

Crocodilus, 211. 
microcephalus, Dasia, 214. 

Diploglossus, 214. 

Euprepis, 214. 

Gerrhonotus, 214. 



microlepidopterus, Sceloporus, 120. 
microlepidopus, Cnemidophorus, 180. 
microlepidotus, Sceloporus, 120. 

Sceloporus grammicus, 119, 120, 
220, 221, 222, 223, 225, 228, 229. 

Sceloporus microlepidotus, 120. 
microlepis, Sceloporus, 120. 
microscutata, Uta, 145. 
microscutatus, Urosaurus, 140, 145, 217. 
micrura, Limnochelone, 19. 
Micrurus affinis mayensis, 225. 
milleri, Anolis, 57, 64, 224. 
minor, Sceloporus jarrovii, 123, 128, 221, 
222, 224, 225, 226, 229. 

Sceloporus torquatus, 128. 
mississippiensis, Alligator, 211. 
mitratus, Brachydactylus, 41. 
mitrella, Cristasaura, 71. 
Mocoa, 159. 
Mocoa cherriei, 157. 
modesta, Anota, 101. 

Barisia, 199, 201, 228. 
modestum, Phrynosoma, 95, 101, 219, 

220, 224, 226, 227, 229. 
modestus, Doliosaurus, 101. 

Gerrhonotus, 201. 

Pterogasterus, 201. 
Molinia, 210. 

raonotropis, Diploglossus, 214. 
monserratensis, Sceloporus, 115. 

Sceloporus magister, 112, 115, 217. 
moreletii, Crocodylus, 210, 211, 218, 
220, 227, 228, 229. 

Mesaspis, 198. 
motaguae, Cnemidophorus, 181. 
mucronatus, Sceloporus mucronatus, 
123, 124, 222, 225, 229. 

Sceloporus torquatus, 124, 125. 
multicarinata, Elgaria multicarinata, 

205. 
multicarinatus, Cordylus (Gerrhonotus), 

205. 
multifasciatus, Gerrhonotus, 206. 
multiscutatus, Cnemidophorus tessella- 
tus, 191. 

Cnemidoohorus tigris, 177, 191, 
217. 
multispinis, Ctenosaura, 74. 
multivirgatum, Plestiodon, 165. 
multivirgatus, Eumeces, 161, 165, 219. 
muralis, Phyllodactylus, 46, 47, 224. 
murinus, Seps, 175. 
mutilatus, Hemidactvlus, 51, 52. 

Peropus, 51, 52,^223, 226. 
Mydas, 17. 

mydas, Chelonia, 17, 221, 223, 224, 225, 
226, 228, 229. 

Testudo, 17. 
Mydasea, 17. 

nana, Elgaria multicarinata, 205, 207, 

217. 
nannodes, Anolis, 63. 
nanus, Gerrhonotus multi-carinatus, 
207. 
Gerrhonotus scincicauda, 207. 



INDEX 



245 



nasuta, Caretta, 15. 

navarri, Heniidactylus, 51, 52. 

nebuloides, Anolis, 58, 66, 219, 220, 221, 

224, 225, 227. 

nebulosa, Chrysemys, 32, 217. 

Dactyloa, 55, 65. 

Pseudemys scripta, 31, 32, 217. 
nebulosus, Anolis, 58, 65, 220, 221, 222, 

223, 224, 225, 226. 
nelsoni, Phrynosoma, 103. 

Sceloporus, 138, 139, 219, 222, 223, 
226 227. 

Terrapene, 34, 35, 223. 

Urosaurus bicarinatus, 141, 147, 
225. 

Uta, 147. 

Uta bicarinata, 147. 
nemoralis, Coleonyx elegans, 42, 43, 

220, 221, 223, 224. 
Neoseps, 155. 
newmanorum, Xenosaurus, 207, 208, 

226. 
nigricauda, Uta, 145. 
nigricaudus, Urosaurus, 140, 145, 217. 
niloticus, Crocodylus, 210. 
nobilis, Elgaria, 206. 

Elgaria kingii, 205, 206. 

Gerrhonotus, 206. 
nolascensis, Uta, 148, 151, 227. 
Norops tropidoiiotus, 60. 

yucatanicus, 60. 
notata, Uma, 89. 

Uma notata, 89, 217. 
nuchalis, Basiliscus (Cristasaura), 72. 

oaxacae, Abronia, 196, 197, 224. 

Gerrhonotus, 197. 
oberon, Sceloporus jarrovii, 123, 129,219. 
obesus, Euphryne, 79, 81. 

Sauromalus obesus, 79, 81. 
obscura, Lepidophyma flavimaculata, 

151. 
obscurus, Gerrhonotus, 200. 

Sceloporus, 109. 
obsoletum, Plestiodon, 160, 165. 
obsoletus, Cnemidophorus gularis gu- 
laris, 182. 

Eumeces, 161, 165, 219, 224, 228. 
obtusirostris, Ecphymotes, 213. 
occidentalis, Cnemidophorus communis, 

182. 
occipitalis, Aneuporus, 213. 

Tropidurus, 213. 
occulor, Lepidophyma smithii, 152, 153, 

225, 226. 

ochoterenai, Abronia, 196, 197, 218. 

Eumeces, 162, 169, 221. 

Gerrhonotus vasconcelosii, 197. 

Phrynosoma, 103. 

Sceloporus, 133, 135, 221, 223. 
octolineatus, Cnemidophorus, 184. 

Cnemidophorus gularis, 183. 
Oedicoryphus, 71, 



oligoporus, Cnemidophorus deppii, 176, 
179, 218, 228. 
Sceloporus, 116. 

Sceloporus horridus, 110, 116, 220, 
221, 222, 223. 
Oligosoma gemmingeri, 159. 
olivacea, Chelonia, 14, 15. 

