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Flowering Plants 

Vol. 8, part 1 

Cyclopaedia of Collectors - Supplement 2 



Accraceae . . . 




Dipsacaceae . 



Passifloraceae . 



Actinidiaceae s. str. 



Droseraceae . 4 







Aizoaceae . 






Pentaphragmataceae . 



Alismataceae . 5: 




Epacridaceae . 



Pentaphylacaceae . 




Ericaceae . 



Philydraceae . 
















Fagaceae . 






Aponogetonaceae < 

: il 






Plumbaginaceae . 







Podostemaceae A 







5: 1 




Po lemon iaceae 



Betulaceae . .5: 










Bixaceae s. str. 



Geraniaceae . 



Portulacaceae . 



Burmanniaceae . 




Gnetaceae . . A 




Primulaceae . 










5: 209, 








Butomaceae . 







Restionaceae . 




Byblidaceae . . 






Rhizophoraceae 5: 







Haloragaceae . 



Salicaceae . 




Campanulaceae . 

6: 107, 














Sarcospermaceae . 









Saururaceae . 






4: 175, 



















6: 193, 


Celastraceae 6: 227, 389, 


Hypericaceae . 




Centrolepidaceae . 






4: 280, 




Ceratophyllaceae . 



Juglandaceae , 



Sparganiaceae . 




Juncaceae . 



Sphenocleaceae . 







Juncaginaceae . 



Stackhousiaceae . 



Clethraceae . 






Staphyleaceae . 






Loganiaceae . 

6: 293, 


Sty idiaceae 










4: 533; 5: 




Malpighiaceae . 




Connaraceae . 5: 




Martyniaceae . 



4: 349; 6 




Molluginaceae . 



Trapaceae . 



4:338, 599; 5: 




Moringaceae . 



Trigoniaceae , 




Myoporaceae . 



Turneraceae . 










Typhaceae . 



Crassulaceae . 











Nyctaginaceae . 



4: 113, 595; 5: 







Nyssaceae . 



Valerianaceae . 



Dichapetalaceae 5: 




Ochnaceae . 



Violaceae . 



Dilleniaceae . 



Oxalidaceae . 



Xyridaceae . 

4: 366, 





Papaveraceae . 












In this 2nd and final supplement (to Cycl. in Fl. Mai. I, 1, 1950 and Suppl. I 
in Fl. Mai. I, 5, 1958, p. ccxxxvii-cccxlii) relatively little space is occupied by 
collectors of former centuries as only few additional data could be unearthed. 
Recent collectors are included as far as known to me, but several did not answer 
my queries and their entries will be very incomplete. 

Contemporary collectors generally pay more attention to the numbering and 
labelling, and taxonomists will not have much difficulty in locating their finds. 
This is one of the reasons that I have decided to close with this second supplement. 
I expect that the chapters 'Obituaries and Biographical Notes' and 'Expeditions 
and Explorations' in Flora Malesiana Bulletin will be continued, and will give 
ample information for those interested. 

June 1973 M. J. van Steenis-Kruseman. 



6. Malesian botanical collections in which errors 

Cyclopaedia p. xxiii-xxix add : 
F. de Lahaie, see C. A. G. Riche. 

C. A. G. Riche and F. de Lahaie, naturalists of 
the voyage in 'La Recherche' and 'L'Esperance' 
in search of La Perouse, 1791-1794, collected in 
Mauritius and Reunion. Part of the plants have 
erroneously been labelled 'Java'. 


Cyclopaedia p. xxx-xxxi, §1, add: 

Archer, Mildred, Natural History Drawings 
in the India Office Library. London 1962. 

Of the numerous collections of drawings several 
of the botanical ones were removed to Kew and 
the British Museum. The remaining collections are 
listed in a Catalogue (p. 71-102). The book 
contains a host of information on the plates, their 
collectors, painters, etc. 

Backer, C. A., Onkruidflora der Javasche 
suikerrietgronden. Add: The final instalment of the 
atlas, containing 220 plates, was published in May 

Register of Photographs ( 71971) in the Collection 
of the Forest Department, Sarawak, Malaysia. 

This stencilled register has been distributed; the 
collection was started in 1963. 

Cyclopaedia p. xxxi-xxxiii, §2, add: 

Reinwardt, C. G. C. At the end of the entry 
add: A collection of Pa yen's non-botanical 
drawings is in the Ethnographical Museum Leyden. 

Rowan, Ellis. A number of drawings from E. 
New Guinea. See: H. J. Samuel, The Story of 
Ellis Rowan. London, Constable & Co., 1961, 
151 pp., ill. 

Surbeck. Miss H. E. Surbeck, zum Brunquell, 
Hallau (Schaffhausen), Switzerland, has some 200 
water colour drawings of Suniatran plants made 
obviously by her father, H. Surbeck, who also 
collected plants. 


Cyclopaedia p. xxxv add to: 

Weed Floras 

(2) Add: The unpublished 220 plates of this 
Flora, which were still preserved at the Pasuruan 
Sugarcane Experiment Station, were published in 
May 1973. 

Cyclopaedia p. xxxvi-xxxvii: 

Flora of (he Malay Peninsula 


(7) Henderson, M. R., Malayan Wild Flowers 
(Dicotyledons pts 1-3 in Mai. Nat. J. 1949-51; 
Monocotyledons publ. by the Mai. Nature Soc., 
Kuala Lumpur 1954). 

(8) (iiLLiLAND, H. B., Common Malayan Plants. 
Smgaporc 1958. Selected drawings with notes. 

(9) HfcNDtRSfjN. M, R., Common Malayan 
Wild/lowers. Longmans London 1961 (in scries 
Malayan Nature Handbooks). 

(10) GiLLiLAND, H. B., A revised Flora of 
Malaya. IIL Grasses of Malaya. 1971. 

(11) Tree Flora of Malaya. 1st vol. 1972. 

For the ferns see under 'Pteridophyta'. 
Cyclopaedia p. xxxvii-xxxviii : 

Flora of Java 

(10) Add: The final instalment of the atlas, 
containing 220 plates, was published in May 1973. 

(14) BACKERf, C. A. & R. C. BaKHUIZEN VAN 

DEN Brink Jr, Flora of Java (Spcrmatophytes 
only). Groningen 1963-1968, 3 vols. With a 
concise plant-geography of Java by C. G. G. J. 
van Steenis Sl Mrs. A. F. Schippers-Lammertse 
in vol. 2 (1965) p. (l)-(72), with photogr. and 2 

(15) Steenis, C. G. G. J. van. The Mountain 
Flora of Java. Leiden 1972. With 20 introductory 
chapters elucidating all involved aspects, 57 
coloured (originally hand-drawn) plates depicting 

(I) No attempt is made to make exhaustive additions, as these can be found in the Flora Malesiana 

Flora Malesiana 

[ser. I, vol. 8^ 

456 species with extensive captions on features not 
seen on the plates, as size of the total plant, 
habitat, occurrence elsewhere, etc. 72 Photographs 
are reproduced to give an impression of the habitats 
and vegetation types. 

Cyclopaedia p. xxxviii-xxxix : 

Flora of Borneo 

(6) Add: Continued in Reports Sarawak 
Museum 1924-34. 


(20) Merrill, E. D., A brief survey of the present 
status of Bornean Botany (Webbia 12, 1950, p. 

(21) Browne, F. G., Forest Trees of Sarawak 
and Brunei and their Products. Kuching, Sarawak 
1955, 369 + xviii pp., 47 pi., map. 

(22) AsHTON, P. S., Ecological Studies in the 
Mixed Dipterocarp Forests of Brunei State. 
Oxford Forestry Memoirs no 25, 1964, 75 pp., 37 
pi., 4 App., tables, figs. 

(23) AsHTON, P. S., A Manual of the Dipterocarp 
Trees of Brunei State. Oxford Univ. Press 1964, 
xii + 242 pp., 58 pi., 20 fig. 

(24) A Discussion on the Results of the Royal 
Society Expedition to North Borneo, 1961 (Proc. 
R. Soc. B, 161, 1964, p. 1-91, 36 fig., 3 tab.). 

(25) Meijer, W. & G. H. S. Wood, Dipterocarps 
of Sabah (North Borneo). Sabah Forest Record 
no 5, 1964, 344 pp., 59 text fig., 30 pi., map. 

(26) AsHTON, P. S., A Manual of the Dipterocarp 
Trees of Brunei State and of Sarawak . Supplement. 
Sarawak Forest Department 1968, viii + 129 pp., 
23 pi., 15 fig. 

(27) Anderson, J. A. R., is engaged in a 
checklist of trees of Sarawak. 

Numerous important papers on the non- 
Indonesian parts were published by J. A. R. 
Anderson, P. S. Ashton, E. F. (W. O.) Brunig, 
E. J. H. Corner, and W. Meijer (mostly mimeo- 
graphed at Sandakan by the Forestry Service); see 
also the Unesco Symposium on ecological research 
in humid tropics vegetation. Kuching 1963 (publ. 
1965). Japanese botanists started with the publi- 
cation of the results of their expeditions. A book. 
Trees of Sabah, is in preparation. 

Cyclopaedia p. xxxix-xl: 

Flora of New Guinea 


(20) Womersley, J. & J. B. McAdam c.s.. The 
Forests and Forest conditions in the Territories of 
Papua and New Guinea. Brit. Commonwealth For. 
Conf. 1957. 62 pp., with tabl. & photogr. 

Numerous important papers resulted from the 
explorations of C.S.I.R.O., the Forestry Division 

at Lae (incl. papers by P. van Royen), and the 
Australian National University, for the eastern 
half of the island; for reports on the western half 
see C. Kalkman, W. Vink, and others. 

A Handbook of Natural Resources of Papuaj New 
Guinea by C.S.I.R.O., Canberra, is in preparation. 
The organization of the volume on Botany and 
Vegetation is in the hands of Dr. Paymans. 

A Manual of the Forest Trees of Papua and New 
Guinea is issued, of which by now 9 parts appeared. 
Dept of Forests, Administration of Papua and 
New Guinea, Port Moresby. 

The Division of Botany, Department of Forests, 
Lae, New Guinea, issues Botany Bulletins, in- 
cluding a 'Manual of the Grasses', and a Check 
List of the Species of the Mount Wilhelm Flora. 

Cyclopaedia p. xl: 

Preservation of Nature 


(5) Report of the Trustees for National Parks, 
1959-1960. Kuching, Sarawak, 35 pp., 8 photogr., 
4 maps. 

(6) Wyatt-Smith, J. & P. R. Wycherley (ed.), 
Nature Conservation in Western Malaysia. Spec. 
Issue Mai. Nat. J., Kuala Lumpur 1961, viii -|- 
261 pp., 45 pi., fig., maps. 

(7) Basjarudin, H., Problems of National Parks 
and Reserves in Indonesia and Emerging Countries. 
lUCN Publ. n.s. no 10, 1968, p. 386-393, map 
(with principal conserved areas). Protected 
animals listed ; survey of conserved areas in 
Indonesia; general organization of conservation 
in Indonesia. 

(8) Basjarudin, H., Nature Reserve and National 
Parks in Indonesia. Present situation and problems. 
Stencilled account for lUCN IIT. M/J.6 session, 
New Delhi, Nov. 24, 1969, 6 pp. 

(9) Wycherley, P. R., Conservation in Malaysia. 
A manual on the conservation of Malaysia's 
renewable natural resources. lUCN Publ. n.s. 
Suppl. Paper no 22, 1969, 207 pp., stencilled. 

(10) National Parks of Malaysia by various 
authors. Spec, double Issue Mai. Nat. J., Kuala 
Lumpur vol. 24, Aug. 1971, Part 3 & 4, p. 111- 
262, fig., pi., maps. 

A MS overall survey: Plant Conservation in the 
Malesian Tropics, has been written by C. G. G. J, 
Van Steenis. Presumably it will be published in 
the future. 

Cyclopaedia p. xli : 



(15) Holttum, R. E., a revised Flora of 
Malaya. Vol. 2. Ferns of Malaya. Singapore 1955. 

1. the etymological use of vernacular names for 

malesian plants 

Cyclopaediap.xliii, 1st paragraph: ^ut contributed to his father's (later brother's) 

Correction : Hugh Low was no Veitch collector, nursery at Clapton. 




Cycl. Fl. Mai. I, 1, 1950, p. Ixvi-kix, and Suppl. 
I.e. 5, 1958, p. ccxxxvii, add to 

14. Selected literature etc. 

Anonymous (1962): Climbing mountains in 
Malaya (Mai. Nat. J. 16, p. 157-196, 3 fig., 2 maps, 
8 photogr.). 

ANON"i'MOUS (1965): Instructions for Collectors. 
No. 10. Plants. Brit. Mus. Nat. Hist. 6th ed. (repr. 
with amendments). London, 72 pp. 

Da\is, p. H. (1961): Hints for hard-pressed 
collectors (Watsonia 4, p. 283-289). 

Dewolf Jr, Gordon P. (1968): Notes on making 
an herbarium (Arnoldia 28, p. (69)-(lll), w. fig. a. 
pi., bibliogr.). 

FosBERG, F. R. & M.-H. Sachet (1965): 
Manual for Tropical Herbaria (Regn. Veget. 39, 
Utrecht, 132 pp., 16 fig.). 

GL7LLAUMIN, A. (1942): Formulaire technique 

du botaniste preparateur et voyageur (139 pp., 
82 fig.). 

Heim, R., Mme S. Jovet-Ast, R. Laine & P. 
BoERELLY (1950): Conseils pour le recolte des 
Cryptogames (Paris, 32 pp., 6 fig.). 

Santapau, H. (1955): Instructions for field 
collectors of the Botanical Survey of India 
(Ministr. of Nat. Res. & Sc. Res. New Delhi, 
16 pp.). 

Simon, C. (1962): Erfahrungen mit wenig 
bekannten Methoden der Herbartechnik (Bau- 
hinia 2, p. 63-69). 

WoMERSLEY, J. S. (1953): Notes on the col- 
lecting of botanical specimens (Papua & New 
Guinea Gazette 8, p. 62-65, 5 fig.). 

WoMERSLEY, J. S. (1957): Paraformaldehyde as a 
source of formaldehyde for use in botanical 
collecting (Rhodora 59, p. 299-303). 


Cycl. Fl. Mai. I, 1, 1950, p. cvii-cxvi: 

Since 1950 a considerable amount of collecting has 
been done in specific areas. A detailed enumeration 
as has been given in the Cyclopaedia would entail 
an immense lot of work and the outcome, still 
approximate, would hardly justify the time of 

myself and others (whose help would be indis- 
pensable) spent on it. On the several Japanese 
expeditions in Malesia hardly any data are known 
to me. 

It is for this reason that I will restrict myself to 
a recapitulation of the larger units. 

Recapitulation: based on more approximate figures than the original one in the Cyclopaedia on p. cxii 
(with correction): 






sg. km 

to 1950 

to 1972 

index 1950 

inde.x 1972 

1. Sumatra 






2. Malay Peninsula 






3. Java 






4. Lesser Sunda Is. 






5. Borneo 






6. Philippine Is. 






7. Celebes 






8. Moluccas 






9. New Guinea 






Grand total 






Within the areas some details are given below: 
Sumatra. In W. Sumatra (W. Meijer a.o.), N. 
Sumatra (W. J. J. O. de Wilde a.o.), S. Sumatra 
(M. JACOB.S a.o.). 

Malay Peninsula & Singapore. Among the 
numerous new collections those on G. Tahan 
must be mentioned. 

Java. Especially the extreme SW. part, Udjon 
Kulon and P. Peutjang ( Mccuwcn I.) by 
several collectors; by HofXiiRwiRp in P. 
Bawcan and extreme H. Java (G. Baluran, Mcru 
Bctiri, f*urwo HIambangan); Nusa Barung 
(Rappard); Christmas Island (MrrciiELi.). 
I^esser Sunda Islands. In Bali, Lombok, Roti & 
Lomblcn, Komodo. Very large collections in 

Florcs (ScHMUTZ), Sumbawa (Kostermans), 
and Timor. 

Borneo. In Sabah. Sarawak, and Brunei 
enormous collections have been made, several 
in series of the forestry services, but also by 
others. Ml Kinabalu in Sabah has been the aim 
of numerous expeditions, amongst which those 
of the Royal Society. 

In Indonesian Borneo (Kalimantan) in the SE. 
part near Bandjcrmasin. 

Philippine Islands. Among the numerous 
collections many in the PNH series. Big collec- 
tion in the Batan & Camiguin (Balanes Is.) by 




Flora Malesiana 

[ser. I, vol. 8^ 

Celebes. SW. Peninsula, some thousands. 
Moluccas. In total some five thousands, in- 
cluding a big collection from Morotai (Koster- 


New Guinea and neighbouring islands. In the W. 

part (Irian) some 10.000, collected by ex- 
peditions and in the BW series. In the E. part 
(Papua, Terr, of New Guinea, Bismarck 
Archipelago etc.) 100.000 collections, including 
many in the NGF and other series. 



Cycl. Fl. Mai. I, 1, 1950, p. cxvii-cxxv, and Suppl. I.e. 5, 1958, p. ccxlv: 


1. Published sources 


Sachse, J. D. W., Verzeichniss von Bildnissen 
von Aerzten und Naturforschern seit den alt est en 
bis auf unsere Zeiten, mit Biographien. 1. Heft 
(Schwerin 1847) {n.v.). 

MusGRAVE, A., Bibliography of Australian 
Entomology 1775-1930 with Biographical Notes 
on Authors and Collectors (R. Zool. Soc. N.S.W. 
1932). Sydney 1932. 

HuLTEN, E., History of Botanical Exploration in 
Alaska and Yukon Territories from the time of their 
Discovery to 1940 (Bot. Not. 1940, p. 289-346, 

List of collectors with extensive data on the 
Alaska itinerary, on p. 294-343. 

Alden, R. H. & J. D. Ifft, Early Naturalists in 
the Far West (Occ. Pap. Calif. Ac. Sc. no 20, 1943, 
p. 1-59, incl. selected bibliogr.). 

Nissen, Claus, Die botanische Buchillustration, 
Geschichte und Bibliographie. Stuttgart 1951. 

The bibliography in vol. 2, gives extensive data 
on draughtsmen, printers etc. of works containing 
botanical illustrations. 

Buck, Peter H., Explorers of the Pacific. 
European and American Discoveries in Polynesia 
(Bern. P. Bish. Mus. Spec. Publ. 43, 1953, 125 pp., 
with ill.). 

Deals with the voyages proper; for the botanical 
results hardly of interest. 

Whittell, H. M., The Literature of Australian 
Birds. A History and Bibliography of Australian 
Ornithology. Perth 1954. 

With biographies, voyages, etc. 

Streit, Pater R., ctd by Pater J. Dindinger, 
Bibliotheca Missionum. Vol. 21, Missionsliteratur 
von Australien und Ozeanien 1525-1950, Freiburg, 
Internat. Inst. f. Missionswiss. Forschung. 1955. 

Hansen, A., Den danske botaniske litteratur, 
Bibliographia Botanica Danica, 1940-59 med 
tilhorende forfatter-biografier. Kobenhavn 1963. 

Prestwich, a. a., / name this Parrot. Eden- 
bridge, Kent, 1963. 

Steenis, C. G. G. J. VAN, Reaping the Harvest. 
Retrieval of Names and Identifications by means of 
Identification and Collection Lists (Fl. Mai. Bull. 
no 26, 1972, p. 2020-2037). 

Also includes Personal collection lists or field 
books, and list of serial collection lists. 

Sternstein, Larry & Carl Springer, An 
annotated bibliography of material concerning 
Southeast Asia from Petermann's Geographische 
Mitteilungen 1855-1966. The Siam Society 1967. 

Also includes the whole of Malesia as accepted 
in the Flora Malesiana. 

Chaudri, M. N., I. H. Vegter & C. M. de Wal, 
Index Herbariorum. Pt II (3) I-L (Regn. Veg. vol. 
86). Utrecht 1972, ed. by Frans A. Stafleu. 
Guide to the location and contents of the World's 
Public Herbaria. 


1. Sumatra and neighbouring islands 

Cyclopaedia p. cxxii add: 

W. Marsden's History etc. is accompanied by a 
folio vol. of 19 pi. + map. 

The plates consist of mostly useful plants, 
animals, etc. 

6. Philippine Islands 

Cyclopaedia p. cxxiv add : 

Quisumbing, E., Botanical Expeditions in the 

Philippines (Proc. 8th Pac. Sc. Congr. 1953, vol. 4 
Bot. 1957, p. 501-542, map). 

Mendoza, D. R., Philippine Orchid Collectors 
(Philip. Orch. Rev. 7, 1959, p. 17-36, 11 portr.). 

History, collectors, bibliography of papers 
dealing with Philippine orchids. 

Quisumbing, E., Botanical Explorations in the 
Philippines (Philip. Geogr. J. 8, 1964, p. 21-38). 

Chronology from 1588-1901, followed by an 
alphabetical list of botanists and/or plant col- 
lectors during the American regime and under the 
Republic of the Philippines. 



Cyclopaedia of collectors — Supplement II 

9. New Guinea and neighbouring islands 

Cyclopaedia p. cxxv add: 

WONIERSLEY, J. S., A brief History of Botanical 
Exploration of Papua and New Guinea (Papua & 

New Guinea Gazette 8, no 2, Oct. 1953, p. 32-39). 

VlNK, W., Botanical Exploration of the Arfak 
Mts (Nova Guinea, Bot. 22, 1965, p. 471-494). 

Review of former botanical explorers and their 
reports, incl. those by Vink and Sleumer. 



Cycl. Fl. Mai. I, 1, 1950, p. cxxvi-cxli, and Suppl. I.e. 5, 1958, p. ccxlv-ccxlvii, add: 

1. General 

Rau, M. A., Herbaria and Botanical Museums 
in the U.S.S.R. (Bull. Bot. Surv. India 6, 1964, 
p. 73-76). 

Miller, Hortense S., The Herbarium of 
Aylmer Bourke Lambert. Notes on its Acquisition, 
Dispersal, and present whereabouts (Taxon 19, 
1970, p. 489-553). 

2. Special^ 

Berlin, Germany 

Edinburgh, Scotland 

Herb. Berlin-Dahlem (Berolinense). 

Hiepko, p., Herbarium Willdenow Alphabetical 
Index (Interdocumentation Cy AG Zug, Switzer- 
land, xvii pp). 

An introduction to this Index, made from the 
handwritten original by L. Krug, with corrections, 
and an English introduction, has been reproduced 
on microfiches by H. L. de Mink. 

Caen, France 

Herb. Caen. 

Hartog, C. den. The Herbarium of the Institut 
botanique de la Faculte des Sciences, Caen (Fl. 
Mai. Bull, no 20, 1965, p. 1270-1271). 

*Cork, Eire 

•Herb. Cork University. 

Parks, H. M., Some notes on the Herbarium of 
University College, Cork (Irish Nat. J. 11, 1953, 
p. 102-106) in.v.). 

Dehra Dun, India 

Herb. Forest Research Institute. 

Raizada, M. B., The Dehra Dun Herbarium: its 
Origin, History and Mission (J. Sen Mem. Vol. 
Bot. Soc. Beng. 1969, p. 487^98). 

Especially dealing with the India collections. 
Much information on early superintendents. 

•Dublin, Eire 

•.Nat. Mus. Dublin. 

Johnson, T. & Miss M. C. Knowlh^, The 
Levinge Herbarium (Sc. Proc. R. Dublin Soc. n.s. 
10. 1'>K)3. p. 122-132). 

Private fern herbarium from India, Ceylon, etc. 
was left by will in 1896. No mention of Malcsian 
collections is made. 

Herb. Royal Botanic Garden. 

Hedge, I. C. & J. M. Lamond (ed.). Index of 
Collectors in the Edinburgh Herbarium (Edinburgh 
1970, 147 pp.). 

The book includes general references, collectors 
in geographical areas, cryptogamic collectors, and 
a Main list of collectors. 