Lepidochelys, 14, 15, 218, 220, 221, 

224, 226. 
olivaceus, Gerrhonotus, 201. 

Sceloporus, 111, 117, 219, 224, 226, 
228. 
oUoporus, Sceloporus variabilis, 129, 

130. 
omiltemanus, Sceloporus, 124. 

Sceloporus mucronatus, 123, 124, 

221, 225. 
Sceloporus torquatus, 124. 
Onychotria, 34. 

mexicana, 34. 
Ophisaurus, 194. 
ophiurus, Gerrhonotus, 204. 

Gerrhonotus liocephalus, 203, 204, 

225, 228. 
Pterogasterus, 204. 

orbiculaire, Agama, 97. 
orbiculare, Phrynosoma, 96, 97, 98. 

Phrynosoma orbiculare, 96, 97, 219, 
220, 222, 223, 224, 225, 226, 228, 
229. 
orbicularis, Lacerta, 94, 95, 97. 

Lacertus, 97. 

Tapaya, 94, 98. 
orcutti, Sceloporus orcutti. 111, 114,217. 
ornata, Cistudo, 36. 

Emys, 31. 

Pseudemys scripta, 31, 33, 218, 222, 
224, 225, 226, 227, 229. 

Terrapene, 34, 36, 219, 227. 

Uta, 142. 

Uta ornata, 142. 
ornatus, Urosaurus ornatus, 142. 

Sceloporus, 127. 

Sceloporus ornatus, 122, 127, 219. 

pacificus, Crocodilus, 211. 
palearis, Cyclura, 76. 
pallida, Clemmys marmorata, 37, 217. 
palmeri, Uta, 148, 151, 227. 
panamensis, Anolis, 61. 
papillosus, Anelytropsis, 170, 226, 228. 
Paraloma, 71. 
parkeri, Ctenosaura, 75. 
parva, Ameiva undulata, 171, 173, 218, 
224. 

Uta, 149. 
parviauriculatus, Eumeces, 162, 166, 

219, 227. 
parviscutata, Uta, 145. 
parvulus, Eumeces, 162, 166, 220, 223, 

226. 
parvus, Sceloporus, 131. 

Sceloporus parvus, 129, 131, 219, 
224, 226. 
paucicarinatus, Elgaria, 205, 206, 217. 

Gerrhonotus, 206. 



246 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



pavimentatus, Eumeces, 160. 

Scincus, 160. 
pectinata, Ctenosaiira, 73, 75, 220, 221, 
222, 223, 224, 225, 226. 
Cyclura, 75. 
peninsularis, Coleonyx variegatus, 42, 

44, 217. 
pentaprion, Anolis, 55, 57, 61, 218. 
Anolis (Coccoessus), 61. 

gercarinatus, Corythophanes, 68, 69,218. 
eripia, 51. 

peronii, 51. 
peronii, Peripia, 51. 

Peropus, 51. 
Peropus, 40, 41, 51. 

mutilatus, 51, 52, 223, 226. 

peronii, 51. 
Perosuchus, 212. 

fuscus, 212. 
perplexus, Cnemidophorus, 184, 185. 
petersii, Anolis, 58, 65, 226, 228. 
Petrosaurus, 55, 90. 

repens, 90, 91, 217. 

thalassina, 90, 217. 
Phrvnosoma, 54, 94, 102. 

'asio, 95, 102, 218, 220, 221, 223, 224. 

blainvillii, 103. 

blainvillii frontale, 103, 

bufonium, 99. 

boucardi, 96, 98, 222. 

boucardii, 98. 

braconniori, 95, 100, 224, 225. 

calidiarum, 101. 

cerroense, 97, 103, 217. 

cornutum, 94, 95, 99, 219, 220, 
224, 226, 227. 

cornutum planiceps, 99. 

coronatum, 94, 96. 

coronatum blainvillii, 97, 103, 217. 

coronatum coronatum, 97, 102, 217. 

coronatum frontale, 97, 103, 217. 

coronatum jamesi, 97, 102, 217. 

ditmarsi, 95, 104, 227. 

douglassii, 96, 100. 

douglassii brachycercum, 96, 100, 
220. 

douglassii hernandesi, 96, 100, 

219, 227. 
frontalis, 103. 
goodei, 102. 
harlanii, 99. 
hernandesi, 100. 
jamesi, 102. 

m'callii, 96, 99, 217, 227. 
modestum, 95, 101, 219, 220, 224, 

226, 227, 229. 
nelsoni, 103. 
ochoterenai, 103. 
orbiculare, 96, 97, 98. 
orbiculare cortezii, 96, 98, 225, 228. 
orbiculare dugesii, 97, 98, 220. 
orbiculare hernandesi, 100. 
orbiculare orbiculare, 96, 97, 219, 

220, 222, 223, 224, 225, 226, 
228, 229. 

planiceps, 99. 



Phrynosoma platyrhinos, 95, 101. 

platyrhinos goodei, 96, 102, 227. 

platyrhinos platyrhinos, 96, 101, 
217. 

regale, 104. 

schmidti, 103. 

solare, 96, 104, 217, 226, 227. 

Solaris, 104. 

spinimentum, 102. 

taurus, 95, 104, 221, 225. 

wiegmanni, 97. 
Phyllodactylus, 41, 46. 

bordai, 46, 47, 221. 

delcampi, 46, 47, 221. 

homolepidurus, 46, 48, 219, 227. 

lanei, 47, 48, 220, 221, 222, 223, 226. 

magnatuberculatus, 47, 48, 221. 

magnus, 47, 48, 218, 221, 224. 

mentalis, 46. 

murahs, 46, 47, 224. 

pulcher, 46. 

tuberculosus, 47, 49, 217. 

tuberculosus homolepidurus, 48. 

unctus, 46, 47, 217. 

(Diplodactylus) unctus, 47, 

xanti, 49. 
Phymatolepis, 140, 144, 

bicarinatus, 140. 

bi-carinatus, 146. 

(Uta) irregularis, 146. 
picta, Testudo, 33. 

Verticaria, 186. 

Xantusia, 155. 
pictus, Claudius, 27, 

Cnemidophorus hyperythrus, 175, 
186 217 

Sceloporus,' 121, 225, 229. 
pilsbryi, Sceloporus, 120. 
piperata, Holbrookia propinqua, 82, 85, 

228. 
planiceps, Phrynosoma, 99. 

Phrynosoma cornutum, 99. 
planifrons, Barisia imbricata, 199, 201, 
224. 

Gerrhonotus (Barissia), 201. 
Platypholis, 160. 
platyrhina, Anota, 101. 
platyrhinos, Doliosaurus, 101. 

Phrj^nosoma, 95, 101. 

Phrvnosoma platyrhinos, 96, 101, 
217. 
Platysternidae, 18. 
Platythyra, 21. 

flavescens, 21, 24, 
Plestiodon bellii, 163. 

lagunensis, 168. 

multivirgatum, 165. 

obsoletum, 160, 165. 

quinquelincatum, 163. 

skiltonianum, 167. 

skiltonianus lagunensis, 168. 

tetragrammus, 165. 
Plethodontidae, 2. 
Pleurodira, 12. 
pleurostictus, Sceloporus, 119. 



INDEX 



247 



plioporus, Sceloporus serrifer, 122, 124, 

218, 226, 227, 229. 
Plistodon sumichrasti, 164. 
Pnoepus, 49. 
podarga, Ameiva undulata, 171, 172, 

226, 228. 
poinsettii, Sceloporus, 121, 122, 125, 
219, 220, 224. 

Sceloporus torquatus, 125. 
polyphemus, Testudo, 28. 
polystictus, Crotalus, 6. 
Poriodogaster, 151. 

grajii, 151. 
praesignis, Aristelliger, 51. 
prezygus, Sceloporus, 107, 109, 218. 
Pristicercus, 55. 
propinqua, Holbrookia, 85. 

Holbrookia propinqua, 82, 83, 85. 
propus, Chamaesaura, 39. 
Pseudemys, 29, 31. 

elegans, 32. 

floridana texana, 31, 33, 224. 

scripta cataspila, 31, 32, 228. 

scripta elegans, 31, 32, 219, 228. 

scripta gaigeae, 31, 33. 

scripta hiltoni, 31, 32, 226. 

scripta nebulosa, 31, 32, 217. 

scripta ornata, 31, 33, 218, 222, 
224, 225, 226, 227, 229. 

texana, 33. 

troostii elegans, 32. 

umbra, 31, 33, 224. 
pseudo-caretta, Chelonia, 16. 
pseudo-mydas, Chelonia, 16. 
Pterogastenes, 203. 
Pterogasterus modestus, 201. 

ophiurus, 204. 
ptychopleurus, Tropidurus, 213. 
pulcher, Phyllodactylus, 46. 
pulcherrima, Emys, 30. 

Geoemvda pulcherrima, 29, 30, 220, 
221, 223, 226. 
pulchra, Anniella, 209. 

Anniella pulchra, 209, 217. 

Holbrookia, 84. 

Holbrookia maculata, 82, 84. 
punctata, Testudo, 37. 
punctatissimus, Sphaerodactylus, 52. 
punctatum, Cinosternon, 26. 
punctilineatus, Cnemidophorus, 189. 
pyrocephalus, Sceloporus, 105, 138, 139, 

220, 221, 222, 223. 
pyrrhocephalus, Sceloporus, 139. 

quadrilineata, Lacerta, 192. 
quadrilineatus, Eumeces, 167. 
quinquecarinata, Cyclura, 76. 
quinquecarinatus, Enyaliosaurus, 76, 224. 
quinquelineatum, Plestiodon, 163. 
quinquelineatus, Scincus, 163. 

rackhami, Xenosaurus, 208, 218. 
radula, Gaigeia, 153, 224. 
rafaeli, Barisia moreleti, 198, 199, 218. 
Gerrhonotus moreleti, 199. 



Rapara, 20. 
rapicauda, Gekko, 49. 

Thecadactylus, 49. 
rapicaudus, Thecadactylus, 49, 229. 
regale, Phrynosoma, 104. 
remivaga, Caretta, 15. 
repens, Petrosaurus, 90, 91, 217. 

Uta, 90. 
Reptilia, 12. 
reticulatus, Crotaphytus, 92, 93, 219, 

224, 228. 
Rhinoclemys, 29. 
Rhinoclemmys mexicana, 30. 
rhinolopha, Iguana iguana, 72, 218, 220, 
221, 222, 223, 224, 225, 226, 227, 228. 
rhinolophus, I[guana] H[ypsilophus], 72. 
rhodostictus, Callisaurus, 87. 
Rhynchocephalia, 37. 
rodriguezii, Anolis, 64. 

Anolis limifrons, 57, 64, 218, 224, 
225, 229. 
rossignonii, Chelydra, 20. 

Emysaurus, 20. 
rostellum, Cinosternum, 25. 
rovirosae, Eumeces, 164. 
rozellae, Celestus, 195, 218. 
rubida, Chelopus, 30. 

Geoemvda, 29, 30, 220, 221, 223, 224. 
rubidus, Cnemidophorus, 191. 

Cnemidophorus tessellatus, 191. 

Cnemidophorus tigris, 176, 191,217. 
rubigenosus, Anolis, 64. 
rubricaudatus, Eumeces gilberti, 161, 

167, 217. 
rubriventris, Sceloporus, 119. 
rudicollis, Barisia, 199, 202, 222. 

Gerrhonotus, 202. 
rufidorsum, Sceloporus, 115. 

Sceloporus magister, 112, 115, 217, 
rufopunctata, Uma, 89. 
ruthveni, Anolis, 60. 

sackii, Cnemidophorus, 181. 