The foreign herbarium of the Glasgow University 
was incorporated in the general herbarium on 
permanent loan, in 1965. In 1966 the collection of 
Hull University (50(X) specimens, mostly from 
New Caledonia) was added. 

•University Herbarium. 

The entire foreign herbarium (excl. cryptogams) 
has been incorporated in the general herbarium 
of the Royal Botanic Garden, Edinburgh, in 1965. 

♦Glasgow, Scotland 

♦Herb. R. Coll. of Science and Technology. 

Lloyd, B., The Herbarium of the Royal College 
of Science and Technology (Glasgow Naturalist 18, 
1964, p. 363-368). 

Includes a.o. duplicates from Wallich, Cuming, 
Buchanan, Sinclair, etc. 

•Hull, England 

•Herb. Hull University. 

In 1966 the collection was added to the Herbar- 
ium at Edinburgh (see there). 

•Liverpool, England 

•Herb. Mus. Liverpool. 

Handbook 6c Guide to the herbarium collections 
in the public museums, Liverpool. Liverpool 1935. 

This beautiful handbook produced by Mr H. 
Stansfield enumerates the more important 
collectors who contributed to the Museum col- 
lections. It givcsbiographics. portraits, photographs 
of their herbarium specimens, and references. 

(I) New herbarium entries arc provided with an asterisk. 


Flora Malesiana 

[ser. I, vol. 81 

London, England 

Herb. Brit. Museum (Nat. Hist.). 

Dandy, J. E., The Shane Herbarium etc. Add: 
London 1958. 

A beautiful work with extensive information 
and a collection of facsimiles of handwritings of 
the contributors to that Herbarium. 

Manila, Philippines 

Philippine National Herbarium. 

In 1964 the number of mounted specimens 
totalled 88,000. These were acquired only after the 
war through new botanical collections, exchanges, 
donations, and purchases. 

Pre-war loans of mounted botanical specimens 
by the former herbarium of the defunct Bureau of 
Science to the Arbold Arboretum, totalling 3299 
with 134 types, were returned to the P.N.H. by the 
Harvard University Herbarium. 

The handwritten register of the F.B. collections 
was saved and is now kept at the Institute of Wood 
Technology, F.P.R.I. College, Laguna. It contains 
identifications, collectors' names, localities, etc. 
(1903-1940). The pertaining herbarium specimens 
were destroyed during World War II, but dupli- 
cates have been distributed. Efforts are made to 
have the data copied and distributed. 

Montpellier, France 

Herb. Inst. Bot. de rUniversite. 

Granel de Solignac, L. & Mile L. Bertrand, 
Les herbiers deVinstitut de botanique de Montpellier 

(Naturalia Monspeliensia Ser. Bot. fasc. 18, 1967, 
p. 271-292). 

Includes Herb. Cambessedes and numerous other 
famous collections. It contains plants collected by 


Oxford, England 

Herb. Oxford University. 

Druce, G. C. & S. H. Vines, The Dillenian 
Herbaria, an account of the Dillenian collections 
in the herbarium of the University of Oxford. 
Oxford 1907. 

Clokie, H. Newman, An account of the Herbaria 
of the Department of Botany in the University of 
Oxford. Oxford 1964. 

Sandakan, Sabah (North Borneo) 

Herb. Forest Department. 

Jan. 31, 1961, unfortunately the mounted 
collections were all destroyed by fire. A new 
collection has been built up. 

Meijer, W., The History and Prospects of 
Forest Botany in Sabah. Forest Dept. 1964. 

Includes information on the Herbarium. 

*Taipei, Taiwan 

*Herb. Bot. Dept. Taiwan University. 

See under Taihoku, Fl. Mai. I, 1, 1950, p. cxl. 

*Herb. Taiwan Forestry Research Institute. 

Includes some Malesian collections made by 
Japanese botanists. 


Cycl. Fl. Mai. I, 1, 1950, cxli-cxlii, and Suppl. I.e. 5, 1958, p. ccxlvii-ccxlviii add: 

GowER, W. H., Herb. Edinburgh, and Kew. 
Greville, R. K., Herb. Edinburgh (also ex 

Lambert, A. B., also in Herb. Edinburgh and 

Florence (with Webb). 
Lehmann, was sold in 1861 (he lived in 


Leveille, H., Herb. Edinburgh (acq. 1919). 
Menzies, A., bequeathed to Edinburgh in 

Moore, Thomas, fern herb, in Kew; dupl. in 

Herb. Edinburgh. 
Seringe, N. C, Herb. Edinburgh (also ex 







Cycl. FI. Mai. 1, 1, 1950, p. 3-4, and Suppl. I.e. 5, 1958, p. ccxlix: 

1. Abbreviations of Herbaria and collecting series To the name changes add: 


See the list by M. Jacobs, Collections cited under Malesia = formerly Malaysia in the Cyclopaedia. 

Abbreviations (Fl. Mai. Bull, no 22, 1968, p. 1571- ^^ the latter name was officially adopted for the 

1578, 1816). newfederationof the Malay Peninsula, Sarawak, 

^ ^ . . , J uu • ..• ^"'l North Borneo, in Flora Malesiana the 

4. Geographical names and abbreviations name 'Malesia' is now used for the geographic 

Add: delimitation of its area. 

F.M.S. = Federated Malay States 

F.R. = Forest Reserve Saba/j = (Br.) North Borneo. 

[7] VII 


•Abbe, Ernst Cleveland 

(1905, Washington, D.C., U.S.A.; x), botanist, 
educated at Cornell University (B.S. 1928; M.S. 
1930), and Harvard University (M.S. 1931; Ph.D. 
1933); from 1935 to the present on the staff of the 
Botany Dept, University of Minnesota, Minne- 
apolis, U.S.A.; from 1959-1962 National Science 
Foundation Fellow, studying Fagaceae and 
Myricaceae especially in SE. Asia. He is usually 
assisted by his wife, L. B. Abbe (see there). 

Collecting localities. From Sept. 1959- 
June I960} Malay Peninsula: Mersing, Ulu 
Gombok F.R., Kuala Lumpur, Eraser's Hill, 
Cameron Highlands; several places in Thailand, 
Cambodia, and Vietnam; Malay Peninsula: 
Temcrloh, Quantan, Jerantut; Sarawak and A^. 
Borneo: Kuching, Brunei, Mt Kinabalu; Philip- 
pines, Luzon: Laguna. — 1962. Early in the year 
on Mt Kinabalu in A". Borneo with W. Meijer. 
E. New Guinea: Mt Otto (June). — 1964. Sarawak 

Collections. The joint collections of Pro- 
fessor and Mrs Abbe will be deposited in: Herb. 
Kew, Singapore, Harvard Univ., Thai Forest Dept, 
Univ. of Minnesota: woods to the Yale Record 
Collection, and the Nat. Mus. Washington, D.C. 
In the Malay Peninsula about 1(X) nos were 
collected with Kadim bin Tassim. 

Literature. (1) Cf. Ann. Rep. Bot. Gard. 
Dept. for 1959, Singapore 1960, p. 4, 7, 11. 

Biographical data. In Who's who in 
America vol. 28. 

•Abbe, Lucy Boothroyd 

(1906, Ithaca, N.Y., U.S.A.; x), biologist, 
educated at Cornell University (A.B. 1928; M.S. 
1930); since 1944 on Biology Dept Staff, Macalester 
College, St. Paul, Minnesota; since 1950 Assistant 
Professor of Biology. Married to Prof. Ernst C. 
Abbe (sec there for Itinerary, Collections, etc.). 

Addison, George Henry 

retired April 22. 1959 (cf. Fl. Mai. Bull, no 14, 
1959, p. 620), and repatriated. 

Adalbert, Albert George Ludwig 
(d. 1972, The Hague, Netherlands). 
For several years he was a botany teacher in a 

secondary school and was later attached to the 

School Museum, both in The Hague. 

Biographical data. Fl. Mal. Bull, no 26, 

1972, p. 1993. 

•.\Kim bin Ambullah, sec Forest Dept North 

•Agra, Mclccio, sec Philippine National Her- 

•Ahmad bin Shukor 

collected vMih \)r R. Hni. in Johorc, 1970: in 

797/ with E. A. Heaslett, and with T. C. Whit- 
more. Herb. Sing. 

Alcasid, ? G.L. 

collected with Edano as early as 1939. 

*.\ldos, B., see Phillippine National Herbarium. 

Aleqaen, Venancio 

was later Assistant Professor at Ilo College, 
University of the Philippines. 

Allen, B. E. G. Molesworth 

Author of papers on ferns, a regular contributor 
to the Malayan Naturalist Journal, and author of 
a book on Malayan Fruits (1967). 

She left Malaya on retirement in June 1963. 

Collecting localities. Md: 1959 . Malaya: 
Ipoh Distr., Perak (with Kadim): G. Bujong 
Malacca (Aug. 10), Bt Batu Suloh via Kinta 
Forest (12), G. Tempurong (13), Oversea Kubor 
Chinese Tin Area (15). 

Allen, Edgar Francis 

Add: Together with his wife he collected living 
plants on Mt Kinabalu in Jan.-Feb. 1966. They 
were brought back to England (at least partly in 
Kew Gardens).^ 

Literature. (1) E. F. Allen: 'A mountain 
hunt yields garden treasures' (Straits Times 
Annual 1973, p. 146-147, photogr.). 

•Alphonso, A. George 

obtained the Kew diploma in 1956 after a two 
years' study course at Kew Gardens. In March 
1957 appointed Curator of the Botanic Gardens 
Singapore. He has a special interest in orchids. 

Collecting localities. 1959. N. Borneo 
(Nov. 14-Dec. 3): Crocker Range, near Tenom, 
Tambunan and Patau, on the Keningau Plateau, 
on the Baiaio and Biah Rivers; Sarawak: at 
Bau. — 1960. Langkawi Is. (on limestone; Nov.- 
Dec.).— 7962. Sarawak (May 30-June 22): Niah, 
Undup R., and Batu Lintang. — 1963. Malaya. — 
1965. Malaya: Trengganu. — 1966. N. Borneo: Mt 
Kinabalu (Sept. 13-30). Possibly many times more. 

Collections. In Herb. Sing.: 84 nos from N. 
Borneo (1959): dupl. Herb. Edinb. Mostly col- 
lecting living plants. 

Alston, Arthur Hugh Garfit 

Collections. Add Java nos 15378-15576; 
Borneo nos 13029-13462 (instead of 13576-15377); 
Sumatra nos 13576-15377. 

Biographical data. Brit. Fern Gaz. 8, 
1958, p. 220-221; Amcr. Fern J. 49, 1959, p. 1-2; 
Taxon 8. 1959. p. 83-86. portr.; J. Soc. Bibliogr. 
Nat. Hist. 3. 1960. p. 383-404 (bibliogr., list of 
new taxa, nomcncl. changes). 

•Altona, Th. 

of the Dutch East Indian Forest Ser\'ice col- 
lected at Rcmbang, Java (1922), in the scries of 

(t) New entries are provided with an asterisk. 



A. M. 

Flora Malesiana 

[ser. I, vol. 8^ 

Beumee. He had no separate entry in the Cyclo- 
paedia (Fl. Mai. I, 1, 1950). 

♦A.M., see A. Menzies. 

*Ampuria, see Forest Dept North Borneo. 


(d. Oct. 4, 1952, Bogor, Indonesia). 

Anderson, James Aidan Robb 

got his Ph.D. at the University of Edinburgh 
(1961, in absentia), on a thesis dealing with the peat 
swamp forests of Sarawak and Brunei.^ 

He was on leave in spring 1971 and would 
return in the middle of the year for a final short 
tour before his retirement. He settled at Edinburgh 
and acts as a consultant to forestry services and 
timber companies. He is writing a checklist of 
Sarawak forest trees. 

Jarandersonia Kosterm. and Goniothalamiis 
andersonii J. Sinclair were named after him. 

Collecting localities. NW. Borneo add: 
G. Mulu and Api up the Baram River (June-July 
1960), limestone hills; on the principal limestone 
hills in Sarawak. Terr, of New Guinea: Mt Wilhelm 
(Sept.). — 1964. Sarawak: 2-weeks' trip to the 
Kelabit Highlands, based at Bario (June 19 
onwards), collecting > 200 nos; G. Selabor, 
Serian Distr.— 7959. With Ilias bin Paie to Ulu 
Melinau, Hose Mts, Mt Murud; Mt Batu Tibang 
(Centr. Borneo) (June). ^ — 7970. Kalimantan (Indon. 
Borneo) : Kutai Nature Reserve with a small party 
of BiOTROP, Indonesia (exploring 6 weeks in May 

Collections. His series should be cited with 
the prefix 'Si', with the date of collecting, but the 
stamped number on the label should be discarded.^ 
Dupl. also in Herb. Edinb. The Mt Wilhelm plants 
numbered 1-22. 

Literature (1) Author of 'Field Check List 
of Trees of Peat Swamp Forests of Sarawak' 
(mimeogr., 4 + 18 pp.); 'The Flora of the Peat 
Swamp Forests of Sarawak and Brunei etc." 
(Gard. Bull. Sing. 20, 1963, p. 131-228, 10 pi., 3 
tabl., 1 map). 

(2) Cf. FI. Mai. Bull, no 25, 1971, p. 1890. 

(3) Cf. Fl. Mai. Bull, no 21, 1966, p. 1434. 

Anderson, Thomas (add:) A. 

Collections. Java and Singapore 
(1861) in Herb. Edinb. 


Anderson, William 

(d. 1778). 

Biographical data. Add: In H. M. 
Whittell: 'The Literature of Australian Birds' 
1954, pt 2, p. 12-13. 


of Osaka City University, spent 5 months (Oct. 
1968-M.a.r. 1969) in many areas of the state Sabah 
(N. Borneo), with Koka wa and Hotta (see there). 
Special attention was given to Tawau Hills and 
Silabukan in the south and to Kinabalu and Trus 
Madi on the west coast . 

Andrews, Charles William 

Collections. Dupl. Christmas I. (Indian 
Ocean) 7597 in Herb. Edinb. 

*Ang Gek Choo 

of the Singapore Herbarium, collected in 
Malaya at least in 1963, and from Feb. 28-Mar. 6, 
1965, at Penang Hill and on Maxwell's Hill in 
Perak (181 nos), and from Oct. 5-16 at Penang 
Hill and in the Cameron Highlands (few numbers). 
Presumably in Herb. Sing. 

*Angian, see Forest Dept North Borneo. 

Anonuevo, Prudencia 

(1926, Bucalbucalan, Sorsogon, Luzon, P.I.; x) 
was employed in the National Museum at Manila 
from Aug. 1947 to Feb. 1951. 

Collecting localities. 1947-51. Philip- 
pines. Luzon: Abra and Mountain Province; 
Palawan; Mindanao: Mt Apo, Cotabato; Basilan. 

Collections. In PNH Manila. 

♦Antonio, Engl-Silv. F. 

collected grasses in Portug. Timor from Oct.- 
Dec. 7967. In Herb. Lisbon. 

*ANU nos^ 

a series of the Australian National University, 
made by staff and students working under Dr 
Donald Walker in the Department of Geography 
of the Institute of Advanced Studies. Apart from 
Dr and Mrs (Pat) Walker*, collectors are: John 
Flenley*, Josselyn Wheeler*, Dr Donald 
MacVean*, and L. Keith Wade* in New Guinea, 
and Martin Kellman* in Mindanao (Philippines). 
See under the separate entries. 

Appelman, Frederik Johannes 

(d. 1965, The Hague, Netherlands). 

Biographical data. Blijdorp Geluiden 13, 
no 4, Apr. 1965, p. 2-3, with portr. ; Panda Nieuws 
1, no 8, Apr. 1965, p. 1; Vakbl. Biol. 45, 1965, p. 
74-76; Meded. Ned. Com. Int. Nat. Besch. 19, 
1965, p. 13-25, with portr. a. bibliogr. 

Aranez, Adoracion T. 

Student, taking a Master's degree in Botany; 
Instructor of Botany in the University of the East. 

*Ardley, John H. 

Entomologist with the Papua and New Guinea 
Department of Agriculture; subsequently Tech- 
nical Adviser-Entomology with William Cooper & 
Nephews, PTY. Ltd. at Concord, N.S.W., 

Collections. Mainly SdSZ.v/aktl(New Guinea) 
plants associated with insect pests or food plants of 
insects. Partly numbered serially under collector's 
name, others without number; in Herb. Lae. New 
Zealand plants in Herb. Domin. Mas. Wellington. 

*Ardzi bin Arshid, see Forest Dept Sarawak. 


Collections. In the Leyden Herbarium 
archives a list was found of some (88) rare dried 

(1) An asterisk refers to a separate entry with more particulars. 



Cyclopaedia of collectors — Supplement II 


plants from Java, which were packed in a case 
addressed to Mr Fritze (Dr A. E. Fritze, Head 
of the Medical Service in the D.E.I.), and later 
received by Blume; presumably in Herb. Leyden, 
dating from the first half of the 19th century. 

*.\riffin, see Forest Dept Sarawak. 

Annit, William Edington de Margrat 

When in Goodenough I. (May/June 1894) he 
visited the summit of Mt Oiamadawa'a (c/. Am. 
Mus. Novit. no 1792, p. 7). 

Arnold, Joseph 

Add: In 1815 he sailed from Australia homeward 
bound in the "Indefatigable". The ship touched 
Batavia (Sept. 3), and burnt out on Oct. 22 in the 
harbour. In this way Arnold was forced to stay 
in Java^ for more than 3 months before having the 
opportunity of a passage to England in the 
"Hop>e". He botanized a.o. at Ciceroa (= Tjiserua) 
and visited G. Tangkuban Prahu. The living plants 
he brought from Java for Sir J. Banks were dead 
on arrival (by rats, seawater, etc.). He evidently 
drew some Java plants. 

He left England for Sumatra in Nov. 1817. 

Arnold's letters from Sumatra are published 
by Dr J. Bastin.^ 

Raffles stated in a letter to Dawson Turner, 
dated 12 April 1820, that Arnold's collections in 
Sumatra were altogether unimportant and that the 
few plants were examined by Dr Horsheld who 
did not think them of much interest. They were 
forwarded with the rest of their collections to Mr 
Brown, i.e. Herb. Brit. Mus. (Dr J. Bastin /./. 28 
Nov. 1972). 

Collections. The collections of shells etc. 
were apparently sold at auction by the Linnean 
Society on 10 Nov. 1863; present whereabouts 
unknown. A copy of their description is in Mitchell 
Library, Sydney (Dr J. Bastin in litt., July 1972). 

Literature. (2) His Java Journal will in due 
time be published by Dr J. Bastin (London). See 
J. Mai. Br.R.As. Soc. 46 (1973), pt. 1 , 92 pp., 1 1 pi. 

f3) J. Soc. Bibliogr. Nat. Hist. 6 (1973) p. 305- 
372,4 pi. (1 portr.). 

Biographical data. Add: Portr. in Str. 
Times Annual for 1971, p. 60. 

*Asah anak Unyong 

Collector for the Forest Dept Brunei, at least in 

Ashton, Peter Shaw 

Ph.D. Cambridge (1962). From Oct. 1962 
Forest Botanist at Kuching, Sarawak. In 1966 
appointed Lecturer in Tropical Botany at the 
University of Aberdeen. 

Author of papers on Dipterocarpaceae, and the 
forests of Brunei and Sarawak, and Tree Manuals. 

Knenia ashtonii Sinclair was named after him. 

Collecting localities. Add: From 1963 
onwards in Sarawak. 1963. 6 weeks' trip to the 
Similajau Forest Reserve, Bintulu, across the 
watershed to Ulu Bclaga and Sibu (Mar.-Apr.); 
10 days in Bako National Park (May)'; 4 weeks in 
the Tatau-Mukah area (Scpt.-Oct.).— /964. To Bt. 
Kajang and the Ulu Mujong part of the Hose Mts 

(7 weeks, from Mar. 10 onwards, collecting c. 800 
nos); Santubong (2| weeks from May 29 onwards); 
Lambir Hills (2nd half of June, coll. 150 nos); 
accompanying T. D. Pennington (see there) for a 
week in Nov. — 7965. Ulu Dapoi, Tinjar (Mar.- 
Apr.), plateau of Usun Apau (coll. 600 nos)\ 
Ulu Temburong (in Brunei) for 3 weeks; Bako 
National Park (July-Aug.). 

Collections. Mostly palms and dipterocarps 
in 1963. Also dupl. in Edinb. Herb. 

Literature. (1) P. S. Ashton: 'The Plants 
and Vegetation of Bako National Park' (Mai. Nat. 
J. 24, 1971, p. 151-162, map, pi. 53a-58b). 

*Asseni, Johannes van den 

(1930, Rotterdam, Netherlands; x), zoologist 
educated at Leyden University (M.S. 1954); from 
Apr. 1956-Apr. 1959 entomologist of the 'Malaria- 
bestrijding' of the Medical Service in New Guinea. 
Later on the staff of the Zoological Laboratory at 
Leyden. Ph.D. at Leyden (1967). 

Collections. Some tens of weeds collected 
at Ifar, Hollandia, North New Guinea (Oct. 1956); 
in Herb. Manokwari and Leyden. 


= Rd. Atje Adiwidjaja (d., Maseng near Bogor, 

*Awing, James, see Forest Dept North Borneo. 

Backer, Cornelis Andries 

(d. 1963, Heemstede, N.H., Netherlands). 

Biographical data, Vakbl. Biol. 43, 1963, 
p. 65-66; Taxon 12, 1963, p. 173-177, w, portr., 
facsim. handwriting; in Backer & Bakhuizen/., 
Flora of Java 1, 1963, p. xi-xx, w. portr.; Blumea 
12, 1963, p. 1-4, w. portr. 

*Baker, C, see sub M. A. Carne. 

*Bakhuizen van den Brink, Karel Johan 

(1918, Buitenzorg, Java; Feb. 28, 1957, on 
board, off the Cocos Islands, on the voyage to 
Holland), younger brother of R. C. Bakhuizen 
Van Den Brink Jr (see there), lastly Assistant on a 
tea estate S of Buitenzorg. 

He is mentioned on the labels as the collector 
of Cyperaceae, etc., in total > 74 nos in the years 
1922-23, evidently in the wake of his father; 
material in Herb. Bog. 

^Baldemor, J., see Forestry Bureau, Manila. 

*Balfour, Andrew Francis 

lieutenant of the Royal Navy, is recorded as the 
collector of 9 plants from the summit of Ternati 
(5000 ft) in the Philippines, and of a specimen of 
'Elang-Elang' from Manila. The Peak of Tcrnate 
is rightly situated in the Mohtccas. In Herb. Edinb. 
(Trans. & Proc. Bot. Soc. Edinb. 12, 1876, p. Ixi). 

♦BalRooy, Max Michael Josephus van 

(1932, Poerbalingga. Java; x). botanist, educated 
at the University of Indonesia, Banuung( 1952-57). 
and at Leyden University (1958-61), F»h.D. 1971 
on a plantgcographical analysis of the Pacific. 
Since 1961 on the stalf of the Rijksherbarium, 




Flora Malesiana 

[ser. I, vol. 8^ 

He visited New Guinea, Australia, Lord Howe 
L, and Java in 1965. ' In 1971 he attended the 
Pacific Science Congress in Australia, and made a 
collecting trip to Tahiti. 

Itinerary 1952-53. W. Java.— 1965 (Mar. 
30-Aug 14).^ Leaving Holland (Mar. 30), travel- 
ling by way of Sydney and Canberra to Lae, Terr, 
of New Guinea (arriving about Apr. 12); two one- 
day trips with Van Royen to Busu R. and 

Markham R.; Apr. 15 starting from Goroka for 
Keglsugl (f. 2600 m) and from there on foot to 
Mt Wilhelm, basecamp near Lake Aunde (3560 m) 
until July 8; Java. 

Collections.^ 993 New Guinea nos collected 
above 2600 m, mosses included; in Herb. Leyden; 
dupl. elsewhere. 

Several unnumbered specimens mostly from W. 
Java in the educational herbarium of the Univ. of 
Indonesia, Bandung, coll. 1952-53. 