Cnemidophorus sackii, 178, 180, 
221, 223, 225. 

Cnemidophorus sexlineatus, 180, 181. 
sagraei, Anolis, 55. 
Saiientia, 2. 
sallaei, Anolis, 67. 
salvini, Sceloporus, 108. 

Sceloporus malachiticus, 107, 108, 
225, 229. 
salvinii, Staurotypus, 27, 224. 

Staurotvpus (Stauremys), 27. 
Sator, 55, 139. 

angustus, 139, 217. 

grandaevus, 139, 140, 217. 
Sauria, 2, 12, 37, 39. 
Saurochelys, 20. 
Sauromalus, 54, 79. 

ater, 79, 80, 217. 

australis, 79, 80, 217. 

hispidus, 79, 80, 217. 

klauberi, 79, 80^ 217. 

obesus obesus, 79, 81. 

obesus townsendi, 79, 81, 227. 



248 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



Sauromalus obesus tumidus, 79, 81. 

slevini, 79, 80, 217. 

townsendi, 81. 

varius, 79, 81, 227. 
scalaris, Cnemidophorus gularis, 182. 

Cnemidophorus sackii, 178, 182, 
219, 220. 

Sceloporus, 105, 137. 

Sceloporus scalaris, 136, 137, 220, 
221, 222, 223, 229. 
scapularis, Gymnodactylus, 42. 
Sceloporus, 55, 105. 

acanthinus, 108. 

aeneus, 137. 

aeneus aeneus, 136, 137, 220, 221, 

223, 225. 

aeneus bicanthalis, 136, 137, 222, 

224, 225, 228. 

asper, 107, 109, 221, 222, 223. 
binocularis, 126. 
bi-seriatus, 118. 
boulengeri, 113. 
bulleri, 122, 125, 222. 
carinatus, 134, 135, 218. 
cautus, 117, 219, 226, 229. 
chrysostictus, 105, 133, 218, 226, 

229. 
clarkii, 113. 
clarkii boulengeri, 111, 113, 222, 

223, 226, 227. 
clarkii clarkii, 111, 113, 219, 227. 
cochranae, 134. 
consobrinus, 118. 
couchii, 105, 129, 132, 219, 224. 
cozumelae, 129, 130, 226, 229. 
cupreus, 134. 

cyanogenys, 123, 125, 224, 228. 
delicatissimus, 131. 
digueti, 114. 
dispar, 120. 
disparilis, 120. 
dugesii, 127. 
dugesii dugesii, 122, 127, 220, 222, 

223. 
dugesii intermedius, 122, 127, 221, 

223. 
edwardtaylori, 110, 112, 224. 
ferrariperezi, 126. 
ferrariperezi binocularis, 126. 
ferrariperezi melanogaster, 126. 
formosus, 107, 108. 
formosus formosus, 107, 108, 224, 

225, 228. 

formosus scitulus, 107, 109, 221. 
fulvus, 135. 

gadoviae, 138, 221, 223, 224, 225. 
goldmani, 136, 137, 219, 226. 
graciosus, 106. 

graciosus vandenburgianus, 119, 

217. 
grammicus, 105, 119. 
grammicus alpha, 120. 
grammicus disparilis, 119, 120, 219, 

220, 221, 222, 224, 226, 228, 229. 



Sceloporus grammicus grammicus, 119. 

221, 224. 
grammicus microlepidotus, 119, 

120, 220, 221, 222, 223, 225, 228, 229. 
guentheri, 108. 
heterolepis, 119, 121, 222. 
heterurus, 120. 
horridus albiventris, 110, 116. 222. 

226. 
horridus horridus, 110, 116, 221, 

223, 225. 

horridus oligoporus, 110, 116, 220, 

221, 222, 223. 
humeralis, 134. 
intermedius, 127. 
jalapae, 136, 225, 229. 
jarrovii, 128. 

jarrovii immucronatus, 123, 128, 

222, 225, 226, 229. 

jarrovii jarrovii, 122, 123, 128, 219, 

220, 227. 
jarrovii minor, 123, 128, 221, 222, 

224, 225, 226, 229. 
jarrovii oberon, 123, 129, 219. 
jarrovii sugillatus, 123, 128, 222. 
lateralis, 132. 

licki, 114. 
lineatulus, 115. 
lineolateralis, 122, 126, 220. 
lunaei, 107. 

lundelli gaigeae. 111, 112, 229, 
lundelli lundelli, 111, 112, 218, 
maculosus, 105, 133, 220. 
magister lineatulus, 110, 115, 217. 
magister magister, 111, 112, 114, 

217, 219, 220, 227. 
magister monserratensis, 112, 115, 

217. 
magister rufidorsum, 112, 115, 217, 
magister zosteromus, 112, 115, 217. 
malachiticus acanthinus, 107, 108, 

218. 
malachiticus malachiticus, 107. 
malachiticus salvini, 107, 108, 225, 

229. 
malachiticus smaragdinus, 107. 
malachiticus taeniocnemis, 107, 218. 
marmoratus, 131. 
megalepidurus, 106, 121, 225, 228, 

229. 
melanogaster, 126. 
melanorhinus, 112, 113. 
melanorhinus calligaster, 110, 113, 

220, 221, 222, 223. 
melanorhinus melanorhinus, 110, 

112, 225. 
melanorhinus stuarti, 110, 113, 218. 
merriami, 105, 132. 
merriami annulatus, 132, 133, 219. 
merriami merriami, 132. 
microlepidopterus, 120. 
microlepidotus, 120. 
microlepidotus disparilis, 120. 
microlepidotus microlepidotus, 120. 
microlepis, 120. 
monserratensis, 115. 