Literature. (1) M. M. J. Van Balgooy: 
'Report of an expedition to New Guinea, Australia 
and Indonesia' (mimeographed, Leiden 1965). 

(2) K. Thomasson: 'Phytoplankton from some 
lakes on Mt Wilhelm, East New Guinea' (Blumea 
15, 1967, p. 285-296). 

Biographical data. Curr. Vitae in Thesis 
Leyden 1971. 


Bally, Walter (F. E.) 

(d. 1959, Geneva, Switzerland). 

Biographical data. Verb. Schweiz. Naturf. 
Ges. 139, 1959, p. 379-384, portr., bibliogr. 

Bangham, Walter Nicholas 

(d. 1954 or 1955, Kisaran, Sumatra East Coast, 
Indonesia). In the fifties Consultant of the Good 
Year Rubber Co. 

Banks, Sir Joseph 

Collections. Also dupl. Cook's 1st Voyage 
in Herb. Edinb. 

Literature. (1) Add: J. C. Beaglehole: 
'The Endeavour Journal of Joseph Banks' (1st ed. 
Sydney 1962, 2 vols). 

(3) Add to 'Illustrations of the Botany e/c.': 
For particulars on the drawings, see Cl. Nissen, 
'Die botanische Buchillustration etc.'' 1951, vol. 2, 
p. 7. 

E. W. Groves: 'Notes on the Botanical Speci- 
mens collected by Banks and Solander on Cook's 
first voyage, together with an Itinerary of Landing 
Localities' (J. Soc. Bibliogr. Nat. Hist. 4, 1962, 
p. 57-62). 

D. J. McGillavry: 'A checklist for the 
illustrations of the botany of Cook's First Voyage' 
(Contr. N.S.W. Nat. Herb. 4, 1970, p. 112-125). 

(5) Add: See also J. Ramsbottom: 'Banks and 
Solander's duplicates' (J. Soc. Bibliogr. Nat. Hist. 
4, 1963, p. 197). 

(6) See further: W. T. Stearn: 'The Botanical 
Results of the Endeavour Voyage' (Endeavour 27, 
1968, p. 305-309, 1 fig., 4 phot.): 'A Royal Society 
Appointment with Venus in 1769. The Voyage of 
Cook and Banks in the Endeavour in 1768-1771 
and its Botanical Results' (Not. & Rec. R. Soc. 
Lond. 24, 1969, p. 64-90, map facing p. 90). 

Biographical data. Add: W. R. Dawson, 
'The Banks Letters. A Calender of the Manuscript 
Correspondence of Sir Joseph Banks etc' (London 
1958); Proc. R. Soc. Queensl. 83, 1972, p. 1-19, 

*Banlugan, G., see Philippine National Herbarium. 

Banzon, Gregoria T. 

Instructor of Zoology, University of the 

Barclay, George 

Literature. (3) Add: P. H. Raven: 'George 
Barclay and the 'California' portion of the Botany 
of the Sulphur' (Aliso 5, 1964, p. 469-477; inch 
list of plants). 

Bartlett, Harley Harris 

(d. 1960, Michigan, U.S.A.). 

Itinerary. 1935. Philippines, add: Mt Iraya, 
Batan I. (July). 

Collections. Philippine collections distrib- 
uted by the University of the Philippines, a.o. to 
Herb. Berkeley. 

Biographical data. Bull. Torr. Bot. Club 
88, 1961, p. 47-56, portr., bibliogr. 

Baskinas, Leticia 

Instructor of Botany, Araneta University; later 
married to a botanical colleague and living in 



Cyclopaedia of collectors — Supplement II 


Batten-Pooll, Arthur Hugh 
(d. 1971 or early 1972, England). 

Baudin, Thomas Nicolas 
Biographical data. In A. A. Prestwich, 

I name this Parrot, 1963, p. 6-7. 

Bauer, Ferdinand Lucas 

Collections. Add: On the whereabouts of 
his original drawings, see Cl. Nissen, 'Die 
botanische Buchiilustration' 1951, vol. 2, p. 9. 

Bayer, Frederick Merkle 

Add (to Suppl. Cycl. Fl. Mai. I, 5, 1958, p. 
cclvii) : 

(1921, Asbury Park, N.J., U.S.A.; x) zoologist, 
B.S. Miami (1948), M.S. Washington (1954), 
Ph.D. 1958. Assistant Director State Museum 
Florida 1942-46; Assistant Marine Lab. Miami, 
1946-47; Assistant Curator (1947-51) and Associ- 
ate Curator (1951-61) of the U.S. National 
Museum; Associate Professor Institute Marine 
Science, Univ. of Miami (1962-63), and Professor 
from 1964- 

He accompanied several expeditions, e.g. to 
Bikini, Palau and the Gulf of Guinea. 

His interests are biological oceanography, 
ecology of coral reefs and taxonomy of Octocorallia, 
Mollusca, and Pterobranchia. 

He occasionally collects ferns. 

•Bebob, Aban, see Forest Dept North Borneo. 

Beccari, Odoardo 

Collections. The type locality Mt Poe (Poi), 
Sarawak, given for several Sarawak plants, is 
according to Dr B. L. BuRrr not the G. Poi used 
as a name on modern maps, but a more south- 
easterly peak in the range, G. Berumput (c/. Fl. 
Mai. Bull, no 19, 1964, p. 1131-1132). Sumatra 
dupl. in Herb. Edinb. 

Literature. (2) Add: For annotations on the 
itinerary cf. W. Vink in Nova Guinea, Bot. 22, 
1965, p. 479, and fig. 6. 

(5) Add: For annotations on the itinerary cf. 
W. Vink in Nova Guinea, Bot. 22, 1965, p. 479- 
481, fig. 6. 

(11) Add: On the drawings, see Cl. Nissen. 
'Die botanische Buchiilustration' 1951, vol. 2, p, 

Becking, Johannus Hendrikus 
(d. 1972, Arnhem, Gld., Netherlands). 
Collections. Add: 20 plants from former 

Dutch New Guinea (Irian), collected in 1961 ; in 

Herb. Wageningen. 

•Behr, Hermann Hans 

(1819, Kothcn, Germany; d. 1904), a medical 
man who was for a long time a resident in South 
Australia. In Dec. 1849 he wrote a letter from 
Manila telling of his plan to stay there and to set 
up as a medical practitioner. Later he settled in 
California. Author of some botanical papers'. 

Aristida behriana I'.v.M. was named in his 

Collections. He made natural history col- 
lections; Australian plants in Herb. Melbourne. 
As yet no Philippine collections known. 


Literature. (1) E.g.: 'Synopsis Genera of 
Vascular Plants San Francisco' (1884). 

Biographical data. Bot. Zeit. 7, 1849, p. 
873; ibid. 8, 1850, p. 335; Dorfler, Bot. Adress- 
buch 1902, p. 240; Britten & Boulger, Biogr. 
Index, 2nd ed. by Rendle, 1931; Index Herb, pt 
2 A-D, 1954. 

Belanger, Charles Paulus 

Collections. Java dupl. (1833, probably 
acq.) in Herb. Edinb. 

Literature. (1) Add: cf. R. Ross in Taxon 
13, 1964, p. 193-196. 

Drawings of the botanical atlas made by Bory 
DE St Vincent and E. Delile. 

Benecke, Franz F. 

collected also in E. Java, Mar.-Apr. 1891; 
dupl. in Vienna. 

Benthem Jutting, W. S. S. van, see sub Feen-Van 
Benthem Jutting, W. S. S. van der. 

*Bergraan, Mrs Dagny 

wife of Sten Bergman, accompanied her 
husband during part of his travels in former 
Dutch New Guinea, from where they returned in 
spring 1959. 

Collecting localities. Former Dutch New 
Guinea. Mostly in Swart Valley at c. 1 600-2000 m 
alt., but also at Steenkool, near Tembuni (Feb.- 
Mar. 1957), and on the S. coast near Asmat (June 

Collections. Herb. State Mus. Stockholm: 
1\1 nos, mostly orchids and ferns, but also mosses*. 

Literature. (1) E. B. Bartram: 'Additions 
to the Moss Flora of Netherlands New Guinea' 
(Svensk Bot. Tidskr. 54, 1960, p. 483^87). 

Bergman, Sten 

Additions to Suppl. Cycl. Fl. Mai. I, 5, 1958, 
p. cclviii: 

Itinerary.' After Beeuw insert: (= Beo). 
Add : Leaving Sorong (Dec. 3) by boat to P. Adi 
on the S. coast of New Guinea by way of Fak-Fak, 
exploring the islet and the opposite mainland near 
Kambala; returning via Fak-Fak, Kokas, and 
Inanwatan to Sorong; returning to Inanwatan 
(leaving Sorong Feb. 5, 1949) for a trip into the 
interior: by canoe to kp. Bahru; Atinjoe, from 
where (15) to Susemok, Fanatof, Aifat River, 
Kotjewir, and returning to Atinjoe (leaving Mar. 
9), Aimaroe (lakes), Teminaboean, Konda, and 
return to Sorong; leaving (Apr. 12) to Jeflio, S of 
Sorong (staying about a month); Sorong; via 
Solol (Salawati) to Batanta /., Wailibit; Sorong, 
leaving June 30 by boat to Manokwari; Ransiki 
and trip into the interior (starting July 13):* Anggi 
Lakes (Anggi Gita), Tombrok, from there (21) to 
Anggi Gigi (Scrorci), and back to Tombrok and 
October bivouac (1500 in); return to Ransiki, by 
boat to Wandammen and Mici, Manokwari and 
Sorong (arriving Oct. 4); to Batanta I. (staying 1 
week); Sorong, leaving (Nov. 2) for Macassar; by 
K.P.M. steamer via Bali to Java, short visit to 
Batavia and Huitenzorg; Singapore and sailing for 
Gotcborg (arriving Jan. 15, 1950). 

Literature, (1) Ciiron. Bot. I.e., is rightly: 
Chron. Nat. 



Flora Malesiana 

[ser. I, vol. 8^ 

Sketch map showing localities visited in 1948- 
49, in Ark. f. Zool. 8, no 2, 1955. 

(3) Add: Transl. from 'Vildar och ParadisfSglar' 
(Stockholm 1950; with route map); 'Paradiesische 
Insel. Urwaldtiere und Steinzeitmenschen in 
Neuguinea' (Wiesbaden 1956, with map) ; 'Through 
primitive New Guinea' (London 1957); 'Mein 
Vater der Kannibale' (Wiesbaden 1961, with map, 
transl. from Swedish). 

(4) For annotations on the route in the Arfak 
Mts see W. Vink in Nova Guinea, Bot. 22, 1965, 
p. 489. 

Biographical data. Portr. in 'Vildar och 
Paradisfaglar' I.e., following p. 192 and 256. 

Bernard, Charles Jean 

(d. 1967, Amsterdam, Netherlands). He was 
active in several international fields, as Nature 
Preservation and the Red Cross. 

Biographical data. F1. Mai. Bull, no 22, 
1968, p. 1515, 1518. 

*Berthe nee Friedberg, Claudine 

(1933, Paris, France; x), studied in Paris and 
was subsequently 'Associee au Laboratoire de 
Botanique Appliquee, Museum d'Histoire Natur- 
elle'. She visited the Lesser Sunda Islands for 
ethnological purposes.^ 

Collecting localities. Lesser Sunda Is- 
lands. 1962. Bali (May-Nov.),^ at Tangenan, 
Kintamani, and in the environs of Sanur. — 1966. 
Timor (May-Oct.),* the central part, both sides of 
the frontier. 

Collections. Several hundreds of plants; 1st 
set in Lab. Ethnobot. Mus. Nat. Hist. Nat. Paris; 
2nd set Timor in Leyden and few dupl. from Bali; 
3rd set in Lisbon. 

Literature. (1) C. Friedberg: 'Les plantes, 
les dieux et les hommes dans I'lle de Bali' (Science 
et Nature no 59, 1963, 3-12). 

C. Friedberg & L. Berthe: 'Note ethnobotan- 
ique sur I'utilisation rituelle de quelques riz 
balinais' (J. Agric. Trop. & Bot. Appl. 10, 1963, 
612-620, 4 pi.). 

(2) Cl. Friedberg: 'D'apr^s les Bunaq de I'lle 
de Timor. Chasse rituelle brulis et herbes a savane' 
(Science de Nature no 92, 1969, p. 23-30, 10 
photogr.); 'Analyse de quelques groupements de 
vegetaux comme introduction a I'etude de la 
classification botanique Bunaq' (Echanges et 
Communications offerts k Cl. Levi-Strauss, p. 
1092-1130, 1 pi.); 'Elements de botanique Bunaq 
receuillis a Lamaknen (Timor central)' (Langues 
et Techniques Nature et Societe II. Approche 
ethnologique. Approche Naturaliste. Ed. Klinck- 
sieck, p. 375-393, fig. 1-9); 'L'agriculture des 
Bunaq de Timor et les conditions d'une equilibre 
avec le milieu' (Journ. Agr. Trop. & Bot. Appl. 
Paris 18, 1971, p. 481-532, map, photogr.); 
'Aper9u sur la classification botanique bunaq 
(Timor central)' (Bull. Soc. Bot. Fr. 118, 1971, 
p. 255-262, 2 schemas). 

Beumee, Johan Gotlieb Benjamin 
(d. 1966, Bennekom, GId., Netherlands). 
Biographical data. F1. Mai. Bull, no 21, 

1966, p. 1371-1372, 1377. 


^Bianchi, Frans 

(1918, Buitenzorg, Java), botanist who studied 
at Amsterdam University. Since Sept. 1946 working 
in the Hugo de Vries Laboratory, Amsterdam 
University; taking his Ph.D. on a genetical 
subject there, and from 1964 Professor of Botany. 

Collections. In August 1938 he collected 
some 3(X) nos on Mt Gedeh-Pangrango in W. 
Java; in Herb. Univ. Amsterdam. 

Billardiere, Jacques Julien Houtou' de la 

Collections. Following up the items in the 
Cycl. Fl. Mai. I, 1, 1950, p. 57-58 and 599, some 
additional data on the history of his collections 
were found in a recently published book*. In 1795 
the senior surviving French officer, the Chevalier 
DE RossEL, embarked with the collections of the 
expedition, in a Dutch ship, with the object of 
placing them at the disposal of Louis XVIII, then 
titular King of France. This ship was captured by 
the British Navy off the Shetland Islands, and the 
collections came into the hands of Sir Joseph 
Banks. Rossel's protest against the seizure of the 
collections resulted in placing them at the disposal 
of Louis XVIII, then in Courland. The latter 
instructed his ambassador to present them to the 
Queen of England. Later, Billardiere's plea with 
Banks resulted in the sending of the lot to France. 

P. W. Webb, who had acquired Billardiere 
plants in 1834, in 1837 wrote Lambert that he was 
sending the latter specimens from New Holland, 
New Ireland, Java, and Amboyna, collector not 
named, but evidently Billardiere. At Lambert's 
sale 2 bundles of plants from Java etc. were bought 
by R. Brown (c/. H. S. Miller in Taxon 19, 1970, 
529); presumably in Herb. Brit. Mus. now. 

Literature. (4) Add: Cf. Sir Gavin de 
Beer: 'The Sciences were never at War' (1960) p. 

(9) Add: The spelling 'Houtou' is authentic 
according to Chevalier. 

Biographical data. F. A. Stafleu: 'Adan- 
son, Labillardiere, de CandoUe, Introductions to 
four of their books' (in the series Historiae 
naturalis classica, J. Cramer 1967, p. 16-63). 

Biro, Lajos 

Collections. Phanerogams and pterido- 
phytes in the Herb. Gen. Ungarian Nat. Mus.; 
identified by Prof. E. Gombocz. 

*Bisset, Norman Grainger 

(1925, Glasgow, Scotland; x) was educated in 
chemistry at Imperial College, University of 
London (M.Sc. organic chem. 1952; Ph.D. 1968). 
Ahli khimia at the Treub Laboratory, Bogor 
Botanic Gardens, 1954-58; Colombo Plan expert 
in chemistry, Malaysia, 1958-62; plant chemist 
at the C.N.R.S. Institut de Chimie des Substances 
Naturelles, Gif-sur-Yvette, France, 1962-67; 
lecturing in pharmacognosy, Dept of Pharmacy, 
Chelsea College, University of London, since 1967. 

In 1964 he carried out a 4 months' mission for 
the C.N.R.S. collecting dammar samples and 
plants of potential medicinal interest for chemical 
studies by various French institutes^ 

Author of numerous phytochemical papers, 
recently especially on the genus Strychnos. 

Collecting localities. 1956. S. Moluc- 
cas.^ With J. VAN BoRSSUM Waalkes (itinerary see 



Cyclopaedia of collectors — Supplement II 


there, Suppl. Cycl. Fl. Mai. I, 5, 1958, p. cclxi). 
W. Java: Rantjabadak and Pelabuhan Ratu. — 
1957. iV. Java: G. Pulusari, near Menes, Tjipining, 
Tjipunaga, Tjigarondong. — 1960-62. Malaysia, 
occasional coll. mostly of dart-poison plants from 
various parts. ^ — 1964.^ Many localities in Sabah 
(mostly with W. Meijer), Sarawak (partly with 
J. A. R. Anderson, P. S. Ashton, and Ilias Paie), 
and Malaysia (mostly with Kochummen). 

Collections. Emphasis on plants of reputed 
or potential medicinal interest. S. Moluccan 
collections numbered in his personal series (570- 
748), but also in that of Van Borssum Waalkes or 
Pleyte. W. Java and Malaysia plants (749-783) 
in BO, K, or L. A few Malaysian collections in the 
KL series. The 1964 collection partly in his 
personal series (784-808), and also in the KEP, 
SAN, and SAR institutional series. 

Literature. (1) Rapport de mission en 
Extreme-Orient du 9 avril au 6 septembre 1964, 
pp. 28. 

(2) N. G. Bisset: 'A phytochemical survey of 
some plants from the South Moluccas' (Phytochem. 
Symp. Kuala Lumpur 1957, p. 125-140). 

(3) N. G. Bisset: 'The arrow and dart poisons of 
South-East Asia with particular reference to the 
Strychnos species in them' (Lloydia 29, 1966, p. 
1-18, with M. C. Woods p. 172-195). 

♦Black, Michael 

Collecting localities. ? 1965. New 
Guinea. — 1968. Malay Peninsula and Java with 
P. J. B. Woods (see there). New Guinea with 
Womersley, Ridsdale, and P. J. B. Woods. 

Collections. Herb. Edinburgh. 

Blanco, Francisco Manuel 

Literature. (1) Add: For the drawings of the 
3rd ed. of the Flora, see Cl. Nissen, 'Die botanis- 
che Buchillustration' 1951, vol. 2, p. 16. 

Bland, Laura Shelford 

collected as early as 1902 in Malaya at Alor 
Gadjah (Aug. 23). 

•Blaxland, G. D., see sub L. H. DEN>as & G. D. 

Blume, Carl Ludwig 

Collections. Delete: Herb. Oxford Univ. 
Dupl. from Ambon (certainly not collected by 
himself) and Java in Herb. Edinh. 

In 1833 J. Frank of The Hague, acting as a 
Lambert's agent, sent him a parcel of Blume's 
plants from Java {cf. Taxon 19, 1970, p. 515). This 
lot was later bought by Pamplin at the sale of L.'s 
herbarium; subsequent history unknown. 

Literature. (2) Add: For particulars on the 
drawings of his publications sec Cl. Nissen, 'Die 
botanischc Buchillustration' 1951, vol. 2, p. 17. 

liocdijn, Karcl licrnhard 

(d. 1964. The Hague, Netherlands). 

Biographical data. Vakbl. Biol. 44, 1964, 
p. 191; Pcrsoonia 3, 1965. p. 325-330, pi. 4, 
bibliogr.; Fl. Mai. Bull, no 20, 1965, p. 1237. 

•BorIc, A. Linn 
Ph.D. Minnesota; Field Research Assistant to 


Prof. Abbe (see there). In Asia he was accompanied 
by his wife (M. P. Bogle) at her on^ti expense, 
except for the Kinabalu trip, for which NSF paid 
her expenses as well. 

Collecting localities. 1961. India (Sept. 
21-Oct. 6; Oct. 31-Dec. 12), Burma (Dec. 18- 
21). — 1962. Malaya: Cameron Highlands (Jan. 
13); Sabah: Mt. Kinabalu (Feb. 2-23) with Abbe, 
Kaul, and W. Meijer. 

Collections. Dupl. in Herb. Edinb.; main 
collection probably in Minnesota. Primarily 
Fagaceae for Prof. Abbe were collected, but also 
material for a thesis. 

*Bojeng bin Sitam, see Forest Dept Sarawak. 

Boorsma, Simon Eliza 
(d. 1959, Amsterdam, Netherlands). 

Boorsma, Willem Gerbrand 

Biographical data. Add: Pharm. Tijdschr. 
N. I. 1, 1924, p. 137; Pharm. Wkbl. Ned. 74, 1937, 
p. 331; Voeding 27, 1966, p. 167-170, portr. 

Booth, H. or J. 

As yet there is no certainty on the Java collector 
of that name. See also sub J. Th. Booth. 

A certain John Booth of the Flottbecker 
Baumschulen ( = nursery) near Hamburg, Germany, 
built a new orchid house in 1844 (Bot. Zeit. 3, 
1845, p. 48). In 1850 a catalogue of the firm of 
James Booth & Sohne offered many orchids for 
sale (Bot. Zeit. 9, 1851, p. 240). In 1861/62 the 
orchid collection of the firm was sold to England 
and they started growing water plants (Hamb. 
Gart. Blum. Zeitschr. 18, 1862, p. 433^36). 
Possibly a son of the family collected in Java for 
somebody in England. 

♦Booth, Jonas Thomas 

(b. 1829; d. ?), who preferred to be known as 
Thomas J. Booth, was a nephew of Thomas 
NuTTALL. He shared his uncle's enthusiasm for 
travel and made botanical collections for the latter 
in N. Assam,' having sailed from Liverpool for 
Calcutta in June 1849. In 1853 Nuttall (and 
Booth) described 22 new Rhododendron species 
from there. 

From a letter to the Gardeners Chronicle,* 
written several years later, it is apparent from the 
signature, that Booth at that time was living at 
Rainhill Nursery, near Prescot. 

It might be possible that the unknown Booth 
who collected in Java in 1862 (see Cycl. Fl. Mai. I, 
1, 1950, 72a), was identical or a relation. 

Literature. (1) Cf. Trans. Bot. Soc. Edinb. 
41, 1972. p. 351-363. 

(2) Cf Gard. Chron. 1862, p. 406. 

•Borgman S. V. D., Father Ernst 

Alcxishafen; a student of the Max Planck Inst, 
fur Zuchtungsforschung at Cologne, spent several 
months in the Eastern and Western Highlands, 
Territory of New Guinea, collecting herbarium 
specimens and the corresponding cytological 

Collecting localities. 1960. Territory of 
New Guinea: 4 weeks on the E. slope of Mt Wilhelm 
in the alpine zone, and again in Sept. and Nov., 


Borssum Waalkes 

Flora Malesiana 

[ser. I, vol. 8^ 

climbing the summit (4700 m) three times; other 
collections in the Upper Chimbu Valley between 
Mogl (near Kundiawa) and Denglagu, above 
Keglsugl airfield up to Komanemambuno (the 
Pengagl River camp of 1959 Archbold Exp.) and 
in the Middle and Upper Wahgi River valley (Mt 
Hagen, Fatima, Banz, Nondugl, Minj, Mingende, 
Kondiu, Nera Gaima). 

Collections. 454 nos, in Herb. Lae, Herb. 
Leyden, and ? Edinb. (as E. Bergman). 

Literature. (1) Author of: 'Anteil der 
Polyploiden in der Flora des Bismarckgebirges 
von Ostneuguinea' (Zeit. Bot. 52, 1964, p. 118-172, 
1 map, 21 fig.). 

Borssum Waalkes, Jan van 

took his Ph.D. at Leyden University in 1966. 
Horticultural Taxonomist, Botanic Gardens, 
Groningen University. 

Collections. Correction: Only Fungi de- 


Possibly a contemporary medical colleague of 

Collections. A specimen 'O. Indien Borst' 
is preserved in Herb. Leyden. The label (of Lin- 
dernia anagallis (Burm. /.) Pennell) was written 
by C. L. Blume. 

Boswezen Nieuw Guinea 

To the collectors in the BW series add: Ir P. J. 
Faber (Kebar), Ir M. Flach (Lake Jamur; Fak- 
Fak), R. F. Hulskamp (Fak-Fak), H. Lorenzo, 
Ir J. LuiTJES (Manokwari; Arfak), B. Mahuze 
(Arguni Bay), Momot (Manokwari), C. M 
MoRTiER (Manokwari, Biak), A. Nusi (Manokwari ; 
Kebar), H. J. van Petersen (Japen; Manokwari), 
Ch. Riekerk (Kebar; Biak), H. C. J. Roeby 
(Kebar; Manokwari), F. Saurwalt (Wildeman 
River), W. E. F. Siebenhaar (Fak-Fak; Arguni 
Bay), A. W. Telussa (Arguni Bay), R. Versteegh 
(Kebar; Manokwari; Ransiki; Sarmi), H. Warent 
(Kokonao), L. Worobai (Manokwari). 

The collector C. Lugtmeyer (Suppl. Cycl. Fl. 
Mai. I, 5, 1958, p. cclxii) rightly has the initials 
H. W. 

Bouillenne, Raymond 
(d. 1971 or early 1972). 

*Bowers, Miss Nancy 

American anthropologist, educated at Columbia 
University (B.S., Ph.D.), Assistant Professor of 
Anthropology, Duke University, Durham, North 
Carolina, U.S.A. She made expeditions to Aus- 
tralian New Guinea to investigate traditional and 
changing agricultural patterns; subsequently (1969) 
Visiting Research Fellow, Austr. National Univer- 

Collecting localities. 1963- . She visited 
Western and Southern Highlands of Australian 
New Guinea, collecting almost all plants in Upper 
Kaugel Valley (W. Highlands), but some items 
from Upper Mendi (S. Highlands Distr.), Porgera 
(W. Highlands Distr.), and Kandep (W. Highlands 

Collections. Herb. Lae: nos 1-807; dupl. 
eventually in Canberra, Wash. U.S. Nat. Mus., 


Kew, and Brisbane. Dupl. of Pandanaceae to the 
B. P. Bish. Mus. Hawaii (Dr St. John), and to Dr 
B. C. Stone, Herb. Univ. Malaya Kuala Lumpur. 

Boxall, Richard 

Collecting localities. Add: In 1891 
Philippines: in Samar and Mindanao. 

Braat, Christoph George Sigismund 
(d. 1891, Velp, Gld., Netherlands). 

*Brand, D., see Forest Dept North Borneo. 

*Brandes, Elmer Walker 

(1891, Washington, D.C., U.S.A.; x), plant 
pathologist, educated at Michigan State College 
(B.S. 1913, M.S. 1915); student at Cornell Uni- 
versity (1916-17), and Ph.D. University of Michi- 
gan (1919). Up to 1959 he made an impressive 
career in sugar-plant investigations, mainly of the 
Dept of Agriculture in Washington. Many were 
the honours bestowed on him. 

He was sponsored by the said Agricultural 
Department to collect sugar-cane varieties in 
Australian New Guinea, together with Jeswiet 
(see Cycl. Fl. Mai. I, 1, 1950) and Pemberton (see 
this Suppl.). In total 292 varieties were brought 

Collecting localities. 1928. E. New 
Guinea: Fly River, Palmer R., Ok-Tedi R., 
Strickland R., Lake Murray, Port Moresby, Rigo, 
Sepik R., Lae. 

Collections. Jeswiet brought back many 
herbarium specimens, but whether the other 
members did, is not known to me. 

Literature. (1) Cf. Hawaiian Plant. Rec. 55, 
1958, p. 213; E. W. Brandes: 'Into primeval 
Papua by seaplane' (Nat. Geogr. Mag. 56, 1929, 
p. 253-332, ill.); 'Cane Growers' Quart. Bull. 
Brisbane 15, 1951, p. 41^2. 

Biographical data. Who was who in 
America 1961-68, vol. 4; Amer. Men of Science, 
Phys. & Biol. Vol., ed. 10. 

Brass, Leonard John 

(d. 1971, Cairns, Australia), was awarded a 
honorary Dr's degree from Florida University in 

In May 1966 he retired from the Archbold 
Expeditions and Museums and settled in Australia, 
where he was offered a honorary curatorship of 
the Herbarium of the North Queensland Natur- 
alist's Club at Cairns. 

Itinerary. Add: 5th Archbold Expedition, 
1956.^" — 6th Archbold Expedition, 1959." From 
Lae to summit Mt Kaindi (Mar. 25-May 28); to 
Mt Wilhelm (15.000 ft) (June 6-Aug. 1); Mt Otto 
from Kotuni on the S. slopes, Mt Michael, Purosa 
in the Okapa region; Kratke Mts with ascent to Mt 
Elandora (8500 ft) (Aug. 2-Nov. 9); Upper Mark- 
ham Valley (Nov. 9-29). 

Collections." Exp. 1959 in U.S. Nat. Herb.: 
3622 nos incl. 342 cellular crypt.; dupl. will be 
distributed in the future. Moss dupl. in Herb. 
Bartram, Farlow. 

Literature. (9) Add: Mosses by E. B. 
Bartram in Brittonia 9, 1957, p. 32-56. 

(10) L. J. Brass: 'Results of the Archbold 
Expeditions no. 79'. 'Summary of the fifth Arch- 
bold Expedition to New Guinea, 1956-1957' (Bull. 



Cyclopaedia of collectors — Supplement II 


Am. Mus. Nat. Hist. 118, 1959, p. 1-70, 1 map, 8 

Mosses by E. B. Bartram in Blumea 10, 1960, 
p. 142-150. 

(11) Cf. extensive data in Fl. Mai. Bull, no 15, 
1960, p. 717-718. 

L. J. Brass: 'Results of the Archbold Ex- 
peditions no. 86'. 'Summary of the sixth Archbold 
Expedition to New Guinea, 1959' (Bull. Am. Mus. 
Nat. Hist. 127, 1964, p. 145-216, map, photogr.). 

(12) For a complete comprehensive Hst, see Fl. 
Mai. Bull, no 21, 1966, p. 1422-1425. 

Biographical data. J. Am. Arb. 52, 1971, 
p. 695-698, portr. 

♦Bray, P. S. 

a British graduate volunteer under the Voluntary 
Service Overseas Programme; he graduated in 1968 
>Aith an external London degree in botany from 
Portsmouth Technical College. At present (1969) 
selling computers. Attached to the Forest Depart- 
ment Kepong from Sept. 7P65-Mar. 7969. He 
occasionally collected in the series of the Forest 
Research Institute with P. F. Cockburn and 
maybe others in Malaya. 

Breda de Haan, J. van 

(d. 1917). 

Biographical data. Teysmannia 28, 1917, 
1 p. in suppl. obit, announcement. 

*Bridg(e)]and, Leon 

an Agricultural Consultant Officer at Rabaul, 
Bismarck Islands, who made a small collection of 
plants in association with a Nutrition Survey 

Collections. At least in 79-^7 in the Territory 
of New Guinea. Numbered in the NGF series; 
Herb. Lae. 

Brockx, W. Ch. L. 

(d. c. 1973, Amhem, Netherlands). 

•Broekhuijsen, Dr H. O. 

^ probably Gerrit Jeronoimo Broekhuysen 
(1908, Java; x), zoologist who studied biology at 
Leyden University (1929-33); Ph.D. 1936. From 
1937-39 at the University Capetown; entomologist 
at Pretoria, 1939-41; freshwater biologist Java, 
1941-46; at present in the Dept of Zoology 
University Capetown, Randebosch, S. Africa. 

Numerous zoological, especially ornithological 

Collections. He evidently collected some 
grasses for Dr H. C. D. de Wit during the 
Japanese occupation of Java (at least on 28-10- 
2603, Jap. chronology); in Herb. Bog. 

Brongersma, Leo Daniel 

Suppl. Cycl. Fl. Mai. I. 5. 1958, p. cclxiv add: 
The Star Mountains Expedition took place in 1959 
(sec under C. Kalkman, B. O. Van Zanten, and 
J. J. F. E. DE Wilde). 

Brooke, Winifred Mary Adelaide 
Collections. Also dupl. in Herb. Edinb. 

Brouwer, H. A. 

(sec Cycl, II. Mai. I, I, 1950) is Hendrik 


Aldert Brouwer (1886, Medemblik, Nether- 
lands; 1973, Amsterdam) who worked with the 
Dept of Mining in the D.E.I, from 1910-17; 
Professor of Geology at Delft 1918-29, and at 
Amsterdam University 1929-57. After his retire- 
ment he still lived in Amsterdam. 

Author of numerous books and papers, mainly 
of the geology of the former Dutch East Indies. 

biographical data. World Who's Who in 
Science 1968. 

Brown, Robert 

Collections. Add: Extensive data on col- 
lecting localities in Australia, key to the letters 
and numbers used in the 'Prodromus', etc., are to 
be found in Stearn's Introduction.^ 

Literature. (1) Add: 'The Prodromus' 
(1810) and 'Supplementum primum' (1830) were 
reprinted (facsimile 1960) with an Introduction by 
W. T. Stearn. 

(2) Add: For particulars on the drawings see 
Cl. Nissen, 'Die botanische Buchillustration' 
1951, vol. 2, p. 61 sub Flinders. 

Biographical data. Austr. J. Sc. 21, 1958, 
p. 127-130; in W. T. Stearn, 'Introduction to R. 
Brown Prodromus etc' (facs. 1960) p. v-lii, portr. 

Browne, Francis George 

left Borneo about 1959 and accepted a post in 
charge of the West African Timber Borer Research 
Unit. He later repatriated. 

*BRUN nos, see Forest Department Brunei. 

Brunig, Eberhard Friedrich (Wilhelm Otto) 

Add: In 1964 he lived in Hamburg, Germany, 
working on a thesis on Kerangas vegetation. 

Author of several papers on the vegetation of 

Collecting localities. Add: Merurong 

Literature. (1) A.o. 'The Merurong Plateau' 
(Borneo Terr. For. Bull. 1, 1959, p. 1-9); 'A 
Guide and Introduction to the Vegetation of the 
Kerangas Forests and the Padangs of the Bako 
National Park, Sarawak' (Symp. Ecol. Res. Humid 
Trop. Veget. Kuching 1963, 1965, p. 289-318, 1 
fig., 1 map, 2 tab.); 'Some observations on the 
status of Heath Forests in Sarawak and Brunei' 
(Proc. Symp. Recent Adv. Trop. Ecol. Varanasi 2, 
1968, p. 451-457, 4 fig., 3 tab.); 'Der Heidewald 
von Sarawak und Brunei' (Mitt. Bundcsforsch. f. 
Forst- u. Holzwirtsch. Reinbek bei Hamburg no 
68, Sept. 1968, 431 pp., roneo, with fig., maps, 
etc.); 'The Classification of Forest Types in 
Sarawak' (Mai. For. 32, 1969, p. 143-179, 7 fig., 
2 tab.); etc. 

♦Brunson, L. W. 

collected some plants together with W. H, 
Lambert (see Cycl. Fl. Mai. I. 1, 1950) in 1945; 
probably during military expeditions. 

Buchanan, Francis, cf. Hamilton. F. Buchanan 
in Cycl. Fl. Mai. I, 1, 1950 and Suppl. Cycl. Fl. 
Mai. 1, 5, 1958. 

*Budcrus, J. 

collected together with D. G. Frodin (see there). 



Flora Malesiana 

[ser. T, vol. 8^ 

♦Biirger, Heinrich 

(1804 or 1806, Hameln on the Weser, Hanover, 
Germany; 1858, Indramaju, Java). Of his edu- 
cation not much is icnown with certainty, but it 
seems that he was, from 1821, a student in mathe- 
matics at Gottingen University. He used the 
doctor's title. In 1823 he went to Holland and 
sailed for Java to be appointed apprentice pharma- 
cist in the hospital at Weltevreden. In 1825 he was, 
at his own request, put at the disposal of the 
surgeon-major Ph. F. von Siebold in Deshima, 
the Dutch settlement in Japan. In 1828 he took 
over part of Von Siebold's function and up to 
1835 he sent large zoological and botanical 
collections from Japan to Leyden. He was com- 
missioned to Java in 1832, and in 1833 joined an 
expedition of the 'Natuurkundige Commissie' to 
the Padang Uplands (see P. W. Korthals) as a 
mineralogist.* He was pensioned in 1843, and 
returned to Java as a private citizen; there, his 
business interests lay in many fields. In 1854 J. 
Motley (see there) was his guest in Batavia. 

Especially Miquel has named many Japanese 
plants in his honour. 

Collections. Japanese plants in Herb. 
Leyden and in several other herbaria. It seems 
probable that he is the 'Burger' (see Cycl. Fl. Mai. 
I, 1, 1950, p. 91) who sent botanical material 
(presumably from Java) to Blume, as his name 
was often written in that way. Dr Leenhouts of the 
Rijksherbarium remembers to have seen Sumatra 
specimens of the Blume herbarium with a 'B' at 
the left upper corner of the label, which by 
exclusion he inferred to be Burger specimens. 

Literature. (1) Cf. L. B. Holthuis in 'Early 
European Investigations on Japanese Carcinology 
(1640-1854)' (Study of Mollusca no 3, 1967, p. 15- 
20) and L. B. Holthuis & T. Sakai: 'Ph.F. von 
Siebold and Fauna Japonica. A history of early 
Japanese Zoology' 1970, p. 37-42. 

(2) H. Burger, Phil. Dr.: 'Aanteekeningen op 
eene reize door een gedeelte der Noord-Westkust 
van Sumatra, ... in het jaar 1833. Etc' (Verh. Bat. 
Gen. K. & W. 16, 1836, (159)-(226)). 

Biographical data. Blumea 11, 1962, p. 

BUsgen, Moritz 

Collecting localities. Add: In Central 
Java also at: Tudir, Wonosemi, Ngliron, Gundih. 

*Bujang, Hj., see Forest Dept Sarawak. 

*Buliner, Ralph Neville Hermon 

(1928, Hereford, England; x), anthropologist, 
B.A. at Cambridge (1952), Ph.D. of the Australian 
National University, Canberra (1961); Research 
Scholar, Austr. Nat. Univ. (1954-57); Lecturer/ 
Senior Lecturer in Social Anthropology University 
of Auckland, New Zealand (1958-67); Professor of 
Social Anthropology, University of Papua and 
New Guinea, Port Moresby (1968- ). Author 
of anthropological papers^. 

Collecting localities. Eastern half of 
New Guinea in Western Highlands District: Baiyer 
Valley (Aug. 1959-60); Kaironk Valley, Schrader 
Range (intermittently from Aug. 1963 to 1968 at 


Collections. Herb. Lae: approx. 350 nos; 
two dupl. of most numbers in Auckl. War Mem. 
Mus., New Zealand. 

Literature. (1) R. N. H. Bulmer: 'Edible 
Seeds and Prehistoric Stone Mortars in the 
Highlands of East New Guinea' (Man 64, 1964, 
Art. 183, p. 147-150); 'Why is the Cassowary not a 
Bird? Zoological Taxonomy among the Karam of 
New Guinea Highlands' (Man n.s. 2, 1967, p. 5-25; 
includes general account of field work in the 
Kaironk Valley region, Schrader Range). 

Biographical data. In New Zealand Who's 
Who, and Who's Who in Australia. 

*Buntar, A., see Forest Dept North Borneo. 

Burbidge, Frederick William 

Collections. Dupl. from A^. Borneo and the 
Philippines in Herb. Edinb. also. 

Burcham, Levi Turner 

Add: Biographical data. Portr. in Hat- 
usiMA, Flora Ryukyus 1971, p. 75. 

Burchard, E. W. 

? identical with W. F. G. Burchard (see there). 
Collections. Dupl. (Rosenstock exsicc.) 
from Sumatra (1907) in Herb. Edinb. 

Burger, see H. Burger. 

*Burgess, Peter Frederick 

(1925, Hertford, England; x), B.Sc. For. 
University College North Wales, and Dept For. 
of St. Johns College Oxford; Malayan Forest 
Service 1946-1956; Deputy Conservator, N. 
Borneo and Sabah 1956-1965; Colombo Plan 
Silvicultural Ecologist, Forest Research Institute 
Kepong, Malaysia, 1967- 

Author of papers in the Malayan Forester, the 
Malayan Nature Journal, and of a book 'The 
Timbers of Sabah'. 

Collecting localities. With P. F. Cock- 
burn (see there) on G. Tapis, Pahang (Sept. 3-14, 
1968).^ Main collecting areas: East Coast of Sabah, 
Ranau (Sabah); Malay Peninsula: G. Inas, G. 
Tapis, G. Stong, G. Pulai and Bt Jelakoi (Johore), 
Lepar and Gua Peningat (limestone hill) in Pahang. 

Collections. Herb. Kepong and Sandakan. 
Dupl. at Kew and Leyden. Collections in West 
Malaysia mostly for ecological purposes. 

Literature. (1) P. F. Burgess: 'An ecological 
study of the hill forests of the Malay Peninsula' 
(Mai. For. 31, 1968, p. 314-325). 

Burkill, Humphrey Morrison 

retired mid-1969; he has now joined the Kew 
staff under the Bentham-Moxon Trust to revise 
Dalziel's 'Useful Plants of West Africa'. 

Collecting localities. Add: 1957. P. 
Satuma, P. Senang, etc. (Jan. 30-Feb. 3, Aug. 2-6); 
Malacca (Apr. 19-20), Eraser's Hill (Apr. 30- 
May 4); Thailand, Kanburi Prov. (Nov. 25-27).— 

1958. P. Satuma, P. Senang, etc. (Feb. 15-19).— 

1959. Johore (June 21-29, July 13-24); Eraser's 
Hill, Pahang (Aug. 25-29) with M. Shah and M. 
NuR BIN Juma(a)t. — 1962. Malaya, Singapore, 
BXidJava, mainly collecting Algae. — 1966. Sarawak: 
Mersing, Tk. Sari (Jan. 1-3); Malaya: P. Pemang- 
gil, Johore (July 4-10). 



Cyclopaedia of collectors — Supplement II 


Certainly at many other places during his stay 
in Singapore. 

Collections.^ Add: Dupl. in Herb. Edinb. 
(acq. 1960). 

Literature. (1) J. H. Kern: 'Juncaceae, a 
new family record for Malaya' (Gard. Bull. Sing. 
17, 1958, p. 91-92). 

Edinb. 26, 1965, p. 325-329); 'Cosmianthemum: 
a Bornean genus of Acanthaceae' {I.e. p. 365-382). 
B. L. Burtt: 'Sarawak Plants'. I {ibid. 29, 1969, 
p. 141-155. f. 1-6, pi. 5-6); 'Studies in the Ges- 
neriaceae of the Old World XXXIII. Some species 
of Cyrtandra, chiefly Bornean' {ibid. 30, 1970, p. 
23^2, 1 fig.). 

Burklll, Isaac Henry 

(d. 1965, Leatherhead, Surrey, England). 

Biographical data. Add: Gard. Bull. Sing. 
17, 1960, p. 341-349, photogr., partial bibliogr.; 
ibid. p. 350-356; Nature 206, 1965, p. 871; Gard. 
Bull. Sing. 22, t967, p. 67-105, incl. bibliogr. and 
list of taxa (proposed, and named after him); Ass. 
Trop. Biol. Bull, no 7, 1966, p. 52. 

Burret, Carl Ewald Max 

(d. 1964, Berlin, Germany). 

Biographical data. Principes 2, 1958, p. 
87-91; Willdenowia 2, 1959, p. 140-142, portr.; 
ibid. 4, 1965, p. 23-31, 1 Abb., bibliogr., portr. 

•Burtt, Brian Laurence 

(1913, Claygate, Surrey, England; x), botanist, 
B.Sc. London (1936); on the staff of the Royal 
Botanic Gardens, Kew (1930-51, interrupted by 
war service 1940-46); at present Principal Scientific 
Officer Royal Botanic Garden, Edinburgh. Monog- 
rapher of Gesneriaceae.^ He visited the Malesian 
region (see below), Pakistan (1958), South Africa 
(1965, 1966), and Malawi (1967). 

Itinerary. 1962," with P. J. B. Woods (see 
there). Malay Peninsula: Kuala Lumpur (Apr. 14), 
Ipoh (19), Perlis (23), Langkawi Js. (Apr. 28-May 
3); Sarawak: Kuching area (May 15), Niah caves 
(June 3), Marudi (10), G. Mulu and Sg. Melinau 
(12), Lambir Hills (July 4), Kuching area (13), 
Kapit area (Wong Pelagus on Rejang and S. Bena: 
July 19), Kuching area and G. Gaharu (30), 
Lundu distr. G. Gading (Aug. 5); Poi Range, G. 
Berumput (Aug. 8-17).— 1967, with A. M. 
Martin. Sarawak (July-Oct.):^ Kuching area 
(July 22-26), Hose Mts (Aug. 2-27), Kuching area 
(Sept. 7-14); in the 5th Division from Long 
Semadoh to Bakelakan, G. Murud and return 
(Sept. 20-Oct. 18). 

Collections. In Herb. Edinb.: nos Burtt 
1603-1823 {Malay Renins. 1962), 1825-2901 
{Sarawak 1962),* 4700-5577 {Sarawak 1967): dupl. 
in Herb. Sarawak. Primary objectives of collecting 
were Gesneriaceae and Zingiberaceae, partly for 

Literature. (1) Author of 'Studies in the 
Gesneriaceae of the Old World' in Notes R. Bot. 
Gard. Edinb. 

(2) 'Report on B. L. Burtt Gesneriad Hunting 
Expedition' (Gloxinian 12, 1962, p. 77-79, 1 map) 
(n.v.); cf. also map in Notes R. Bot. Gard. Edinb. 
29, 1969, p. 142. 

(3) Cf. Mimcogr. internal report: 'Expedition 
to Sarawak, 1967" (li. L. Burit& A. M. Martin), 
and map in Notes R. Bot. Gard, Edinb. 29, 1969, 
p. 142. 

(4) B. L. BuRrr & R. M. Smith: 'A new species 
of Zingiberaceae from Sarawak' ( Irans. Bot. Soc. 
Edinb. 39, 1964, p. 502-511); 'A new genus of 
Acanthaceae from Sarawak* (Notes R. Bot. Gard. 


Butterworth, W. J. 

Biographical data. 
Annual for 1971, p. 60. 

Portr. in Str. Times 

*Buwaya, see sub H. C. Conklin. 

Buysman, Maarten, was erroneously written 
Buisman in Suppl. Cycl. Fl. Mai. I, 5, 1958. 

*Buzacott, James Hardie 

(1902, Brisbane, Queensland, Australia; x) was 
educated in the University of Queensland (B.S. 
1933, M.S. 1938). From 1925^7 Entomologist, 
Queensland Bureau of Sugar Experiment Stations; 
subsequently Senior Plant Breeder with the same 

He published extensively on cane breeding and 

In 1951 he and C. G. Hughis (see there) were 
sponsored by the Queensland Bureau of Sugar 
Experiment Stations to collect cane varieties in 
Australian New Guinea.^ They were accompanied 
by J. N. Warnir as a representative of the 
Hawaiian Sugar Planters' Association. 

Collectincj locai rrihs.* 1951. E. New 
Guinea: Port Moresby (Apr. 17 21). Lae. Mark- 
ham K.. Kain;intutu, Aiyura. Cioroka, Chimbu, 
Mt Hagen, Mendi, Laloki. Sogcri. Leaving New 
Guinea by plane from Port Moresby on June 3. 