INDEX 



249 



Sceloporus mucronatus aureolus, 123, 

124, 225, 229. 
mucronatus mucronatus, 123, 124, 

222, 225, 229. 
mucronatus omiltemanus, 123, 124, 

221 225. 
nelsoni, 138, 139, 219, 222, 223, 

226, 227. 
obscurus, 109. 

occidentalis biseriatus, 117, 118, 217. 
ochoterenai, 133, 135, 221, 223. 
oligoporus, 116. 
olivaceus, 111, 117, 219, 224, 226, 

228. 
omiltemanus, 124. 
orcutti licki, 111, 114, 217. 
orcutti orcutti. 111, 114, 217. 
ornatus, 127. 

ornatus caeruleus, 122, 127, 219. 
ornatus ornatus, 122, 127, 219. 
parvus, 131. 
parvus parvus, 129, 131, 219, 224, 

226. 
parvus scutulatus, 129, 131, 222, 

226. 
pictus, 121, 225, 229. 
pilsbryi, 120. 
pleurostictus, 119. 
poinsettii, 121, 122, 125, 219, 220, 

224. 
prezvgus, 107, 109, 218. 
pyrocephalus, 105, 138, 139, 220, 

221, 222, 223. 
pyrrhocephalus, 139. 
rubriventris, 119. 
rufidorsum, 115. 
salvini, 108. 
scalaris, 105, 137. 

scalaris scalaris, 136, 137, 220, 221, 

222, 223, 229. 

scalaris slevini, 136, 138, 219, 220, 

224, 227. 
scalaris unicanthalis, 136, 138, 222. 
fierrifer plioporus, 122, 124, 218, 

226, 227, 229. 
serrifer serrifer, 122, 123, 218, 229. 
siniferus, 105, 134. 
siniferus cupreus, 134, 225. 
siniferus siniferus, 134, 218, 221, 

223, 225. 
spinosus, 106, 107, 116. 
spinosus caeruleopunctatus, 110, 

116, 225. 
spinosus floridanus, 117. 
spinosus spinosus, 110, 111, 116, 

126, 217, 220, 221, 222, 223, 225, 

228, 229. 
squamosus, 134, 135, 218. 
stejnegeri, 107, 109, 221. 
taeniocnemis, 107. 
teapensis, 129, 130, 218, 225, 227, 

229. 
torquatus, 105, 106, 121, 
torquatus binocularis, 122, 126, 

224. 
torquatus cyanogenys, 125. 



Sceloporus torquatus melanogaster, 122, 
126,217, 221, 222, 223, 226, 229. 

torquatus minor, 128. 
torquatus mucronatus, 124, 125. 
torquatus omiltemanus, 124. 
torquatus poinsettii, 125. 
torquatus torauatus, 122, 125, 220, 

221, 222, 223, 225, 229. 
undulatus, 106, 107. 

undulatus consobrinus, 117, 118, 

219, 220, 224, 229. 
undulatus virgatus, 117, 118, 219, 227. 
utiformis, 105, 135, 220, 221, 222, 

223, 226. 
vandenburgianus, 119. 
variabilis, 105, 130. 
variabilis marmoratus, 130, 131, 

219, 224, 228. 
variabilis olloporus, 129, 130. 
variabilis smithi, 130, 131, 225. 
variabilis variabilis, 130, 218, 221, 

222, 225, 226, 228, 229. 
viviparus, 108. 
westphalii, 127. 
zosteromus, 115. 

schiedii, Anolis, 57, 58, 63, 64. 

Dactyloa, 63. 
schlegelii, Dermochelys, 13. 

Sphargis coriacea, 13. 
schmidti, Anolis, 56, 58, 60, 220. 

Cnemidophorus hyperythrus, 175, 
187, 217. 

Eumeces, 164. 

Phrynosoma, 103. 

Urosaurus ornatus, 142, 143, 219. 

Uta ornata, 143. 

Verticaria hyperythra, 187. 
schottii, Urosaurus ornatus, 141, 144, 

223, 226, 227. 
Uta, 144. 

Uta ornata, 144. 
schwartzei, Eumeces, 160, 162, 218, 227, 

229. 
Scincella, 155, 156. 

assata assata, 157, 160, 218. 

assata tavlori, 157, 160, 218, 220, 

221, 222, 224. 
caudaequinae, 157, 158, 224, 226. 
cherriei cherriei, 157, 218, 227. 
cherriei ixbaac, 157, 158, 218, 229. 
cherriei stuarti, 157, 158, 224, 228. 
gemmingeri forbesorum, 157, 159, 

222. 
gemmingeri gemmingeri, 157, 159, 

222, 224, 228. 
incerta, 156. 
laterale, 157, 159, 219. 
silvicola. 157, 158, 224, 228. 

scincicauda, Tropidoiepis, 205. 
Scincidae, 40, 155. 
Scincus agilis, 156. 

lateralis, 159. 

pavimentatus, 160. 

quinquelineatus, 163. 

sloanii, 156. 

telfairii, 156. 



250 BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



Scincus ventralis, 203, 214. 

scitulus Sceloporus formosus, 107, 109, 

221. 
scorpioides, Kinosternon, 21. 
scutulatus, Sceloporus parvus, 129, 131, 

222, 226. 
sealous, Cnemidophorus gularis, 183. 
semicristata, Cyclura, 74. 
semifasciatus, Cnemidophorus gularis, 
184. 
Cnemidophorus sackii, 178, 184, 
219. 
Seps murinus, 175. 

septemvittatus, Cnemidophorus, 184. 
sericea, Verticaria, 186. 
sericeus, Anolis, 58, 67, 218, 224, 226, 
227, 228. 
Cnemidophorus gularis, 183. 
Serpentes, 2, 12, 37. 
serpentina, Chelydra, 20, 21, 218, 228. 