Flora Malesiana 

[ser. I, vol. 8^ 

Collections. 1 65 varieties of cane (SaccAfln/m 
and Erianthus mainly) were collected and herbar- 
ium specimens might be in existence. 

Literature. (1) J. H. Buzacott & C. G. 
Hughes: 'The 1951 Cane Collecting Expedition to 
New Guinea' (Queensl. Bur. Sugar Exp. Stat., 
Cane Growers' Quart. Bull. 15, 1951, p. 35-72). 

(2) Cf. Hawaiian Plant. Rec. 55, 1958, p. 213; 
Cane Growers' Quart. Bull. 15, 1951, p. 43-54. 

Biographical data. In Bibl. Austr. Entom. 
1775-1930 (R. Zool. Soc. N.S.W. 1932, p. 32). 

BW nos, see Boswezen Nieuw Guinea. 

Campbell, Charles 

Add: His botanical MSS possibly came into the 
hands of J. Arnold (see J. Bastin in J. Soc. 
Bibliogr. Nat. Hist. 6, 1973, p. 365 (note 245), but 
probably were lost with the fire on the 'Fame' (see 

*Caiiright, James Edward 

(1920, Delaware, Ohio, U.S.A.; x), botanist 
educated at Miami University (Oxford, Ohio), and 
Harvard University (Ph.D. 1949); Teaching Fellow 
in Botany, Harvard Univ., 1946-49; Associate 
Professor, Department of Botany, Indiana Uni- 
versity, Bloomington, U.S.A., 1949- . With 
the assistance of a Guggenheim Fellowship he 
spent five months in a guest house in Kebun Raya, 
Bogor, Java; subsequently he worked for about the 
same time at the Herbarium of the Singapore 
Botanic Garden. 

Collecting localities. Sept. 79<50-Feb. 
1961 in W. Java: Gardens at Bogor and Tjibodas, 
and a few plants from Udjong Kulon Nature 
Reserve; throughout Malaya, in Johore: G. Pulai 
(Mar. 2, 8, 24); G. Panti (Mar. 4, 12); Sg Dohal 
and Sg. Kayu swamp forests (Apr. 21); Mersing to 
Endau (Apr. 24); Kukup region, Pontian Distr. 
(May 5); Kota Tinggi to Kg. Sedili Besar (May 19, 
June 23); during trip in Malacca, Negri Sembilan, 
Pahang (Eraser's Hill), Perak (chiefly on Maxwell 
Hill), Penang, and Selangor (June 9-17); numerous 
Singapore collections from Botanic Gardens, Bukit 
Timah, Labrador Reserve, MacRitchie Reservoir 
Reserve, Mandai Rd Distr., and Seletar Reservoir 

Collections. About 350 nos from above- 
mentioned localities, including much preserved 
material. Dupl. of most ranalean specimens from 
Kebun Raya Bogor in Herb. Bog. ; Myristicaceae 
deposited in Herb. Un. Cincinnati; other Malaysian 
collections not yet distributed. His collections will 
eventually be deposited in the Indiana Un. Herb. 

Biographical data. Amer. Men of Sc. ed. 
9 and 10; Who's who in America 1961. 

Cantor, Theodore Edward 

See sub White (Cycl. Fl. Mai. I, 1, 1950) (1809, 
Copenhagen, Denmark; d. 1854), M.D. Halle 
1833, entered the uncovenanted Bengal Medical 
Service in 1835, and from 1837-38 worked as a 
surgeon to the Bengal Marine Survey. He volun- 
teered as Assistant Surgeon to one of the regiments 
despatched to China (Opium Wars!) by the E.I.C. 
and stayed in Chusan from July 1840-Mar. 1841, 
making zoological^ and botanical collections 
(partly a failure because of ineffective preservatives) 


and numerous sketches of plants etc. which are 
still in the India Office Library.* 

Later he served in Malaya, and was in Penang 
in 1844. 

Bambitsa cantori Munro was named in his 

Collections. Doubtful whether any Malayan 
plant collections exist. 

Literature. (1) Author of 'The Zoology of 
Chusan' (Calcutta 1842) and several zoological 

(2) Cf. M. Archer in 'Natural History Drawings 
in the India Office Library' (London 1962) p. 51- 
53, 76. 

Biographical data. Biogr. Index Britten 
& Boulger 2nd ed. by Rendle, 1931. 

*Carlquist, Dr Sherwin 

Associate Professor of Botany at Claremont 
University, and anatomist-palynologist-phylogenist 
of the Rancho Santa Ana Botanic Gardens, 

In 1962 he travelled 14 months in the Pacific 
(incl. Society Is. and New Guinea) and had plans 
to do further collecting in tropical Queensland, 
and Indonesia, financed by a National Science 
Foundation Grant. 

He wrote a book on natural history, based upon 
his trip and research.^ 

Itinerary. Terr, of New Guinea. In 1963 he 
accompanied P. Van Royen (see there) on a trip 
from Lae to Rawlinson Range and Saruwaket 
(Feb. 20-Mar. 3); Mt Piora, 3719 m, W of Lae 
with E. Henty etc. (Mar. 9-13). 

Collections. 850 nos of dried and pickled 
material, particularly of Scaevola, Stylidiaceae, etc. 

Literature. (1) S. Carlquist: 'Island Life' 
(New York 1965). 

*Carne, M. A. & C. Baker 

Itinerary.^ 2nd Colonial Sugar Refining Co. 
(of Australia) Expedition in SE. New Guinea. 1914. 
Setting out from Buna (May 31) and collecting in 
the country lying west of Buna down to the northern 
slopes of the Hydrographers Range and as far 
north as the Kumusi R.; after 6 weeks return to 
Buna, setting out again on July 22, and during the 
2nd trip collecting behind the Hydrographers 
Range on the Bariji R., behind Oro Bay and NW 
in the Koena and Mambare R. district; leaving 
Buna (Aug. 19), proceeding along the Kokoda 
Trail to Port Moresby. 

Collections. 106 varieties of sugar-cane. 
Some varieties may be represented in herbaria. 

Literature. (1) Cf. Hawaiian Plant. Rec. 55, 
1958, p. 213; Cane Growers' Quart. Bull. 15, 
1951, p. 40. 

*Carocci-Buz!, V. P. 

collected in Hort. Bot. Bogor. in 1957. Material 
in Herb. Bog.; dupl. in other herbaria. 

*Carolyn, see Philippine National Herbarium. 

Carr, Cedric Er(r)oll 

Collecting localities. Details mentioned 
under 1935-36. SE. New Guinea (Cycl. Fl. Mai. I, 



Cyclopaedia of collectors — Supplement II 


1, 1950) substitute by;^ Centr. Division: from 
Jan.-Aug. 1935 in the lowland country around 
and to the NW. of Port Moresby, mainly at 
Kanosia (Jan., Feb., Apr.), Veiya (Mar.), Rouna 
(1300 ft, Apr.-July), and Koitaki (1500 ft, Apr.- 
July); towards the Owen Stanley Range (Aug. 16) 
via Hailogo (3000 ft, Aug. 31-Sept. 4), the S. 
slopes of the Range, camping at Boridi (4700 ft) 
(Sept.-Dec); Northern Division: leaving Boridi 
(Dec. 3), camp at 6000 ft near Alola on the N. side 
of the Range, Lala River (5500 ft, up to early Jan. 
1936); subsidiary camp nearer the Gap (8000 ft), 
collecting up to 10,000 ft (Jan. 12-30); down to 
Isuarava (3500-4500 ft), Lala River and Yodda 
River below Isuarava at 3500 ft (Jan. 31-Mar. 15), 
at Kokoda (1200 ft. Mar. 17-May 23), Fara River 
(May 24). Besides there are a few numbers from 
Saputa (200 ft), Inapa (500 ft), and Buna (sea-level) 
(Apr. 5-8, 1936), either collected by Carr himself 
en route or by native collectors. 

He never reached Mt Victoria. 

Collections. Add: The remainder of his 
1935-36 New Guinea collection was shipped from 
Singapore to the British Museum in 1954-55, and 
has since been distributed to the recipients of 
Carr's sets. With it were 6(X) gatherings of orchids 
with minute field tickets, but unnumbered, 
evidently some of the last plants gathered before 
his death, or cultivated at camps. The latter were 
numbered (between 16,600-17,232) in theB.M. and 
have been distributed. Besides in the already 
mentioned herbaria, a few numbers are at Lae, 
Tucuman, Geneva, and Am. Arbor* 

Dupl. from Borneo (1933) and New Guinea 
(1935-36) in Herb. Edinb. 

Literature. Add: (5) Cf. E. W. Groves in 
Fi. Mai. Bull, no 16, 1961, p. 826-827. 

(6) Cf. E. W. Groves in Fl. Mai. Bull, no 15, 
1960, p. 719; ibid, no 16, 1961, p. 826-827. 

•Carrick, John 

(1914, Glasgow, Scotland; x) was educated in 
the University of Glasgow (B.Sc. 1938); 1940-46 
in Military Service, from 1943 onwards in India; 
1946-51 Lecturer in Mycology and Plant Pathology 
at the University of Manitoba, Winnipeg, Canada; 
from 1952 Lecturer in Plant Physiology at the 
University of Malaya, Singapore and Kuala 
Lumpur; Senior Lecturer since 1960. His appoint- 
ment as Field Pharmacognosist (Colombo Plan) 
terminated end 1967. In 1968 he moved to Adelaide, 
S. Australia. Liaison Officer at Kew in Apr. 1970. 

Plant collections have been made in connection 
with a physiological-ecological study of lowland 
kcrangas vegetation and soil. 

Collecting localities. 1959. NW. Borneo, 
Sarawak: Bako National Park (Apr. 18-May 11), 
with E. C. Enoch; Santubong (Apr.). — 1961. 
Malaya: Jambu Bongkok Forest Reserve, Treng- 
ganu (Mar.); Mcnchali Forest Reserve, Pahang 
(Mar.); Tanjong Hantu Forest Reserve, Perak 
(Apr.); Sarawak: Bako National Park (Apr.). — 
1967. Padang area of Jambu Bongkok (Feb. -Mar.). 

Collections. Main set in Herb. Univ. Mai. 
Kuala Lump.; Sarawak dupl. in Herb. Sing. (6(X) 
nor), and in Iferb. for. Dep. Sarawak. No dupli- 
cates were made of the F-orcst Reserve collections. 
His personal numbers as far as the survey goes end 
with JC 1670 (P.C.S.M. no 2600). 

Carrier, Charles Lionel 

retired from the Malayan Forest Service in Mar. 
1964; was still living in Malaya, at Ipoh, in 1970. 

Biographical data, Mai. For. Apr. 1964, 
p. 81, portr. 

♦Carroll, D. 

(1897, Malaya; 1955, ?) was attached to the 
Sarawak Forest Department from its start in 1921, 
as Forest Ranger, special grade, in charge of the 
administration of Kuching, Lundu and Sadong 
forest districts. By the time he had retired in 1954 
he had won a M.B.E. and been promoted to the 
senior staff.^ 

Collections. Several hundreds of numbers 
for the Forest Department, from 1922 onwards; 
Herb. Sarawak. 

Literature. (1) Cf. P. S. Ashton in Fl. Mai. 
Bull, no 21, 1966, p. 1432-1435. 

♦Carter, Dr Walter 

(1897, Leeds, England; x), naturalized American 
entomologist; B.S. Montana (1923), M.S. Min- 
nesota (1924), Ph.D. (1928), from 1925-30 em- 
ployed by the U.S. Dept of Agriculture; Head 
Entomological Dept Pineapple Research Institute 
Hawaii, 1930-62; in 1962 emeritus professor; in 
Jamaica 1962-65. He made numerous travels, and 
visited Africa, Malaya, Java, the Philippine Islands, 
Australia, and Fiji in 1937-38. From 1964-65 in 
the employ of the U.N. for Technical Assistance. 

Collections. Living orchids, also from Java, 
Celebes, and Fiji, turned over to Mr Nakamoto 
in Hawaii.^ 

Literature. (1) 'Carteria', The story of a 
collection (Bull. Pac. Orch. Soc. Hawaii 26, 1968, 
p. 3-8, 3 phot.). 

Biographical data. In Amer. Men of 
Science. Phys. and Biol. Sciences, 11th ed. 1965. 

Castro, Arturo P. 

(1914, Tondo, Manila, P.L; x), B.S.E. graduate. 
Southeastern College, Pasay City. Fiom 1941-49 
in the employ of the National Museum Manila as a 
Helper; from Apr. 1949 Laboratory Technician, 
Ateneo de Manila. He accompanied a scientific 
expedition to Bataan with Dr Gilliard (see there). 

Collecting localities. Add: A'. Luzon: 
Abra and Mountain Prov.; Mindanao; Palawan. 

Collections. Besides plants he collected 
animals and ectoparasites of birds etc. 

C.B.D., see Dalhousie, Countess of (Cycl. Fl. 
Mai. I, 1, 1950). 

Cer6, Jean-NlcoIas (de) 

Director of the Royal Garden at Montplaisir; 
rightly: Monplaisir. 

Biographical data. Add: Madel. Ly- 
Tio-fane (cd.): 'Mauritius and the Spice Trade. 
The Odyssey of Pierre Poivre' (Publ. no 4 of 
Mauritius Arch. Publ. Fund 1958, w. portr. facing 
p. 100). 

CF nos, sec Conservator of Forests series. 

♦Chai, Paul Piang Kong 

(1941, Kuching, Sarawak; x), graduated at the 
University of Malaya (lions Bot.) in 1970. From 




Flora Malesiana 

[ser. I, vol. 81 

Jan. 1970 appointed Forest Botanist, Forest 
Department, Sarawak. He will be engaged on an 
ecological study of mangrove forest in Sarawak 
and will continue the preparation of a manual on 
the non-Dipterocarp trees of Sarawak. 

He collects in the series of the Sarawak Forest 
Department, and accompanied H. P. Nooteboom 
on his Sarawak exploration in 1970, continuing 
another week's collecting after N.'s departure. 


Collecting localities. From 7970 on- 
wards throughout Sarawak. 

Collections. Herb. Sarawak and with 
Nooteboom's collection in Herb. Leyden. In the 
'S' series; dupl. in many herbaria. 

Chamisso, Adalbert von 

Literature. (1) Add: For itinerary in Alaska 
cf. Bot. Not. 1940, p. 298-299. 

Biographical data. Add: Occ. Pap. Calif. 
Ac. Sc. no 20, 1943, p. 21-27. 

*Chan Yee-Chong 

Assistant of the Forest Research Institute 
Kepong, Malaya. 

Collecting localities. 1969. Malaya: 
Kedah, Ulu Muda (Jan. 15-24) with P. S. Bray; 
Langkawi and. Pedis with Chelliah (Mar. 10-19). 

Collections. In Herb. F.R.I. Kepong. 


collected with G. H. S. Wood (see there) in A^. 


Borneo in the SAN series. See Forest Dept North 

Cheang Koh Choy 

collected in Malaya in 1963; Superintendent of 
the Penang Gardens. 

Cheesman, Lucy Evelyn 

(d. 1969, Highwood, near Chelmsford, Essex, 

Literature. (2) Add: 'The Land of the Red 
Bird' (London 1938). 

Biographical data. New Scientist 6, 1959, 
p. 80-81; Entom. Monthly Mag. 105, 1969, p. 

*Chelliah, S. 

Assistant of the Forest Research Institute at 
Kepong, Malaya, collected some hundreds of 
plants in Kedah, Penang, Johore, Selangor, Perak, 
Trengganu and Pahang in 1966-67; Bt. Bauk F. R., 
Trengganu (Nov. 12-14, 1968). 

*Chew Wee-Lek 

(1932, Singapore; x), botanist, educated at the 
University of Malaya, where he took his B.Sc. 
(Hons) in Botany under Prof. Holttum in 1956; 
in Oct. of that year he joined the staff of the 
Singapore BotanicGardens. He went to Cambridge, 
England, in Sept. 1957, and took his Ph.D. under 
Prof. Corner at Cambridge University in June 
1960. Since 1963 Keeper of the Singapore Her- 
barium and Botanist at the Botanic Gardens, 
Singapore, later Acting Director. He took part in 
two Royal Society expeditions to Mt Kinabalu 
under Prof. Corner. He joined the staff of the 
Royal Botanic Garden Sydney from Apr. 1970. 

His main interests are Urticaceae and Piperaceae 
of Malesia, and lianas of Malaya and Singapore. 

Litsea chewii Kosterm. was named after him. 

Collecting localities. 1957. Malaya: 
Trengganu (Mar. 27-31) and Kelantan (Apr. 1-6), 
Penang (7-8), Kedah (9-12), with Kiah bin Haji 
Salleh; Kedah, Perils, and Langkawi Is. (May 
10-29).— 1960. Java: Udjung Kulon and P. 
Peutjang (Oct. 26-Dec. 10, with Dr Kostermans, 
Mr KocHUMMEN, and Mr Gutierrez). — 1961. 
Sabah: Mt Kinabalu (May 29-Aug. 16, Royal 
Society Expedition, see under E. J. H. Corner). 
Malaya: Pahang at Tasek Bera (Oct. 24-31).— 
1962. In Pahang with M. D. Poore and F. Mbrton. 
Sarawak: Niah Caves (June 3-7), Baram R. (8-12), 
Mt Mulu (June 13-17), Mt Api and Benarat 
(20-29). — 1963. Sarawak: Seburan Limestones in 
Bau (July 12-15), Mt Gading (17-21), Sg. Priam 
(22), Mt Mentawa and Maja, near Kuching (25- 
28). Malaya: Cameron Highlands, collecting at 
Sg. Pauh Valley, Mt Berumbun, Bt Siku, Bt Ruil, 
Mentigi F. R., Sg. Boh, Sg. Bertam Valley, Mt 
Brinchang, Mt Jasar (Oct. 2-12). Sarawak: Bau 
and Tiang Bekap (Oct. 11-29).— 1964. Sabah: 
Mt Kinabalu at the Pinosuk Plateau between 8000 
and 5000 ft alt. (Jan. 10-May 20, 2nd Royal Society 
Expedition under Prof. Corner). — 1966. Sarawak: 
Bt Lambir (Feb. 3), Marudi F. R. (5-6); Baram 
Distr. : Kuala Melinau Paku (8-10); Sg. Tutoh, Sg. 
Melinau (11-12); G. Api and vicinity (13-17); Sg. 
Terikan in Limbang Distr. (18-19). Malaya: G. 
Bubu in Perak (Aug. 13-18), Cameron Highlands 



Cyclopaedia of collectors — Supplement II 


in Pahang (Oct. 1-4).— 1967 Sarawak: G. Api 
Benarat, Tiang Bekap limestones near Kuching, 
on Mt Maja and Mentawa (Mar. 10-14); Mt 
Santubong (15-23). 

Besides, numerous daily collecting forays in 
Singapore and nearby Johore. 

Collections. About 3000 nos of flowering 
plants of which those from 1957 mainly in the 
SFN series, those from Kinabalu {1961, 1964) in 
the RSNB series (Royal Society North Borneo 
numbers), and the rest in his own C.W.L. series 
(to be cited in his own name). The RSNB series 
distributed by Kew, the others by Herb. Sing. 
Dupl. in Leyden, Kew, Arn. Arbor., U.S. Nat. 
Mus., Copenh., Aarhiis, Berkeley, Saraw., Kepong, 
Edinb., and specialists. 

Chinnery, Ernest William Pearson 

Add: He was also the collector of 40 sugar-cane 
varieties for the Colonial Sugar Refining Co. of 
Australia in 1921 (cf. Hawaiian Plant. Rec. 55, 
1958, p. 213; Cane Growers' Quart. Bull. Brisbane 
15, 1951, p. 40). 

Collecting localities. Add: 7927. Pa/jwa; 
in the district of Korigo, Orama, and Durom on 
the side of Mt Obree in the Owen Stanley Range 
at c. 700-1000 m alt. 

*Chong, J., see Forest Dept North Borneo. 

♦Christiansen, Sofus Emil 

(1930, Salby near Haslev, Denmark; x) was 
educated at the University of Copenhagen; cand. 
mag. (geogr.) 1955; 1962 Assistant at the Geo- 
graphical Institute, University of Copenhagen. 

As a member of the Danish 'Noona Dan 
Expedition' (see there for Itinerary, Collections 
etc.) he collected some botanical specimens at 
Yalon (Mussau, Bismarck Arch.), and in Rennell 
(Solomon Is.) in 1962. In 1965 and 1966 he made a 
small collection in Rennell and Bellona (Solomon 

♦Cid, Miss F. 

Instructor of Botany in a private school in 
Pangasinan Province, Luzon. 

Cinatti, Ruy 

Collections. Add: In Herb. Leyden: 3S0 nos. 

♦Clark & Roberson 

collected a grass, numbered 1024, in moist rice 
paddies (Aug. 1945). Found by Veldkamp in 
MSS J ansen together with specimen of /'jt'«(/ora;j/i- 
15 sp. 

•Clarke, William Carey 

(1929. Marblehead, Mass., U.S.A.; x), took 
his Ph.D. in Geography at Berkeley (1968); 
presently Professor of Geography and Anthro- 
pology, California State College at Hayward, Cal., 
U.S.A.; 1968-69 appointed as Associate Professor 
of Geography, University of Hawaii, Honolulu. 
Member of exp)cdition to Bismarck Mts, Terr, of 
New Guinea, 1964-65, for study of cultural 
ecology of montane shifting cultivators (under the 
direction of A. P. Vavda of Columbia University).' 

Collecting Lf>CAi.rrii'.s. J 964 65. Terr, of 
New Guinea: Simbai Valley, Bismarck Mts. 


Collections. Nos 1-132 in Herb. Lae; 
certain specimens forwarded elsewhere, e.g. 
Pandanus to B. C. Stone, Kuala Lumpur; ferns to 
Holttum, Kew. 

Literature. (1) W. C. Clarke: 'From 
Extensive to Intensive Shifting Cultivation: A 
succession from New Guinea' (Ethnol. 5, 1966, 
p. 347-359); 'Soil Fertility and Cultivation 
Practices in New Guinea' (with J. M. Street) (J. 
Trop. Geogr. 24, 1967, p. 7-11). 

Clason, Else Willem 

(d. 1971, Groningen, Gr., Netherlands). 

Biographical data. Jaarboek, Versl. «& 
Meded. Kon. Ned. Bot. Ver. over 1971, 1972, p. 5. 

Clemens, Mary Strong 

(d. 1968, Chermside Garden Settlement, Bris- 
bane, Australia). 

Collecting localities. Add: Philippines: 
Mt Pulog (Mountain Prov., Luzon) with Ramos & 
Edano (Feb. 1925), where they detected the 
genera Pimpinella, Potentilla, and Koordersio- 
chloa, new to the Philippines. 

Collections. Also dupl. {Borneo; fairly com- 
plete set from New Guinea) in Herb. Edinb. 

Biographical data. F1. Mai. Bull, no 23, 
1969, p. 1669-1671. 

*Cockburn, Peter Francis 

(1946, Brighton, England; x), botanist, educated 
at University College, Oxford 1964-67. He arrived 
in Malaya in Aug. 1967 under Voluntary Overseas 
scheme, and worked at Kepong from Sept. 1967- 
Dec. 1968, with T. C. Whitmore. He made prolific 
collections in virgin territory along the east coast 
of Malaya and wrote up several families for the 
Tree Flora of Malaya. Mar. 1969 appointed 
Forest Botanist, Sabah, on contract. Co-author of 
the Tree Flora of Malaya. 

Collecting localities. Malaya. 1967. 
Ulu Kelantan, Sungei Lebir with T. C. Whitmore 
and B. C. Stone (Aug. 31-Sept. 24); Ulu Kelantan, 
Dabong, Relai F. R. (Oct. 15-Nov. 2).— 1968. G. 
Blumut, Johore (Feb. 2-8); Tg. Penawar, SE. 
Johore (19-24); Sungei Sedili Kechil (25-28); G. 
Panti, Johore (29-Mar. 6); Ulu Endau, Johore 
(Mar. 19-Apr. 1); Losong F. R. and Ulu S. Anak 
Andau (2-6); G. Lawit, Besut N. Trengganu (Apr. 
22-May 6); Ulu Sungei Trengan (May 27-June 
16), 6 collecting camps at various points on the 
river; G. Mandi Angin\ Trengganu, with T. C. 
Whitmore; G. Tapis, Pahang, with P. F. Burgess 
(Sept. 3-14); Bt. Chini, S. Pahang (Oct. 1-14), 
with P. S. Bray. A few collections made at Fraser's 
Hill, Tasck Bcrak (Pahang) and Ulu Kali on single 
day trips. — 1969. Sabah (N. Borneo). Kalabakan 
F. R. Tawau (May 5-12); Madai F. R. Mostyn 
(13-15); Lahad Datu Distr. (16-21); Sepilok F. R., 
Sandakan (June 5-7) 16 miles S of Sook on 
Pcnsiagan Road in the Witti Range (June 9-16); 
Loro F. R.. Kudat Distr. (17-20); Sepilok F. R. 
(Aug. 25-28); Patau Sclingan (Sept. 24-25); Pulau 
Timlnin Mala, Darvel liay (Oct. 5-9); Crocker 
Range F. R. (Oct. 12-20). 

Collections. 2110 collections in Malaya, 
numbering Kl-P 1 1595! -1 16000 and FRI 7000- 
8050. 8101 85(X). 10501-1 1 1 10; full set at Kepong, 
dupl. in Kuala Lump., Sing., and some to Arn. 



Flora Malesiana 

[ser. I, vol. 8^ 

Arbor., Sarawak and Sandakan. Sabah collections 
full set at Sandakan, dupl. to Kuala Lump., Sing., 
and partly to other herbaria. 

Most collections concentrated on tree species; 
also collecting Indotrislicha malayana in W. 



Literature. (1) P. F. Cockburn: 'Gunong 
Mandi Angin, Trengganu: a botanical visit' (Mai. 
Nat. J. 22, 1968-69, p. 164-170). 

(2) Cf. Blumea 18, 1970, p. 152-155). 

Collenette, Irish Sheila Darnton 

Add: She made a trip to Mt Kinabalu and 
Mesilau R. with H. Sleumer, H. P. Fuchs, and 
others in July 1963. 

Some plants were collected with Fuchs numbers 
(see there). 


♦Collins. John D. 

transport man and field assistant to the 6th 
Archbold Expedition 1959. Occasionally he 
collected plants with L. J. Brass or alone. 

Rhododendron collinsii Sleum. was named after 

Collecting localities. 1959. Terr, of 
New Guinea: Mt Otto (Aug. 12), Mt Michael (31). 
For the complete itinerary see L. J. Brass. 

Collections. See sub L. J. Brass. Plants 
collected by him evidently labelled L. J. Brass & 
J. D. Collins. 

*Coniber, Jim B. 

student gardener at Kew from Apr. 1951 until 
1955. He then went to Sapong Estate, near Tenom, 
Sabah (N. Borneo). 

Collections. ? All from the Tenom area, 
Sabah, rather few in number. According to Mr L. 
Forman (Kew) his 796-^ collections range from 
nos 100-162; list for 1965 for nos between 4001- 
4194, including many ferns. At Kew. 

In 1964 he sent some living plants to Kew 

Commerson, Philibert 

Literature. (1) Add: An appendix to 
Commerson's diary of his journey with Bougain- 
ville consists of a list of plants collected in the 
island of Buru. 

Conklin, Harold C. 

Collecting localities. Add: Mindoro 
(Dec. 7957-Feb. 7955); N. Luzon, Mountain Prov. 
(Feb. 1963) with Buwaya. 

Conover, J. T. 

Biographical data. Portr. in Hatusima, 
Flora Ryukyus 1971, p. 68. 

Conservator of Forests (CF) series 
Add: SuDEN, Suleiman bin Manja, Ujang. 

*Coode, Mark J. E. 

(1937, Plymouth, England; x), botanist, edu- 
cated at Cambridge (B.A. 1961), who made ex- 
peditions to E. Congo (1959) and Turkey (1962, 
1965) ; assisted Dr P. H. Davis with the preparation 
of vols 1 and 2 of the Flora of Turkey, when on the 
staff of the University of Edinburgh. 

In 1966 appointed Senior Botanist at the Botany 
Division, Forestry Department at Lae, Territory 
of New Guinea. Initially working on Melanesian 
Terminalia.^ Author of a Foresters' Manual of 
Combretaceae (1969). 

Collecting localities. 1966. Territory of 
New Guinea: Lae, local collecting (Oct.); around 
Popondetta with J. S. Womersley in the latter's 
numbering series (Nov. 16-18); Lae, local col- 
lecting (Dec.).— 1967. New Ireland"^ with P. Katik 
and T. E. Cropley : around Kavieng (Jan. 22-23), 
Lavongai, c. 26 miles downcoast from Kavieng 
(24); around Katu, Lavongai, Lossuk, etc., based 
at Katu Plantation (26-Feb. 4); to Lemeris and 
Namatanai, a few roadside collections and some 
inland (6-8), round Kavieng (10-11), leaving 
Kavieng for Kaut (13), vicinity of Kaut (14-17), 
back at Kavieng (20) and return to Lae. Terr, of 



Cyclopaedia of collectors — Supplement II 


\ew Guinea: Goroka-Parabo near Okapa East. 
Highlands Distr., with Womersley (18); collecting 
around Purosa (S of Okapa) (19), Okapa environs 
(20), Purosa (22); Gimi Valley, SW of Okapa (23); 
Okapa Valley, SE of Okapa (24); to Lae (25); to 
loma. Northern Distr. (31), loma area (June 1), N 
of loma (2), S of loma (4), Popondetta (5), to Lae 
(6); S. Highlands Distr., with McVean and L. K. 
Wade; Mendi (July 13), Mt Giluwe via Komia and 
Kagaba (14); camp at c. 10500 ft, collecting by 
campsite (15-16); higher altitude collecting (17); 
back to Mendi (18) and Lae (19). New Britain: 
Hoskins on the N. coast, local collecting (Dec. 18); 
Tabai Rikam near Hoskins (19-20), Kwala Kessi 
near Hoskins (21). Lae (22).— 1968. Terr, of New 
Guinea: Kassam Pass (Jan. 8), Kassam (9-16), and 
to Lae (17). Near Lae (Feb.). To Madang (Mar. 5), 
proceeding to Utu; road N of Madang (7), Aione 
on Ramu R. (8); Aione area (9-13), and return to 
Lae (14); Wau (Apr. 22); near Lae (May). To Edie 
Creek (June 6), Edie Creek-Kaindi (7-11); Mt 
Hagen (Sept. 9), Kagaba (11), around Kagaba 
(12-19).— 1969. Via Rabaul to New Ireland: 
Namatanai (Jan. 11), collecting on coast road 
around Karu (11-13), Kavieng (14), Namance I. 
(15-17), Kavieng (18), back via Rabaul (20) to Lae 
(21). To Bulolo-Wau-Kaindi (June 5-6) with P. 
Wardle; laUbu, S. Highlands Distr. (16), Giluwe 
camp at c. 11000 ft, collecting between 11000- 
12500 ft (18-20), lalibu (21), around lalibu and 
Maral (22-24), Mt Hagen (village) (25), Lae (26). 
Short visit to Bougainville (Solomons) collecting 
round Tonoleki (Aug. 9-14). Via Rabaul and 
Kavieng to Einira I. (coll. Sept. 4), via Kavieng (5), 
Taskul I. and New Hanover (6), back to Kavieng; 
Namance I. (8) and Kavieng; down coast road and 
across New Ireland to Lemusmus and back to 
Kavieng (9), Kavieng (10), Rabaul and Lae (11). 
To Bulolo-Wau (Okt. 1 1-12).— 1970. Via Rabaul 
(Jan. 16) to Keravat and back (17); flight to Manga 
(5. New Ireland) for reconnaissance: Tranlan, 
Muliama (20), Danfu bridge (21), up Danfu R., 
camping at 800 ft (22), Danfu area from above 
camp (23-Feb. 20), back to Manga (20), by road to 
Lei (24) and down coast to Matkamlagira (arrived 
27); attempt to cross Weitin-Kamdaru divide 
(some Sands /Jo.y); Matkamlagira (Mar. 3). — ^1971. 
Papua: Tapini (Apr. 29) by helicopter to top Mt 
Strong with P. F. Stevens,^ summit area and upper 
forest (May 1-3); walk to Torura (5), to Tapini by 
helicopter and to Waitapc (6); helicopter to Mt 
Scratchlcy^ (7), flying back (8), Waitape (8-11). 

Collections. Herb. Lae, a.o. 300 nos New 
Ireland: dupl. in Herb. Edinh. etc. 

Literature. (1) Author of 'A new Terminalia 
from Melanesia' (Kcw Bull.; in press). 

(2) Cf. Fl. Mai. Bull, no 22, 1968, p. 1541. 

(3) M. J. E. Coode & P. F. Stevens: 'Notes on 
the Mora of Two Papuan Mountains' (Pap. New 
Ciuin. Sc. .Soc. Proc. 1971, 23, 1972, p. 18-25; 
with Appendix of plants collected above 33(K) m, 
p. 23-25). 

Cook, James 

Literature. (2) Add: J. C. Beaolf.hole: 'The 
Journals of Capl. James Cook. The Voyage of the 
I ndcavour. 1 768 - 1 77 1' ( 1 955 Cambridge, I lakluyt 
Society, vol. 1). 

(6) Add: See also 'Captain Cook's artisLs in the 


Pacific 1769-1 779', compiled by Anthony Murray- 
Oliver (New Zealand 1969, pp. xxiv, 168, 130 pi.). 

Coolidge, Harold Jefferson 

Add: A member of the Pacific Science Council, 
representing the U.S. Nat. Academy of Sciences, 
for many years. After his retirement by the end of 
1971 he was awarded a Honorary Life Fellowship. 


Coomans de Ruiter, Louis 
(d. 1972, Hilversum, Netherlands). 

*Cooper, David Edward 

(1928, Christchurch, New Zealand; x), gradu- 
ated M.Sc. (in chemistry) at Auckland University. 
Successively working in the Organic Dcpt of this 
University, at Auckland Hospital Laboratory, and 
since 1956 with Auckland Farmer's [-rcezing Co. 
Laboratory. He joined the New Zealand New 
Guinea Expedition I%1 (leader C. K. Pun)' 
which was supported by the New Zealand Alpine 
Club and the Mt Everest Foundation. The objec- 
tive was to climb in the Carstcns/ Mts as well as to 
carry out some collecting and scientific work. 
Cooper made the botanical collection. 

Itinerary. 1961. Dutch West New Guinea: 
walking from Tiowe (near the confluence of the 
Ualiem and Gi Maleie Rivers), travelling up the 
Cii Maleie, crossing over into the West Halicm; 
following the West Ualiem to its head and crossing 
the plateau into the iiaga Valley; subsequently to 
the North Wall of the Carstensz Mts (June), 



Flora Malesiana 

[ser. I, vol. 8^ 

crossing the Zendilerong R. and the head of the 
Kemaboe River. 

Collections. About 42 specimens, partly 
sterile and mouldy; grasses and ferns in Herb. 
Canterbury Un. Christchurch; other material in 
Herb. Auckland War Mem. Mus. (on behalf of 
R. C. Cooper), and phytochemical specimens in 
Auckland University (on behalf of Prof. L. H. 
Briggs); some dupl. in Herb. Leyden. 

He brought home living plants, seeds, and 
cuttings also. 

Literature. (1) D. Cooper: 'Some botanical 
and phytochemical observations in Netherlands 
New Guinea "New Zealand New Guinea Ex- 
pedition 1961" ' (Econ. Bot. 25, 1971, p. 345-355). 

C. K. Putt submitted an expedition report to 
the Mount Everest Foundation. 

Copeland, Edwin Bingham 

(d. 1964, Chico, Cal., U.S.A.). 

He wrote a 'Fern Flora of the Philippines' in 3 
volumes (1958-60). 

Biographical data. Am. Fern J. 54, 1964, 
p. 177-188, portr., partial bibliogr. 

*Coppock, Frank 

forest officer in various parts of New Guinea, 
collected some tens of specimens in the NGF 
series, E. New Guinea, at: Keravat {New Britain) 
(1955), Brown River (Oct. 1959), Trans-Vanapa 
(Oct. 1959), and at Sangara, Popondetta (May 
1960). In Herb. Lae. 

*Cordero Jr, Paciente A. 

(1941, Burauen, Leyte, Visayas, P.L; x) was 
educated in botany at the Far Eastern University 
(B.S.); since December 1963 Botanical Collector 
and later Museum Technician in the Botany 
Division (Herbarium) of the National Museum; 
later in charge of the Algal section, a group in 
which he specializes. He is working on his Masters 
degree at Kagoshima University under Dr T. 

Collecting localities. 1963. Manila and 
suburbs, Luzon; Lipuun /., Quezon, Palawan and 
nearby islets, with D. R. Mendoza and R. A. 
Espiritu. — 1964. Palawan (Apr.-June). Batanes & 
Camiguin Islands together with a group of Kago- 
shima University, Japan (Nov. 9-30). — 1965. 
Batan & Sabtan Islands (Apr. 9-May 7) with G. T. 
Velasquez and A. Timbol; Lubuagan, Kalinga- 
Apayao Prov., Luzon, with R. M. del Rosario. — 
1966. Puerto Galera, E. Mindoro, Luzon (with R. 
M. DEL RoSARio and R. A. Espirttu); Bolinao, 
Pangasinan, Luzon. — 1967. Luzon: Dalton Pass 
(with Mendoza) (Mar. 3-18); Biliran I. (May 2- 
June 3); Leyte; Visayas. 

Collections. In the PNH series, mostly 
Algae (> 1000). 

Corner, Edred John Henry 

Later Reader, and in 1966 appointed Professor 
of Tropical Botany. 

He received the Darwin Medal of the Royal 
Society in 1960, and the Linnean gold Medal in 

Leader of several expeditions including the 
Royal Society Expedition to the Solomon Islands 
in 1965.^ 

April 1972 he went for a study visit to Bogor, 
but left for Singapore 3 weeks after his arrival. 

Collecting localities. 1934 add: Jason 
Bay.*— 1941. Langkawi (Nov. 13-22); G. Baling in 
Kedah (25).— 1959. Sarawak; Bako National 
Park.— 1960. Papua, Central Distr.: Waitape 
(5000 ft) (Aug. 24) and probably elsewhere in 
Australian New Guinea. — 1961. N. Borneo, Royal 
Society Expedition to Mt Kinabalu (see also Chew 
Wee Lek).* — 1964. N. Borneo, 2nd Royal Society 
Expedition to Mt Kinabalu (Jan. 14-May 23).»— 
1965. Royal Society Expedition to the Solomon 
Islands,' partly accompanied by T. C. Whitmore 
(see there). 

Collections. Add: Dupl. Mai. Penins. in 
Herb. Edinb. The Royal Society Expeditions 
collections (numbered RSNB and RSS) will be 
distributed from Kew. 

Literature. (1) Add: and of several other 
books on theoretical botany, etc. 

(4) J. H. Kern & C. G. G. J. van Steenis: 'An 
interesting new record from the Malayan beach: 
Spilanthes urens Jacq., its synonymy and distri- 
bution' (Gard. Bull. Sing. 17, 1959, p. 273-275). 

(5) Cf. Fl. Mai. Bull, no 16, 1961, p. 804-806. 
E. J. H. Corner: 'The Royal Society Expedition 

to North Borneo 1961' (Emp. For. Rev. 1962, p. 
224-233, route map, 4 photogr.); ditto 'Reports' 
(Proc. Linn. Soc. Lond. 175, 1964, p. 9-56, 24 

See also 'A discussion on the Results of the 
Royal Society Expedition to North Borneo, 1961' 
(Proc. Roy. Soc. B 161, 1964, p. 1-91). 

(6) E. J. H. Corner: 'Exploring North Borneo' 
(New Scientist no 366, 1963, p. 488-490, 3 photogr.; 
'Mount Kinabalu East' (Sabah Soc. J. 4, 1965, p. 
170-817, 7 fig., 8 pi.). 

(7) A discussion of the Results was organized by 
Corner; published in Phil. Trans. R. Soc. B 255 
(1969) p. 185-631, 196 fig., by several authors. 

Biographical data. Biol. J. Linn. Soc. 2, 
1970, p. 322-324. 

Cowley, Ebenezer 

(see Cycl. Fl. Mai. I, 1, 1950) made an earlier 
(Dec. 1892) attempt to collect sugar-cane varieties 
in Papua (Mabudauan, Samarai, Thursday I.), 
but the material was so thoroughly fumigated that 
not a single bud survived.' 

Literature. (3) Cf. Cane Growers' Quart. 
Bull. Brisbane 15, 1951, p. 37-38. 

*Cox, Elizabeth Monica (nee Howard) 

(1908, Durham, S. Africa; x) got married to Mr 
J. K. Cox in 1931. She made joint collections with 
her husband (see there), both in W. Africa and N. 

*Cox, John Kidman 

(1906, Lahore, Pakistan; x) was educated in 
India and South Australia; in 1931 appointed to 
the Colonial Agricultural Service, posted to the 
Gold Coast; transferred to North Borneo in 1947; 
retired in May 1960 as Director Department of 
Agriculture, Jesselton, N. Borneo. Now settled in 
Devon, England. 

Collecting localities. 1948-58. N.Borneo 
including on Mt Kinabalu. 




Cyclopaedia of collectors — Supplement II 


Collections. Mostly made jointly with his 
wife, E. M. Howard Cox (see there), first in W. 
Africa (nos 1-142, mostly in Kew Herb.), and later 
in A^. Borneo (nos 143-1052). Early Borneo plants 
partly in Herb. Sing., some in Kew (252-257, 261, 
300, 306-347), some in the Brit. Mus. (250, 258- 
266, 301-305, and various ferns), small coll. of 
22 orchids in Herb. Un. Calif., and a collection of 
orchids & dicots (900-1015) in Herb. Leyden. 

Their prime interest was in Orchidaceae of 
which they maintained living collections; her- 
barium specimens were mostly limited to plants 
difficult to identify or which were of special interest. 

His collections are consecutively numbered. 

♦Graven, Lyn A. 

botanical collector and successor of P. Darby- 
shire, joined C.S.I.R.O. in 1964 and collected in 
Bougainville {Solomon Is.) that year. In 1966 in 
New Guinea. See sub R. Schodde, and R. D. 
HooGLAND (Sepik area); last collection.* 

In about 1971 he has been appointed assistant 
in the Land Surveys Division of C.S.I.R.O., 

Literature. (1) L. A. Craven: 'Searching for 
Rhododendrons in New Guinea' (The Rhododen- 
dron 7 (1), 1968, p. 11-16, 3 photogr.). 

Creagh, Charles Vandeleur 

Literature. (1) Add: W. Meijer: 'The 
Contributions of Governor Charles van de Leur 
Creagh to Botanical Exploration in Sabah, 1888- 
1895' (Sabah Soc. J. 5, 1969, p. 67-68). 

•Cropley, Timothy Eric 

(1947, Sydney, N.S.W., Australia; x), medical 
student Sydney University, and a temporary 
member of the staff at Lae; he accompanied M. J. 
E. Coode (see there) with a survey party in New 

Collections. Herb. Lae; New Guinea dupl. 
1967 in Herb. Edinb. 

*Crosbie, A. 

Staff surgeon of the Challenger Expedition, 
1873-76 (itinerary see Moseley). 

Collections. Some dupl. of bryophytes and 
vascular plants of the Aru Is. in Herb. Edinb. 

Cruttwell, Norman Edward Garry 

In 1970 Reverend Canon; stationed at Agaun, 
Daga country since 1966. 

He is interested in Rhododendrons also.* 

Collecting localities. Add: He visited 
the Daga' once or twice a year on foot; later at the 
Head Mission Station in the Daga at Agaun 
(3200 ft). He collected also at Tua (4500 ft), 
Kanasura (2500 ft), Boncndo, Birat (Milne Bay 
Distr.). and Mt Garatun (5000 ft).— ? 1968. 2nd 
Visit to Mt Simpson (Matawan) with Paddy Woods 
(first time climbed from the N. side, this time 
from the W.).—I972. With R. Pullen, J. I-. 
Veldka-mp and P. F. Steve.ns on Mt Suckling in 

Collections.* Dupl. in Herb. Edinb. (1952- 

); partly collected with P. J. B. Woods, 

Galore, e/f. On Kew labels somclimcs N.E.G.C.* 

He collected some !5(X) nos in Mcnapi and Daga 

districts. Recent collections in Herb. Lae. 

Literature. (1) Add: 'Rhododendron time 
in the Daga' (The Rhododendron 6 (3), 1967, p. 
7-9, 3 photogr.); 'The Rhododendrons of the 
High Daga' (The Rhododendron 8, 1969, p. 11- 
14); 'A new Rhododendron' (I.e. (2), p. 8-9, 1 fig.). 

(3) H. Smith: 'Two new gentians from Eastern 
New Guinea' (Kew Bull. 1957, p. 225-227, 2 fig.). 

(4) Cf. Index Coll. Edinb. Herb. 1970, p. 72. 

(5) N. E. G. Cruttwell: 'The Ascent of South 
Suckling' (The Rhododendron Dec. 1972, p. 7- 



Cubitt, George Eaton Stannard 

(d. Sept. 17, 1967, aged 91 years, probably at 
Winchester, England). 

Biographical data. Mai. For. 30, 1967, p. 

Cuming, Hugh 

Collections. In addition to the information 
concerning the numbering given in the Cyclo- 
paedia (Fl. Mai. I, 1), it seems worthwhile to give 
a more detailed extract of the list published by 
Merrill, cf. I.e. Literature (6). 

Collection labelled "Ins. Philippinae 1841": 1- 
434. Vascular cryptogams. Most of^ the species 
from the Philippines, a few from Malacca, Singa- 
pore, etc. Apparently properly labelled. — 435-694. 
Luzon.~-(m-702. Corregidor.—703-l41S. Luzon.— 
1479-1603. A/zWoro.— 1604-1673. Mindanao.— 
1674-1732. 5amar.— 1733-1757. Levte.—ll5S- 
1 789. Cehu.— 1 790- 1810. Negros.—l 811-1857. 
Bohol. — 1858-2153. Philippines, not localized 
(exceptions 2052; 2053-2058).— 2052. Malay Penin- 
sula.— 205^-205%. Singapore.— 2154-2242. Cellu- 
lar cryptogams chiefly from the Philippines. — 
2252-23W. Malay Peninsula.— 2400-2427. Singa- 
pore. —242»-244i. Sunialra.—2444-24(A. St. 



Flora Malesiana 

[ser. I, vol. 8^ 

A. DE Candolle in his Prodromus vol. 17 
(1873) p. 316 gave information on the numbering 
of the 'Plantae Cumingianae' as far as not col- 
lected in the Philippmes. It is less complete, 
however, and partly incorrect. 

Also dupl. in Herb. Edinb. 

Biographical data. Add: In Hemsley & 
Hooker, Biol. Centr.-Amer. 4, 1887, App. p. 133. 

Cunningham (or Cuninghame), James 

His 2nd voyage to China took place in 1700 
(not 1701) o/b Eaton-Fryatt. According to Dandy 
on both voyages Batavia {Java) was touched {not 
in June 1700, however, cf. SL. 3321, f. 52, in which 
letter it is stated that they did not go ashore in 
Borneo either in July of that year). His 2-years' 
imprisonment in Cochinchina took place after the 
massacre at Pulo Condor(e). He arrived at Batavia 
in April 7707. His last letter to Sloane is dated 
Calcutta, Jan. 4, 170|.* 

Collections. In Sherardian Herb. Oxford: 
ferns from Chusan and Malacca.* 

Literature. (3) C/. J. E. Dandy: 'The Sloane 
Herbarium' (1958) p. 117-122, pi. 5; letters in 
Sloane MSS vols 3321, 4025, 4038, 4039, 4040, 
4041, 4064, f. div., from Cuninghame to Petiver 
and Sloane. 