Testudo, 20, 21. 
serratus. Laemanctus. 70, 218, 221, 222, 

224, 226, 228, 229. 
serrifer, Sceloporus serrifer, 122, 123, 

218, 229. 
severus, Claudius, 27. 
sexlineata, Lacerta, 174, 185. 
sexlineatus, Cnemidophorus, 177, 185. 
shawianum, Cinosternon, 23. 
shawii, Cyclura, 74. 
Siderolamprus, 194. 

enneagrammus, 194. 
silvicola, Scincella, 157, 158, 224, 228. 
silvicolum, Leiolopisma, 158. 
similis, Ctenosaura, 73, 218, 224, 226, 
227, 228, 229. 
Cyclura (Ctenosaura), 73. 
Iguana (Ctenosaura), 73. 
siniferus, Sceloporus, 105, 134. 

Sceloporus siniferus, 134, 218, 221, 
223, 225. 
sinistra, Ameiva undulata, 171, 174, 

220, 222, 223, 225. 
skiltonianum, Plestiodon, 167. 
skiltonianus, Eumeces, 161, 167, 217. 
slevini, Coleonyx variegatus, 42, 45, 217. 
Sauromalus, 79, 80, 217. 
Sceloporus scalaris, 136, 138, 219, 

220, 224, 227. 
Streptosaurus, 91, 217. 
Uta, 91. 
sloanii, Scincus, 156. 
smaragdinus, Sceloporus malachiticus, 

107. 
smithi, Sceloporus variabilis, 130, 131, 

225. 
smithii, Lepidophyma, 152. 

Lepidophyma smithii, 152, 218. 
solare, Phrynosoma, 96, 104, 217, 

226, 227. 
Solaris, Phrynosoma, 104. 



sonoriense, Cinosternum, 26. 

Kinosternon, 22, 26, 219, 220, 226. 

Kinosternum, 26. 

Thyrosternum, 26. 
sonoriensis, Coleonyx variegatus, 42, 
45, 227. 

Dipsosaurus dorsalis, 77, 79, 226, 227. 
Spasmocnemis, 51. 
speciosa, Blepharactisis, 192. 
spengleri, Testudo, 29. 
Sphaerodactylus, 41, 52. 

anthracinus, 52, 213. 

cinereus, 52. 

fantasticus, 52. 

glaucus, 53. 

glaucus glaucus, 52, 53, 218, 225, 

227, 229. 

glaucus torquatus, 52, 53, 225, 226. 

inornatus, 53. 

lineolatus, 52. 

punctatissimus, 52. 

sputator, 52. 

torquatus, 53. 
Sphargis coriacea schlegelii, 13. 
spinifera, Amyda, 18. 
spinimentum, Phrynosoma, 102. 
spinosus, Sceloporus, 106, 107, 116. 

Sceloporus spinosus, 110, 111, 116, 
126, 217, 220, 221, 222, 223, 225, 

228, 229. 
splendidus, Callisaurus, 87. 

Callisaurus draconoides, 86, 87, 217. 
sputator, Sphaerodactylus, 52. 
Squamata, 12, 37. 
squamata, Eretmochelys, 17. 

Uta, 148, 149, 217. 
squamosus, Sceloporus, 134, 135, 218. 
stansburiana, Uta, 148. 

Uta stansburiana, 148. 
Stauremys, 27. 
Staurotypus, 21, 27. 

biporcatus, 27. 

marmoratus, 27. 

salvinii, 27, 224. 

(Stauremys) salvinii, 27. 

triporcatus, 27, 227, 228. 
steindachneri, Diploglossus, 195. 
stejnegeri, Cnemidophorus, 191. 

Cnemidophorus tessellatus, 191. 

Kinosternon flavescens, 22, 24, 219, 
220 226. 

Sceloporus^ 107, 109, 221. 

Uta stansburiana, 148, 150, 217, 
219, 220, 227. 
stellata, Uta, 148, 150, 217. 
Stenodactylus fuscus, 45. 

variegatus, 44. 
stictogrammus, Cnemidophorus sackii,. 

178, 183, 219, 227. 
Streptosaurus, 55, 91. 

mearnsi, 91, 217. 

slevini, 91, 217. 
striatus, Celestus, 194. 

Cnemidophorus guttatus, 180. 



INDEX 



251 



stuarti, Ameiva undulata, 171, 173, 218, 
224, 227. 

Anolis, 63. 

Leiolopisma cherriei, 158. 

Lygosoma cherriei, 158. 

Sceloporus melanorhinus, 110, 113, 
218. 

Scincela cherriei, 157, 158, 224, 228. 
sup;illatus, Sceloporus jarrovii, 123, 128, 

222. 
sulcata, Chalcides, 39. 
sulcifrons, Anolis, 61. 
sumichrasti, Epaphelus, 192. 

Eumeces, 161, 164, 218, 228. 

Gvmnophthalmus, 192, 224. 

Plistodon, 164. 
suspectum, Heloderma, 193, 227. 
Swanka, 21. 

leucostoma, 25. 

maculata, 23, 25. 
sylvatica, Gaigeia, 153, 154, 222. 

Lepidophyma, 154. 
symmetrica, Uta, 144. 

Uta ornata, 144. 
sj'mmetricus, Urosaurus ornatus, 142, 
144, 217, 227. 

Tachy bates, 49. 

taeniata, Abronia . taeniata, 196, 197, 

222, 225, 228. 
taeniatus, Abronia, 197. 

Gerrhonotus, 197. 
taeniocnemis, Sceloporus, 107. 

Sceloporus malachiticus, 107, 218. 
Tapaia, 94. 
Tapaja, 94. 
Tapaya, 94, 95. 

boucardii, 98. 

cornuta, 99. 

cortezii, 98. 

dugesii, 98. 

hernandesi, 100. 

hernandezii, 100. 

orbicularis, 94, 98. 

orbicularis longicaudatus, 97. 
Tapayaxin, 97. 
Tapayia, 94. 

taurus, Phrynosoma, 95, 104, 221, 225. 
taylori, Anolis, 57, 61, 221. 