(4) Cf. H. Newman Clokie: 'Account of the 
Herbaria, Dept. Bot. Univ. Oxford' (1964) p. 76. 

Curtis' collector, see Hamad. 

Cuzner, Harold 

(d. Sept. 25, 1956, Manila, Luzon, P.I.) brought 
up in the U.S.A. (B. Sc. 1905). Shortly after 
graduation he came to the Philippines to work in 
the Bureau of Forestry; in 1909 transferred to the 
College of Agriculture, U.P. He resigned in 1953. 

Biographical data. Bull. Nat. Res. Counc. 
Philip. 42, 1958, p. 6 (erroneously giving 1878 as 
year of birth). 

C.W.L., see Chew Wee-Lek. 

Daniker, Albert Ulrich 

(d. 1957, Zurich, Switzerland). 

Biographical DATA.Yierteljahrschr.Naturf. 
Ges. Zurich 102, 1957, p. 373-374; Verb. Schweiz. 
Naturf. Ges. 137, 1957, p. 316-323, bibliogr., 

Dampier, William 

Collections. Add: Also Java specimens in 
Sherardian Herb. Oxford. 

*Dan bin Hj. Bakar, see Forest Dept Sarawak. 

Dansereau, Pierre 

Add: From 1961-66 Ass. Director New York 
Botanic Garden, and adj. Professor of Botany 
Columbia 1961- ; Sr Curator of Ecology 
1966- ; Thomas Alva Edison visiting scholar 
at Cranbrook Institute of Science, 1967-68; at 
present back in Canada, at the 'Institut d'Urba- 
nisme, Universite de Montreal.' 

Biographical data. Who's who in Science 


*Darbyshire, Paul James 

(1937, Cheadle, Cheshire, England; x). Botanical 
Assistant, Royal Botanic Gardens, Kew, 1957-60. 
Joined C.S.I. R.O. Division of Land Research and 
Regional Survey, Canberra, in Nov. 1960, as 
botanical collector; towards the end of 1963 
(71964) he left C.S.I.R.O., and is now living in 

Collecting localities. 1961. Territory of 
New Guinea: Sepik Distr.: (May 30-Aug. 5), 
Aitape Subdistr. with R. D. Hoogland (see 
there); (Aug. 15-Sept. 19) Lumi Subdistr., 
Torricelli Mts {nos 172^86).— 77962 or 1963. 
Coastal area near Port Moresby, from Rigo to 
lokea.— 1964 (or 11963). Safia area. Northern 
Distr., Papua (July-Aug.). 

Collections. Herb. Canberra (C.S.I.R.O.); 
dupl. in Lae, Leyden, Am. Arbor., Brit. Miis., 
Brisb., U.S. Nat. Herb., Herb. Bog., Manila, 
Geneva, Berlin, Edinb. Wood samples deposited 
with the Div. of Forest Products, C.S.I.R.O., 
Melbourne; Dept of Forests, Lae; and C.S.I. R.O. , 

*Darke, Professor 

was assisted by C. E. Ridsdale (see there) for 
making ethnobotanical collections near Cape 
Gloucester, New Guinea, in 1966. 

*Dawa, Mrs E. D., see Philippine National 

*Dawson, F. 

collected Digitaria fuscescens (Presl) Henr. in 
the Lake Archbold Area, New Guinea in August 

He was possibly employed by the Unevangelized 
Fields Mission, Hollandia. 

Collections. Herb. Melbourne. 

Day, Alice W. 

Add: Eria alicae Quisumbing has been named 
after her. 

Day, John 

Collections. Add: Ferns in Herb. Edinb. 

Biographical data. Add: Gard. Chron. 
1888, p. 88. 

*Day, Miss Merle F. 

(born about 1930), from Australia, was em- 
ployed in the Buitenzorg Botanic Garden (Hort. 
Bog.) from about 1955-58. She travelled at least 
in E. Java and Timor. She was a member of the 
'Kebun Raya Indonesia Timor Expedition', Feb.- 
Mar. 1956. 

Collections. Herb. Bog.: at least poor 
specimens from Timor. Some dupl. in other 


collected an unnumbered specimen of Digitaria 
timorensis (Kunth) Balansa and some other 
plants in Singapore (Apr. 17, 1860); in Herb. Paris. 

Delsman, Hendricus Christoffel 
(d. 1969, Netherlands). 



Cyclopaedia of collectors — Supplement II 


♦Dennis, L. H. and G. D. Blaxland 

collected 125 strains of sugarcane in E. New 
Guinea, near Milne Bay, in the Loiiisiade Archi- 
pelago, Trobriand Is., d' Entrecasteaux Is., and 
along the NE. coast of New Guinea to Buna, in 
1908. The collection was made on behalf of the 
Colonial Sugar Refining Co of Australia.^ It is 
unknown to me whether dried specimens were 

Literature. (1) Cf. Hawaiian Plant. Rec. 55, 
1958, p. 213; Cane Growers' Quart. Bull. 15, 1951, 

♦Dentan, R. K. 

An anthropologist from New York, made 
currently ethnobotanical collections in Perak, 
Malay Peninsula, in 7962 (Feb. 8-20, Apr. 6-28). 

He was assisted by Kadim bin Tassim (see there). 

Deschamps, Louis Auguste 

Itinerary. Add: In vol. 1 the itinerary of the 
Voyage is referred to Billardiere. Deschamps 
sailed in the 'Recherche' and the dates from his 
MSS are a little different from Billardiere's, i.e. 
Waigeou (Aug. 16-27, 1793), Bouro (Sept. 3-16). 

Collections. Add: Notwithstanding the fact 
that the British Government had paid the prize- 
money for D.'s captured collection. Sir Joseph 
Banks promised him to send it to France by the 
first opportunity. In spite of Bank's efforts, these 
scientific collections were never returned to 
France as far as known.' The MSS at Saint-Omer 
(France; see Hocquette sub Biogr. Data) must be 
copies of the BM collection of MSS or the reverse. 

Literature. (5) Add: Cf. Sir Gavin de Beer: 
'The Sciences were never at War' (1960) p. 122- 

(7) Cf. also A. Guillaumin: 'Deschamps 
botaniste herborisant en Nouvelle Caledonie en 
1793' (Bull. Mus. Nat. Hist. Nat. 3« ser. no 47, 
1972, Bot. 3, p. 85-87). 

Biographical data. M. Hocquette :'Louis- 
Auguste Deschamps 1765-1842. Sa vie-son oeuvre' 
(Bull. Trimestr. Soc. Acad. Antiq. Morinie, 
Mdmoire, 39, 1970, 77 pp., 10 pi.). 

*Diederix, Dirk Oebele Jan 

(b. 1923), in 1963 Assistant Palynologist with 
Brunei Shell, Scria. He accompanied H. P. Fuchs 
on a trip to Badas F. R. in Brunei. Plants were 
collected with Fuc hs (see there) collecting numbers. 

Dilmy, Anwari 

Add: In 1968-69 for one year in Europe, 
studying new methodologies of ecology. After his 
return Rector of the University at Bandjcrmasin 

Collecting localitifs. 1963. Bali: Mt 
AgungCOct. 1-17); E.Java.G. Baung K)ct. 21).— 
1964. Bali: Mt Agung' and limestone hill Prapat- 
Agung (Mar. 9-Apr. ^)).—J965. Bali (July). 

Collections. Add: Herb. Bo^.: 8'J nos Bali 
1963, collected with Dr. ■[AkNAVs( hi (Rumania) 
and Prof. Jacovliv (Sovjet Russia); 120 nos Bali 
1964,^ with Made Taman and Nkiji. 

Literature. (1) A. 'Pioneer Plants 
found one year after the 1963 Eruption of Agung 
at Ball' (Pac. Sc. 19. 1965. p. 498 501). 



*Dissing, Henry 

(1931, Hansted, Denmark; x) was educated at 
the University of Copenhagen (B.Sc), later 
Assistant at the Mycological Institute. He specializes 
in Fungi (genus Helvella).^ From July 16-Sept. 5, 
1962, he participated in the Danish 'Noona Dan 
Expedition' (see there for Itinerary, Collections, 

Literature. (1) Cf Dansk Bot. Ark. 25 
(1966) p. 1-172. 

*Dizen or Dizon 

collected some tens of plants at Quezon City, 
Luzon, P.I. 

*DockrilI, Alick William 

(born in a suburb of Sydney, N.S.W., Australia) 
had a job in the Lae Herbarium, but returned to 
N. Queensland; author of numerous papers on 
Australian orchids; he did some general botanical 
collecting in the NGF series in the Morobe and 
Sepik districts of New Guinea; in Herb. Lae, 
dupl. elsewhere, also in Herb. Edinb. 

Docters van Leeuwen, Willem Marius 

(d. 1960, Leersum, U., Netherlands). 

Collecting localities. Delete: Bali etc. 

Literature. (4) must be deleted. 

Biographical data. Vakbl. Biol. 40, 1960, 
p. 85-86; Proc. Linn. Soc. Lond. 172, 1961, p. 
131-132; Studentenalmanak Amsterdam 1961, p. 
240-241, portr. 

Docters van Leeuwen-Reynvaan, Jenny 
(d. 1963, Amersfoort, U., Netherlands). 

Donk, Marinus Anton 
(d. 1972, The Hague, Netherlands). 

*Dornstreich, M. D. 

made an ethnobotanical collection in the 
Territory of New Guinea, >246 nos, numbered in 
the NGF-M.D.D. series. East Sepik region was 
visited in Sept. 7965. 

Collections. In Herb. Lae; some dupl. in 
Leyden Herb. 

Doux, J. A. le, see Ledoux, J. A. 

•Dozy, Jean Jacques 

(1908, Rotterdam, Holland; x), geologist edu- 
cated at Leyden University (Ph.D. 1935); in the 
same year he entered the service of the B.P.M. 
(Royal Dutch Oil Co.), being stationed respec- 
tively in Dutch New Guinea (1936-carly 1937), 
Guatemala (1937-39), Ecuador (1949-43), Persia 
(1944), U.S.A., Venezuela (1946-47). The Hague 
(Holland). Indonesia (1949-54). and since May 
1955 in Ihe Hague as Head of the Exploration 
B. Int. P.M. 

In 1936 he accompanied Colijn's expedition to 
Carstens^ Mts, West New Guinea (for itinerary cf. 
sub V. J. WissLL, who took care of botanical 

Collections. 18 Alpine plants from Carstcns 
Mts in private herbarium, brought home for his 
wife. The collection is occasionally on loan for 
study in Leyden Herbarium, and will in the future 
be presented to that institution. 



Flora Malesiana 

[sen I, vol. 8^ 

*Dransfield, John 

(1945, Liverpool, England; x) graduated at 
Gonville and Caius College, Cambridge (1967 
B.A., 1st class in Botany); Sept. 1967-Sept. 1968 
research on the Malayan palms Eugeissona and 
Johannesteijsnwnnia in Malaysia, stationed at the 
Univ. of Malaya, Kuala Lumpur; research on the 


above palm genera in the Botany School at 
Cambridge, under the supervision of E. J. H. 
Corner, Oct. 1968-Sept. 1970; Ph.D. Univ. 
Cambridge Dec. 1970. In October seconded by the 
British Government to the Indonesian Government 
under the Technical Cooperation Programme of 
the Colombo Plan, to work in the Lembaga 
Biologi Nasional, Bogor, Indonesia, at least up till 
Sept. 1972. 

He is doing taxonomic research into rotans and 
other palms, and most of his publications deal 
with palms. ^ 

Collecting localities. 1967. Malay Pe- 
ninsula. In Oct. Selangor, Perak, Johore; Selangor 
(Nov.); Negri Sembilan, Selangor in Dec. — 1968. 
In Jan. Selangor, Cameron Highlands; Taman 
Negara and Pahang (G. Tahan)^ with T. C. 
Whitmore (Feb.-mid Mar.). Sarawak and Brunei 
(Bako National Park, G. Mattang, Bintulu, 
Andalau, Tutong) (mid-Mar.-Apr.). Malay Penin- 
sula: in May G. Blumut, Johore, with T. C. 
Whitmore; Selangor; Kledang Saiong and Lumut 
in Perak, and Eraser's Hill in Pahang in June; 
Johore and Selangor in July; Trengganu, Selangor 
in Aug. — 1970. Indonesia, W. Java (several times 
in Kebun Raya): Tjibodas (Oct. 11, 15); Janlappa 
F. R. and Djasinga (Oct. 17); Tjibodas (Oct. 24); 

Langkong, distr. Sukabumi (Nov. 13); Tjiletuh 
(G. Handjuang F. R. Distr. Sukabumi) (24-26); 
Telaga Warna and G. Geger Bintang (Dec. 6); 
Pelabuan Ratu (11), Tjibodas (15), G. Salak 
(Tjitjurug) (20), Tjisolok (22), Tjibodas (25).— 

1971. Tjibodas (Jan. 8); Tjibalanac, Tjipatudja, 
distr. Tasikmalaja. S. Sumatra: Kepahiang, 
Tjurup (Feb. 1 2-1 3) ; Lampong Distr. : Waikambas 
(16), Karang(b)erak S of Kota Agung (20-22). W. 
Java: Situ Patengan (27), G. Tangkuban Prahu 
(28); E. Java: Nglijep, G. Tengger, Sumber 
Berantas (Mar. 15-20); W. Java: Nature Reserve 
Depok to hunt for Orania (24), Pelabuan Ratu 
(27), G. Salak (ridge above Tjiomas; Apr. 3); 
Udjong Kulon (Apr. 12-22); Dungus Iwul, 
Djasinga (May 15); Pelabuan Ratu (22). Kali- 
mantan Timur (= E. Borneo): Balikpapan, Samar- 
inda, Kutei Nature Reserve (May 26-June 24). 
W. Java: G. Pangrango (July 3); Lengkong and 
Tjiletuh, distr. Sukabumi (July 15-18); G. Salak, 
gorge above Tjiapus (25). A'^. Sumatra: Atjeh 
(mid-Aug.-Sept.) with a Japanese expedition. — 

1972. Centr. Sumatra: Tjagon Alam, Berbak near 
Nipah Pandjang, Djambi; trip from Djambi, via 
Bangko to Sg. Penuh, then to Kaju Aro tea estate, 
Barisan Range; waterfall on Btg. Sangir, lower 
slopes Mt Kerintji and Danau Bento; returned to 
Tapan and Bt Tinggi to Padang; with E. de Vogel 
to S. Sumatra: Lampong Districts and S. Ben- 
coolen. With E. de Vogel to SE. Borneo. 

Collections. Largely palms, but occasion- 
ally flowering plants and many bryophytes, lichens 
and Fungi. In Malaya nos 450-922; in Indonesia 
from 950 onwards (in July 1971 up to 1779). 
From Malaya 1st set in Kew, dupl. to Ithaca, 
Leyden, Kepong, Singapore, Univ. of Malaya; 
from Indonesia 1st set in Herb. Bog., dupl. in time 
to Leyden, Kepong, Singapore, Univ. of Malaya. 

Literature. (1) J. Dransreld & T. C, 
Whitmore: 'A Podostemacea new to Malaya. 
Indotristicha malayana' (Blumea 18, 1970, p. 
152-155, pL l,fig. 1). 

(2) J. Dransfield: 'The Javanese palm flora, 
first impression' (Berita Biologi 1, pt 3, 1971, p. 
21-25); 'The genus Johannesteijsmannia H. E. 
Moore Jr' (Gard. Bull. Sing. 26, 1972, p. 63-83). 

Drescher, Friedrich Carl 

(d. Dec. 19, 1957, Djakarta, Indonesia). 

Biographical data. Treubia 24, 1958, p. 
131-134, portr. 

Druce, George Claridge 

Add: In the same year he visited India, Ceylon, 
China, Korea, and Japan. 

Biographical data. Add: Cf. H. Newman 
Clokie: 'An Account of the Herbaria of the Dept. 
of Botany in the University of Oxford' 1964, p. 
49-52; bibliogr. in Rep. Bot. Soc. and Exch. 
Club Brit. Isles for 1931, p. 804. 

Dumont d'Urville, Jules Sebastien Cesar 

Literature. (4) Add: For drawings of the 
Botany of the Coquille see Cl. Nissen, 'Die 
botanische Buchillustration', 1951, vol. 2, p. 53 
sub Duperrey. 

*Dunstone, E. 

collected few numbers in the NGF series; Herb. 




Cyclopaedia of collectors — Supplement II 


Durant, C. L. 

(see Cycl. FI. Mai. I, 1, 1950) is credited with 
the collecting of some Ficus specimens in Java. 

*Duuren, G. J. van 

collected Loranthaceae in the vicinity of Ukua 
Estate, via Kairuku, Papua, at least between 

Collections. In Brisbane Bot. Dept. and in 
Herb. Univ. Queensl. 

Literature. (1) Cf. B. A. Barlow: 'Contri- 
butions to a knowledge of the Loranthaceae of 
New Guinea' (Proc. R. Soc. Queensl. for 1963, 75, 
1964, p. 13-18, 3 fig.). 

Dw>er, R. E. P. 

(d. Oct. 3, 1959, Sydney, Australia) was finally 
Director of Agriculture, Stock and Fisheries, 
Territory of Papua. 

Biographical data. Papua & New Guinea 
Agric. J. 12, 1959, p. 47-48, portr. 

♦Dyhrberg, A. S. 

Danish zoologist who was for several years 
employed at the Zoological Museum of Indonesia 
at Bogor. Later Chief Display Division of the 
National Museum of Canada at Ottawa. At present 
Director of the Zoo at Copenhagen. 

C o L L E c T I o N s . Collect ions of botanical material 
made during his employment in Indonesia were 
made in co-operation with Dr Kostermans and 
with staff instructed by him at P. Dua, and 
Udjong Kulon, S^V. Java. Files, data etc. ac- 
cording to Mr Dyhrberg have been kept by 
Kostermans. D.'s specimens scrappy and mostly 
thrown away. 2500 plants from Sumatra (Koster- 
mans never collected there) were presented to 
Copenhagen in 1961 ; I have no information on the 
state of the collection. 

Dijkstra, F. 

of the Forestry Service, New Guinea, cf. P. van 
ROYEN 1961. 


a Papuan who collected some numbers in the 
series of T. G. Hartley (see there). 

Edaiio, Gregorio E. 

(d. 1960). 

Collecting localities.* Add: Sulu and 
Job (Feb. 1957); collecting till 1959. 

Collections. He collected mosses too.' Also 
dupl. in Herb. Edinb. [Philip. Is.), partly with J. 

Literature. (1) For the main collecting 
places cf. Philip. J. Sc. 89, 1960, p. 285-286. 

(2) Cf. A. NrxjucHi in Sc. Rep. Tohoku Univ. 
scr. iv (Biol.) 29. 1963, p. 145-151, 23 fig. 

Biographical data. Philip. J. Sc. 89, 1960, 
p. 283-286, portr, 

*Eddowes, Peter 

Utilization Officer, New Guinea, made col- 
lections from various parts of Papua and New 
Guinea, in the 1960s, a.o. islands in Milne Bay 
Distr.; Herb. lute. 

In 1968 attached to the Forest Products Re- 
search Centre, Department of Forests, Hohola, 
Port Morciby, Papua. 


*Eddy, Alan 

in charge of the mosses in the British Museum, 
joined an expedition to the Finisterre Mts etc. in 
1964-65 (see sub A. C. Jermy and others). He 
was to collect mosses, liverworts, and lichens, but 
might have collected some phanerogams. In 1969 
he accompanied Dr A. C. Jermy on an expedition 
to Central Celebes (Latimodjong Mts) and S. New 
Ireland (Bismarck Archipelago). 

Eder, M. P. 

(Suppl. Cycl. Fl. Mai. I, 5, 1958, p. cclxxiv) is 
probably identical with Eden (Cycl. Fl. Mai. I, 1, 
1950); collected in the FB series. 

Edwards, Joshua Price 

(d. Feb. 21, 1966). 

Finally Director of Forestry, Federation of 
Malaya; retired in May 1953. 

Biographical data. Mai. For. 15, 1952, 
p. 177, portr.; ibid. 29, 1966, p. 3-5. 

♦Edwards, P. J. 

collected with P. J. Grubb (see there) in E. 
New Guinea, 1970-71. 

E^houd, Jan Piet Karel van 
(d. 1958, HoUandia, New Guinea). 

*Ehrendorfer, Friedrich 

(1927, Vienna, Austria; x), botanist educated at 
Vienna University (Ph.D. 1949), and attached to 
the Botany Department there from 1949-64 
(Assistant, later Keeper). From 1955 teaching 
Systematic Botany, in 1962 Extraordinary Profes- 
sor in Vienna. Visited the U.S.A. in 1952 and 1959. 
End 1964 Ordinary Professor at Graz University 
and from 1965 Head of the Institute for Systematic 
Botany and Director of the Botanic Garden, 
University of Graz; from 1971 Professor at Vienna 

In Sept. 1964 he was awarded a Senior Fellow- 
ship of the Australian Academy of Science and a 
travel grant, which did materialize only in the 2nd 
half of 1966.* 

His main interests are systematics, phylogenetic 
affinities and evolutionary patterns. The outcome 
of his studies resulted in several papers. 

He made numerous excursions in Europe and 
the Mediterranean. 

Collecting localities.' 1966. En route 
visiting Thailand, Singapore (Aug. 16-20), col- 
lecting during various excursions on the island. 
From Queensland paying a visit (Aug. 30-Sept. 11) 
to Papua and New Guinea: around Lae, Goroka, 
Wau, Bulolo and Port Moresby. 

Collections.' Mostly made in Australia, but 
also in Thailand and Malcsia (sec above). New 
Caledonia, and Ceylon. His Singapore plants are 
numbered 6600/1301 to 6600/1905; the New 
Guinea plants from 6600/3901 to 6600/5713. At 
present in the Botanical Institute, but private; 
presumably the 1st set will go to Vienna; many 

I^rriKATURE. (I) Cf Fl. Mai. Bull, no 22. 
1968, p. 1539-1540. 

•Eichler, Hansjocrg 

(1916. Ravensburg, SW. Germany; x), botanist, 
educated at Berlin, Vienna and Halle Universities 



Flora Malesiana 

[ser. I, vol. 8^ 

(Dr. rer. nat., Halle 1950; Ph.D., Adelaide 1959). 
From 1936-43 he worked in the Berlin Museum, 
from 1944-53 in the Kaiser-Wilhelm-Institut fiir 
Kulturpflanzenforschung at Vienna; in 1953 a 
guest at the University of Parma, and from 1954- 
55 a guest at the Rijksherbarium Leyden, revising 
Maiesian Ranunculaceae with a German research 
grant. Since 1955 Keeper of the State Herbarium 
of South Australia, Adelaide. In 1973 Curator of 
the C.S.I. R.O. Herbarium, Canberra. 

Collecting localities. 1965. Territory of 
New Guinea: Aiyura (Apr. 15), Kainantu (15-16), 
Gabari near Lae (18), Keglsugl-Mt Wilhelm (22- 
28), Tomba, along Mt Hagen-Wabag road (30), 
Kubalis, Wabag-Laiagam road; Lake Iviva; 
Lagaip-Kandep divide (May 1), Merimanta; Lake 
Birip (2). 

Collections. Collected widely in Central and 
Southern Europe, but lost first collection (about 
8000 nos) in Berlin, 1943; new collection c. 19.000 
of which c. 7000 are Australian. About 100 New 
Guinea plants, nos 18186-18287, in Herb. Lae and 
Adelaide. Selected dupl. were sent to other herbaria. 

Elbert, Johannes 

Literature. (3) Add: Cf. also B. Hagen in 
Peterm. Geogr. Mitt. 56,^ 1910, 306-308, w. ill. 
and map. 

*Elleh, Saju, see Forest Dept North Borneo. 

Elmer, Adolph Daniel Edward 

Add: Collected in California prior to 1903. 