Leiolopisma assatum, 160. 

Scincella assata, 157, 160, 218, 220, 
221, 222, 224. 

Uta, 148, 150, 227. 
teapensis, Sceloporus, 129, 130, 218, 

225, 227, 229. 
tehuanae, Lepidophyma smithii, 152, 

218, 224, 
Teiidae, 39, 40, 170. 
telfairii, Leiolopisma, 156. 

Scincus, 156. 
temporalis, Barisia moreleti, 199, 218. 

Gerrhonotus moreleti, 199. 
tenuis, Liolaemus, 213. 
teres, Cyclura, 74. 



Terrapene, 29, 34, 

Carolina, 34. 

coahuila, 34, 36, 219. 

goldmani, 35. 

klauberi, 34, 35, 227. 

mexicana, 35. 

mexicana mexicana, 34, 35, 226, 
228. 

mexicana yucatana, 34, 35, 225. 

nelsoni, 34, 35, 223. 

ornata, 34, 36, 219, 227. 

triporcata, 27. 

yucatana, 35. 
tesselata, Ameiva, 188. 
tesselatus, Cnemidophorus, 3, 176, 188. 
tessellatus, Cnemidophorus, 3. 

Cnemidophorus tessellatus, 189. 

Gerrhonotus, 203. 
Testudines, 2, 12. 
Testudinidae, 19, 27. 
Testudo acutirostris, 20. 

agassizii, 28. 

berlandieri, 28. 

caouana, 15, 16. 

caretta, 16. 

cephalo, 16. 

clausa, 34. 

concentrica, 36. 

concinna, 31. 

coriacea, 13. 

imbricata, 16. 

mydas, 17. 

picta, 33. 

polyphemus, 28. 

punctata, 37. 

serpentina, 20, 21. 

spengleri, 29. 

viridis, 17. 
tetragrammus, Eumeces, 161, 165, 226, 
228 

Plestiodon, 165. 
texana, Anniella, 209. 

Cophosaurus, 81. 

Holbrookia, 82, 85, 219, 220, 224, 
226, 227, 228. 

Pseudemys, 33. 

Pseudemys floridana, 31, 33, 224. 
texanus, Cophosaurus, 85. 
thalassina, Petrosaurus, 90, 217. 

Uta, 90. 
Thalassochelys (Colpochelys) kempii, 
15. 

Thecadactylus, 41, 49. 
rapicauda, 49. 
rapicaudus, 49, 229. 
thermophila, Holbrookia, 84. 
Holbrookia elegans, 84. 
Holbrookia maculata, 82, 84, 226, 
227. 
thomasi, Ameiva undulata, 171, 173, 

218. 
Thyrosternum, 21, 
sonoriense, 26. 
tigris, Cnemidophorus sexlineatus, 182. 
Cnemidophorus tigris, 177, 189, 
217, 227, 



252 



BULLETIN 199, UNITED STATES NATIONAL MUSEUM 



torquata, Agama, 126. 

torquatus, Sceloporus, 105, 106, 121. 

Sceloporus torquatus, 122, 125, 220, 
221, 222, 223, 225, 229. 

Sphaerodactylus, 53. 

Sphaerodactylus glaucus, 52, 53, 
225, 226. 

Tropidurus, 121. 
townsendi, Sauromalus, 81. 

Sauromalus obesus, 79, 81, 227. 
Trachvderma, 192. 

hbrridum, 192, 193. 
Trachypilus, 55. 
tridactylus, Bipes, 38, 221. 

Hemichirotes, 38. 
triliratum, Cinosternum, 23. 
Trionicidae, 18. 
Trionychidae, 18. 
Trionychoidea, 12, 18. 
triporcata, Terrapene, 27. 
triporcatus, Staurotypus, 27, 227, 228. 
Tropidogaster, 94. 
Tropidolepis, 105, 205. 

formosus, 108. 

intermedius, 127. 

scincicauda, 205. 
tropidonotus, Anolis, 56, 59, 60, 218, 
224, 225, 227, 228, 229. 

Norops, 60. 
Tropidopilus, 55. 
Tropidurus bocourti, 213. 

occipitalis, 213. 

ptychopleurus, 213. 

torquatus, 121. 
tuberculata, Iguana, 72. 

Uta, 147. 

Uta bi-carinata, 147. 
tuberculatus, Urosaurus bicarinatus, 

141, 147, 220, 222, 226, 227. 
tuberculosus, Gekko, 50. 

Phyllodactylus, 47, 49, 217. 
tumidus, Sauromalus obesus, 79, 81. 
turcica, Lacerta, 50. 
turcicus, Hemidactylus, 40, 50. 

Hemidactylus turcicus, 50, 218, 
227, 228, 229. 
typica, Cnemidophorus mexicanus, 181 

Uma, 54, 89. 

exsul, 89, 219. 

notata, 89. 

notata cowlesi, 89, 90, 227. 

notata notata, 89, 217. 

rufopunctata, 89. 
umbra, Emys, 33. 

Pseudemys, 31, 33, 224. 
unctus, Diplodactylus, 47. 

Phyllodactylus, 46, 47, 217. 

Phyllodactylus (Diplodactylus), 47. 
undulata, Agama, 105. 

Ameiva undulata, 71, 174, 224. 
undulatus, Cnemidophorus, 174. 

Sceloporus, 106, 107. 
unica, Uta, 147. 

unicanthalis, Sceloporus scalaris, 136, 
138, 222. 



unicolor, Cnemidophorus, 180. 
unicus, Urosaurus, 141, 147, 219. 
uniformis, Anolis, 59, 60. 

Anolis humilis, 56, 60, 218, 227. 
Urosaurus, 55, 140. 

auriculatus, 140, 146, 220. 

bicarinatus anonymorphus, 141, 
146, 218, 221, 225. 

bicarinatus bicarinatus, 141, 146, 
221, 223, 225. 

bicarinatus nelsoni, 141, 147, 225. 

bicarinatus tuberculatus, 141, 147, 
220, 222, 226, 227. 

clarionensis, 141, 142, 220. 

gadovi, 140, 145, 222, 223. 

graciosus, 140, 141, 144, 217. 

irregularis, 140, 146. 

microscutatus, 140, 145, 217. 

nigricaudus, 140, 145, 217. 

ornatus caeruleus, 141, 143, 219. 

ornatus chiricahuae, 142, 143. 

ornatus graciosus, 145. 

ornatus linearis, 142, 143, 219, 227. 

ornatus ornatus, 142. 

ornatus schmidti, 142, 143, 219. 

ornatus schottii, 141, 144, 223, 226, 
227. 

ornatus symmetricus, 142, 144, 217, 
227. 

unicus, 141, 147, 219. 
ustus, Anolis, 58, 66, 218, 229. 
Uta, 1, 55, 146, 147. 

anonymorpha, 146. 

auriculata, 146. 

bicarinata, 146. 

bicarinata anonymorpha, 146. 

bi-carinata bi-carinata, 146. 

bicarinata nelsoni, 147. 

bi-carinata tuberculata, 147. 

caerulea, 143. 

clarionensis, 142. 

concinna, 148, 150, 217. 

elegans, 148, 149. 

gadovi, 145. 

graciosa, 144. 

gularis, 144. 

irregularis, 146. 

lateralis, 144. 

(Phymatolepis) lateralis, 144. 

mannophorus, 148, 149, 217. 

martinensis, 148, 149, 217. 

mearnsi, 91. 

microscutata, 145. 

nelsoni, 147. 

nigricauda, 145. 

nolascensis, 148, 151, 227. 

ornata, 142. 

ornata chiricahuae, 143. 

ornata lateralis, 144. 

ornata linearis, 143. 

ornata ornata, 142. 

ornata schmidti, 143. 

ornata schottii, 144. 

ornata symmetrica, 144. 

palmeri, 148, 151, 227. 

parva, 149. 



INDEX 



253 



Uta parviscutata, 145. 

repens, 90. 

schottii, 144. 

sle villi, 91. 

squamata, 148, 149, 217. 

stansburiana, 148. 

stansburiana elegans, 148, 150, 217. 

stanslMiriana hesperis, 148, 149, 217. 

stausburiaua stansburiana, 148. 

stansburiana stejnegeri, 148, 150, 
217, 219, 220, 227. 

stellata, 148, 150, 217. 

syinmetrica, 144. 

taylori, 148, 150, 227. 

thalassiiia, 90. 

tuberculata, 147. 

unica, 147. 
utiformis, Sceloporus, 105, 135, 220, 

221, 222, 223, 226. 
utowanac, Anolis, 57, 62, 226. 

vandenburghi, Cnemidophorus, 191. 
vandenburgianus, Sceloporus, 119, 217. 

Sceloporus graciosus, 119, 217. 
variabilis, Sceloporus, 105, 130. 

Sceloporus variabilis, 130, 218, 221, 
222, 225, 226, 228, 229. 
variegatus, Coleonyx, 44. 

Coleonvx variegatus, 42, 44, 217, 
227/ 

Eublepharis, 44. 

Steiiodactylus, 44. 
variolosus, Cnemidophorus, 190. 
varius, Sauromalus, 79, 81, 227. 
velox, Cnemidopliorus gularis, 184. 
vent rails, C'allisaurus, 88. 

Callisaurus dracouoides, 86, 88, 227. 

Callisaurus ventralis, 88. 

Homalosaurus, 86, 88. 

Scincus, 203, 214. 
venusta, Emys, 31. 
veraepacis, Anolis ustus, 67. 

Verticaria, 174. 
beldingi, 186. 
caerulea, 186. 
ceralbensis, 188. 
espiritensis, 187. 
franciscensis, 187. 
hyperythra beldingi, 186. 
hyperythra hyperythra, 187. 
hyperythra schmidti, 187. 
picta, 186. 
sericea, 186. 



verus, Cnemidophorus gularis gularis, 

182. 
vigilis, Xantusia, 154, 217. 
virgata, Chelonia, 17. 
virgatus, Sceloporus undulatus, 117, 118, 

219, 227. 
viridiflava, Rarisia, 108, 200, 224. 
viridifiavus, Gerrhonotus, 200. 
viridis, Anolis, 55. 

Testudo, 17. 
vittatus, Basiliscus, 71, 218, 220, 221, 

222. 223, 224, 225, 227, 228, 229. 

webbii, Elgaria multicarinata, 205, 207, 
217. 

Gerrhonotus, 207. 

Gerrhonotus scincicauda, 207. 
westphalii, Sceloporus, 127. 
wiegmanni, Anolis, 63. 

Phrynosoma, 97. 
wislizenii, Crotaphvtus, 94. 

Gambelia, 93, 94. 

Gambelia wislizenii, 94, 217, 219, 
227. 

xanti, Phyllodactylus, 49. 
Xantusia, 151, 154. 

gilberti, 154, 155, 217. 

henshawi, 154, 155, 217. 

picta, 155. 

vigilis, 154, 217. 
Xantusiidae, 39, 151. 
Xenosauridae, 40, 207. 
Xenosaurus, 207. 

fasciatus, 207, 208. 

grandis, 207, 208, 225, 229 

newnianorum. 207, 208, 226. 

rackhami, 208, 218. 
Xerobates agassizii, 28. 

berlandieri, 28. 

yucatana, Cistudo, 35. 

Terrapene, 35. 

Terrapene mexicana, 34, 35, 225. 
yucatanicus, Norops, 60. 

Zablepsis, 154. 

zosteromus, Sceloporus, 115. 

Sceloporus magister, 112, 115, 217. 



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