Collections.* Herb. Edinb.: dupl. Philippines. 

Literature. (8) A fairly complete set of some 
10.000 handwritten original field notes is among 
the Reliquiae Bartlettianae at Michigan. (5549- 
6768 very incomplete, 6917-6961, 7037-18480, 
20003-22694), obtained from Elmer's widow. 
Merrill made typed copies of the nos 7037-18477, 
presumably at Berkeley, Harvard, and Kew. 

Biographical data. Add: In J. H.Thomas, 
Flora S. Cruz Mts California, 1961, p. 35, photogr. 
p. 33. 

*EIsener, Adolf 

(1932, Ibach-Schwyz, Switzerland; x) amateur 
botanist who studied theology; in the Franciscan 
Order. From 1961 stationed in different localities 
in West Borneo as 'Pater Agatho'. He is an 
excellent plant photographer and occasionally has 
made herbarium specimens, although lack of time 
of late {vide letter of June 1969) prevents these 

Collecting localities. 1964. W. Borneo 
{Kalimanten): NE. G. Raja, Njaumkop, 7(X)m 
(Aug. 15, Sept. 14). 

Collections. In Herb. Leyden, small col- 

*Elsworthy, G. 

Cape Rodney, Papua; is mostly interested in 
orchids, but also collected Gesneriaceae and a few 
other plants. 

Collecting localities. 1962. Papua: 'Edit 
Creek, 2 weeks in Musa Mts (Oct.-Dec.) (Owen 
Stanley Range). 

Collections. 400 nos living orchids in 
Edinburgh Botanic Garden. 


♦Enoch, I. 

from the Botany Department University of 
Malaya, collected in Pahang (1954), with J. 
Carrick (see there) in Sarawak {1959). Possibly on 
several occasions. 

Enriqucz, Colin Metcalfe (Dallas) 
Add: (b. 1! 

Enriquez, Gloria L. 

graduate student of zoology, later working on 
microbiology in the U.S.A. 

Ernst, Alfred 

(d. 1968, Zurich, Switzerland). 

Biographical data. Vierteljahrschr. Nat- 
urf. Ges. Zurich 113, 1968, p. 371-371a, 417-418; 
Fl. Mai. Bull, no 23, 1969, p. 1669; Verh. Schweiz. 
Naturf. Ges. 1969, p. 267-268; Osterr. Akad. Wiss. 
119, 1970, p. 337-342. 

Esche, H. 

of the Indonesian Forest Service (see Suppl. 
Cycl. Fl. Mai. I, 5, 1958, p. cclxxvii). 

Collecting localities. Add: 1953. W. 
Borneo (Nov.): subdistr. Semitau, at Sintang; 
subdistr. Upper Kapuas, village of Nanga Njaban. 

Collections. Herb. Bog.: W. Bornean 
plants; Maiesian dupl. in Herb. Leyden (> 100). 

Eschscholtz, Johann Friedrich 

Biographical data. Add: Occ. Pap. Calif. 
Ac. Sc. no 20, 1943, p. 27-28. 

*Esmade, F. 

Superintendent Tungao Operation. 
Collecting localities. Mindanao: Agu- 

Collections. In U.S. Nat. Herb. Wash. 

Espinas, Anachto 

(see Suppl. Cycl. Fl. Mai. I, 5, 1958) numbered 
in the FB series. 

*Espiritu, Rogelio A. 

(1938, Sta. Cruz, Manila, Philippines; x) joined 
the National Museum, Philippine National 
Herbarium, in 1960 as Botany-Aide. From 1963 
onwards he accompanied H. G. Gutierrez, D. R. 
Mendoza, p. a. Cordera, R. del Rosario, M. 
Jacobs, and E. J. Reynoso on collecting tours in 
the Philippines. No field numbers of his own. 

♦Evans, G. B. 

collected ferns^ on Ipoh Temple limestone 
(Perak) and on Bt. Takun in Selangor, Malaya, 
early 1966. Possibly ferns only. 

Literature. (1) Cf. Chin See Chung: 'The 
limestone Flora of Malaya' (Academic exercise, 
Univ. Malaya 1973, p. vii). 

Evans, Ivor Hugh Norman 

(delete the asterisk in Suppl. Cycl. Fl. Mai. I, 5, 

(b. 1886, Cambridge, England) came to South- 
east Asia in 1910 in the service of the Chartered 
Company of British North Borneo. In April 1912 
appointed Assistant Curator and Ethnographical 
Assistant at the Perak Museum, Taiping, Malaya; 



Cyclopaedia of collectors — Supplement II 


from 1920 Ethnographer at the Federated Malay 
States Museum. After his retirement (1932) he 
settled for a time in Suffolk, but returned to Kota 
Belud in Borneo in 1938, resuming serious anthro- 
pological work with a study of the religion of the 
Dusuns; interned during the war. After 1945 the 
greater part of his life was spent in Labuan, but 
following 1946 he returned for a period to Kota 

Collections. From at least 1914 onwards in 
Malaya, partly with FMS nos. 

Biographical data. Sarawak Mus. J. 8 
(n.s. 11), 1958, p. xviii-xx; J. Mai. Br. R. As. Soc. 
33, 1960, p. 109-110. 

•Everaarts, Ary Pieter 

(1950, Maassluis, Z.H., Netherlands; x) student 
in tropical horticulture at the Agricultural College 
Wageningen, made collections during a study-stay 
in Indonesia. 

Collecting localities. 7972. W. Java 
(April): Tjipanas, Lembang. SE. Kalimantan 
(Borneo): Meratus Mts, G. Sarempaka, together 
with E. F. DE VOGEL. 

Collections. Java plants, nos 1-1 10 in Herb. 
Bog., dupl. in Leyden and Wageningen. Borneo 
plants, nos 1585-2142 in ser. de Vogel, in Herb. 
Bog., dupl. in Leyden and Wageningen. 

Everett, Alfred Hart 

Add: (1848, Norfolk I., Pacific; 1898, London, 
England), whose father ran a Penal Colony, sailed 
with the family to England about 1851. 

He first interested Charles Hose in collecting 
and studying (C. F. Cowan in litt. Feb. 12, 1968). 

Collections. Add: Herb. Edinb.: plants 
from Natuna Is (1893). 

Biographical data. Add: Novit. Zool. 5, 
1898, p. 606; A. A. Prestwich, I name this 
parrot . . . , 1963, p. 33-34. 

•Everett, B. 

returned to the U.K. in August 1970 after a year 
at Kepong (1969-70) as a volunteer under the 
British Service Overseas arrangements. He spent 
much of his time collecting and helping prepare 
drafts for the Tree Flora of Malaya. 

Everett, Harold Hart 

(d. c. 1932, London), younger brother of Alfred 
Hart Everett. Primarily a planter and anthro- 
pologist. He settled in Sarawak about 1871, where 
he pioneered planting Cutch. Joined by Ernest 
Hose. 1896 (C. F. Cowan in litt. Feb. 12, 1968). 

•Ewart, J. W. 

collected some plants from P. Penang (Sept. 24, 
1938): bad material in Herb. Sing. 

I- and FA numbers from Sarawak, sec Fl. Mai. 
Bull, no 21, 1966, p. 1433. 

•Faber, P. J., see Boswczen Nicuw Guinea. 

•Fabia, M. P., sec Forest Dept North Borneo. 

lairchild, David Grandison 

Bio(;raphical data. Add: Huntia I, I9M, 
p. 71-78, porlr., 79-102. bibliogr. 

Farquhar, William 

Biographical data. Add: Portr. in Str. 
Times Annual for 1971, p. 60. 

FB nos, see Forestry Bureau Manila. 

*Fedorov, A. A. 

Curator of the Leningrad Herbarium, joined 
Kostermans (see there) on a Siimbawa expedition 
in 196L- Evidently he previously visited Ceylon.^ 

Literature. (1) Author of The Dipterocarp 
Equatorial rain forest of Ceylon' (Trans. Mosc. 
Soc. Nat. 3, 1960, p. 305-332, 9 fig.; in Russian w. 
English summary). 

(2) A. A. Fedorov: 'Botaniceskie putesestvia 
Sovetskie botaniki v Indonezii' (Bot. Zurnal 47, 
1962, p. 1844-1849, 5 photogr.). 

Feen-van Benthem Jutting, W. S. S. van der 

Add: End 1964 she retired on pension, and was 
appointed Honorary Doctor of the Just. Liebig 
University at Giessen, Germany. 

Biographical data. Beaufortia no 130, vol. 
11, 1964, 21 pp., w. portr. a. bibliogr.; in Folia 
Civitates (University of Amsterdam) 18,'^ 16 Jan. 

Finet, Achille Eugene 

Biographical data. Add: Adansonia 2, 
1962, p. 147-159, portr. 

Finlayson, George 

Collections. .\dd: Dupl. Mai. Penins. in 
Herb. Edinb. 

Finsch, Friedrich Hermann Otto 

Biographical data. Add: A. A. Prest- 
wich, I name this parrot . . . , 1963, p. 34-35. 

FL numbers are part of the CF series (Kepong) 
and must be cited as CF (see Fl. Mai. Bull, no 21, 
1966, p. 1433). 

*Flach, M., see Boswezen Nieuw Guinea, 

Flemmich, C. O. 

left Singapore under the Malayanisation scheme 
on Feb. 24, 1960, for Australia. He later returned 
to England. 

Shorea Jlemmichii has been named after him. 

Biographical data. Mai. For. 23, 1960 
p. 137. 

*Flenley, John Roger 

(1936, Ormskirk, England; x), palaeo-ecologist, 
who graduated, B.A. in Botany, at Cambridge 
(1958); Ph.D. at Canberra (1968).' He was a 
member of the Cambridge Botanical Expedition 
to Ethiopia in 1957, and collected under Dr D. 
Walkkr (sec there) in New Guinea. He received a 
grant to visit Malaya in connection with pollen 
analysis. Lecturer in iiiogcography. Department of 
Cjcography, University of Hull, England. He 
makes an investigation into Ouatcrnary vegela- 
tional history in SV:. Asia, performing field work 
in Indonesia, in 1972 accompanied by a Research 
Assistant R. J. Morlky. 

Coi.i.E< I iN(i i.ocALiriis. 1964 65. Terr, of 
New Guinea: Wabag region* and short visit to Mt 




Flora Malesiana 

[ser. I, vol. 8^ 

Wilhelm.— 1969. Malaya: Tasek Bera (Feb.).— 
1972. Indonesia. W. Java: Jakarta (May 5), 
spending several weeks at Bogor Botanic Gardens, 
visiting swamps and lakes. Flying to Sumatra 
(June 10) with Morley and M. KosiM Kardin, a 
graduate biology student of Bandung as field 
assistant and interpreter. Sumatran swamps and 
lakes were studied, viz: N. Sumatra: Laut Tawar, 

Ronggurnihuta swamp, Lake Sipingan near 
Siborong-borong; W. Sumatra: Telaga Batu Piano 
near Bukittinggi; Rawang Biluluk, Talu Dalam, 
Telaga Danau Kacik (= Katjik), and Danau di 
Atas, all near Alahan Pandjang ; Central Sumatra, 
Djambi: Danau Bento, and Danau Padang, in 
Kerinci (= Kerintji); Rawang Payo, Rawang 
Pulau Tengah, Rawang Sikijang, Rawang Bukit 
Bintang, all at Lologedang, Kerinci; Danau 
Gunung Tujuh, Kerinci; Danau Landah Panjang, 
and Danau Sati, G. Kerinci. W.Java: Situ Gunung, 
Ranoe Upas. Return to Bogor by Sept. 13, and to 
Britain on the 21st. 

Collections. //er6. Univ. Ccnfteno, numbered 
in the ANU series (Austr. Nat. Univ.), ANU 
2000-2900; dupl. in Lae, Leyden, Kew, Am. Arbor. 
Ferns only sent to New South Wales. Plants from 
Malaya in Univ. Kuala Lump., Kew, dupl. Leyden. 

In 1972 671 nos by R. J. Morley, J. R. Flenley 
& M. K. Kardin, 1-647 from Sumatra, 648-671 
from W. Java; in Herb. Hull Univ., dupl. at 
Leyden, Bogor, Kew, Sing., Bandung, Am. Arbor. 
The Sumatra collection includes the first record of 
Isoetes for that island. Mosses were collected from 
all ecological sites, and also at Lake Toba and on 
G. Pangrango (W. Java). 

Literature. (1) On a thesis: 'The present and 
former vegetation of the Wabag region of New 

(2) J. R. Flenley: 'The vegetation of the Wabag 

region, New Guinea Highlands: a numerical 
study' (J. Ecol. 57, 1969, p. 465-490, 8 fig., 4 tab.). 

♦Flood, L 

P. W. Leenhouts found a specimen of Atalaya 
salici folia collected in Timor, in Herb. Kew. The 
specimen cited by Danser (see Cycl. Fl. Mai. I, 1, 
1950, p. 168a) is also at Kew. Probably both 
specimens were collected by the same man who 
possibly collected plants as a sideline only. At 
least no Flood is mentioned in the list of col- 
lectors whose plants are at Kew (Kew Bull. 1901, 
by B. D. Jackson). The zoological collector Flood 
of the early 20th century will have nothing to do 
with it. 

Forbes, Henry Ogg 

Collections. Herb. Leyden, add: 461 New 
Guinea plants (purch. 1888); Herb. Leningrad: 
New Guinea dupl. 

His numbers run as follows: 1-137 (Bantam, 
W. Java), 138-166 (Cocos Islands, Indian Ocean), 
167-602 (Kosala, Bantam, W. Java), 603-1441 
(W. Java, by error marked SE. Java), 1442-1897 
(S. Sumatra), 1898-2161 (S. Sumatra, Blalau 
region), 2162-2699 (5. Sumatra, Pasumah Lands), 
2700-2911 (5. Sumatra, Musi region), 2912-3256 
(S. Sumatra, Upper Musi region), 3257-3320 
(Moluccas, Amboina), 3321-3419 {Timor Laut 
Is. = Tanimbar Is.), 3420-4159 {Timor, Lesser 
Sunda Is.); 1-955 {Papua). 

Biographical data. Add: Ibis, ser. 13, vol. 
3, 1933, p. 135-136. 

*Forest Department Brunei'^ 

P. S. Ashton*, and Hasan Pukol collected on 
behalf of this department. 

Forest Department North Borneo (Sabah)^ 

at Sandakan. To the list of collectors mentioned 
in Suppl. Cycl. Fl. Mai. I, 5, 1958, p. cclxxvii, add: 
Agim bin Ambullah, J. Ampuria, Angian, James 
Awing, Aban Bebob, D. Brand, A. Buntar, 
Charington, J. Chong, Saju Elleh, M. P. Fabia, 
A. GiBOT, A. Hashim, p. Hee {1928), Jaswir, A. 
Jawanting, Kamis {1935), Kandilis {1936), 
Kapis bin Sisiron & Kilang (Meijer's collectors), 
HuJAH Kabun {1931), V. Kinted, Kuripin, J. K. 
Lajangah, Lee Koh Hat {1935), Lunau {1932), 
L. Madani, S. Mohammed Maidin {1931, 1934), 
W. Meijer*, Michael Anak, G. Mikil, Muin bin 
Chai, M. a. Mujin, Nordin (w. Singh), 
Othman*, Patrick Ping(kun) {1938), Abdul 
Rahim* {1949), Masirom Rundi, Patrick P. 
Sam, Sikajat {1939), H. T. Sinanggul, J. singh, 
PuTAN TiKAU, Suah Tingguan, James A. Wing, 
Y. K. Wong, Yasin Dangi, Zeano. There cer- 
tainly will be many others. 

*Forest Department Sarawak 

The herbarium of the Sarawak Museum is 
amalgamated with that of the Department. The 
numbering of the Department has been very 
confusing and it is recommended to read the 
article on it by P. S. Ashton (Fl. Mai. Bull, no 21, 
1966, p. 1432-1435). 

(1) An asterisk refers to a separate entry with more particulars. 



Cyclopaedia of collectors — Supplement II 


Recent (1960s) in the Forest Department: 
Ardzi bin Arshid, Ariffin, Bojeng bin Sitam, 
Hj. Bujang, Dan bin Hj. Bakar, Galau, A. 
MuAS, Ilias Paie, Rashid bin Taggoi, ROSLl, 
Rehal, Sainuddin bin Bolhassan, Wasli bin Hj. 

Forest Research Institute, Bogor 

Add: JoENOES (or Junus Dali) {Peutjang I. 
near W. Java 1957; S. Sumatra 1958). 

Collections. Add: As to the numbering of 
wood samples taken with the herbarium specimens, 
and the list which makes it possible to match them, 
see Fl. Mall Bull, no 22 (1968) p. 1545-1546. 

Forest Research Institute, Kepong 

See sub Conservator of Forests series (Cycl. Fi. 
Mai. I, 1, 1950), KEP series (up to 1966), and the 
recent FRI series. 

Forestry Bureau, Manila, P.I. 

Add as collectors in the FB series: J. Baldemor 
{Luzon 1915), J. R. Gillis {Corregidor 1920), J. 
Lazaro (1916), M. Simeon (Luzon 1921). 

Forsten, Eltio Alegondas 

MiQUEL was wrong in ascribing a plant from 
P. Dwars in de Weg as collected by Forsten (c/. 
Cycl. Fl. Mai. I, 1, 1950, p. 179 sub Lit. 3). 
Rightly the plant was collected by J. C. van 
Hasselt and the original label of the collection 
was one of Kuhl & Van Hasselt. 

Forstman, Miss A. G., see Vorstman, A. G. 

Fraga, Z. C. 
see sub E. J. Reynoso, 1965. 

♦Frake, Charles O. 

collected some plants in the PNH (Philippine 
National Herbarium) series in 1957. Grasses and 
sedges only? From Mindanao. 

Frey-Wyssling, Albert Friedrich 

Dr. h.c. at Nijmegen, Netherlands. 

Collecting localities. Add: 1922. Su- 
matra Exist Coast: Sg. Pantjar. 

Collections. From above-mentioned local- 
ities in Herb. Ziirich and Leyden. 

FRI nos, see Forest Research Institute, Bogor. 

FRI scries 

of the Forest Research Institute at Kepong 
replaced the KEP series in 1966. 

Friedberg, Claudinc Berthe n6e, c/. sub Berthe. 

•Frodin, David Gamman 

(1940, Chicago, III., U.S.A.; x) botanist, 
educated at the universities of Chicago (B.S. 
1963), Tennessee (M.S. 1964), and Liverpool 
(M.Sc. 1965). In autumn 1967 he started work on a 
Ph.D. with Prof. Corner at Cambridge University 
(England), concluding in Nov. 1970 with a thesis 
on Schefflera. From (Jet. 1965 to Oct. 1966 
employed in the Division of Botany, Lac, New 
Guinea, as a temporary scientific olliccr. Subse- 
quently he visited a number of herbaria, and made 
short Held trips in Australia and the Far East (up 
to June 1967). From February 1971 Lecturer in 


Botany in the University of Papua & New Guinea 
at Boroko, near Port Moresby. In addition to 
teaching systematics and ecology, it is planned to 
continue studies on Schefflera and other Araliaceae 
and to publish a detailed account of collectors 
(1871-1942) in eastern New Guinea. 


Courtesy E. F. Greenwood, Keeper of Botany, 
City of Liverpool Museums. 

Collecting localities. 1965. New Britain 
(Bismarck Archipelago): for 2 weeks (Nov.) with 
W. Zavattard (a timbermerchant)lromKandrian 
by boat along the S. coast to Gilnit on the Itni R., 
thence on foot across the island by way of Airagil- 
pua village to Borgen Bay and Cape Cilouccster; 
side trip to Mt langis, reaching summit (i7()0m) 
from the NE. ; New Guinea (Morobc, Dec): 1 
week in the Bulolo-Wau area wilh I). J. Hiir. — 
1966. 2 J weeks (Jan.) in the mid-Waua Valley (with 
J. between Garaina and the Papuan 
border (lat. 8'S); collecting mainly near Wakaia 
village (r/. T. Ci. HAKriiv). -New Britain (Mar.- 
Junc), western part, for the greater part attached 



Flora Malesiana 

[ser. I, vol. 8^ 

to a survey from the Dept of Forests; a helicopter 
used to reach several localities. From base camp 
at Malcni^elo I. on S. coast, visiting: Johanna 
River near Whiteman Range; from Arawc base 
camp: lower Puiie River, upper Pulie River near 
Benim and Arawe Plantation (with E. E. Henty), 
river E of Sauren (sago and Casitarina swamps in 
part), Marklo I. (in Arawe Is.) andSubdidi village 

Frodin (behi.idj, in front from lefi to right: 
Nava Kwapena, Sister Marietta Garmer, 
David Ollerenshavv', Andrew Yanieb. U.P.N.G. 
Plant Systematics class, June 1972, Tapini Hotel, 
Central District. 

(in limestone country); from base camp at Borgen 
Bay: Aisega village; from base camp at Eleanora 
Bay, N. coast: Mt Penck and vicinity (up to the 
summit, 600 m ; a few distinctly montane species on 
windswept, foggy peak) as well as to a forest on 
limestone country some 20-25 km S of Cape Ruge. 
Proceeding (May 19) by air to Cape Gloucester 
after brief rest at Talasea; by boat and foot to 
Sag Sag, whence again climbing Mt Tangis as well 
as Mt Talawe (1900 m); return to Cape Gloucester 
(June 1-2) and thence to Rabaul (2-4), visiting 
Keravat briefly; return by air to Lae (4). — New 
Guinea (Madang) : mid-June U- weeks in lowland 
forest and grassland around Aiome (near former 
station of the Neu-Guinea Kompagnie, Arumene 
on the Ramu R.). Southern Highlands District 
(July-Sept.); Doma Peaks Expedition (5 weeks): 
proceeding via Mt Hagen to Tari by air; by land- 
over to Tigibi and thence on foot to base camp 


Ibiwara, joining Vink and Kalkman (see there, 
also for literature). Frodin spent 3 weeks at Lei 
bivouac (2800 m) and 2 weeks at Habono bivouac 
(2200 m). Sepik Distr. (Sept.). By air from Tari 
(with Vink and Kalkman) to Telefo(l)min, 
proceeding on foot to the limestone Hindenburg- 
r^ap Range; camp for 1 week at c. 2400-2450 m; 
returning by air to Lae via Wewak and Madang. 
In Nov. day-trip with C. E. Ridsdale to Markham 
Point near Lae {Nepenthes). — 1967. Malay 
Peninsula (Apr.): a few collections around F.R.I. 
Kepong and day-trip to new road to G. Ulu Kali, 
E of Kuala Lumpur. — 1971. Papua. First half of 
the year in the vicinity of Port Moresby and on 
the Sogeri Plateau, partly with Mrs A. N. Millar 
as well as with students; field trip (June 19-25) to 
Kerau, about 12 km due NE. of Tapini, 2200 m 
alt. (area generally N of the region examined by 
the 1st Archbold Exp. 1933-34, i.e. the strip from 
Mt Tafa through Ononge to Mt Albert Edward; 
field trip to Bulolo-Wau area (4 days, end of Aug.); 
to Musgrave R., near lawarere, and along road 
between there and Koitaki. — 1972. New Caledonia 
(1st week Jan.); first half of year in vicinity of 
Port Moresby, Motupore I. in Bootless Inlet 
( Field Station of Univ. P. & N.G.) ; Owen's Corner 
with R. PuLLEN (June, day trip); participation on 
Mt Suckling Expedition (see P. F. Stevens, J. F. 
Veldkamp, R. Pullen, etc.) (June 20-July 4): 
from Biniguni ascended to Maiyu I and Maiyu II 
camps and to the summit region of Mt Sewie (2 
days on top, 3000 m), return by the same route. 
Trobriand Is.: Kiriwina (Oct. 26-Nov. 9), base 
camp at Kaibola in the north. — 1973. Bisinarck 
Archipelago (Jan. 13-Feb. 8): Gazelle Peninsula 
(Kerevat, Vudal, Bainings Road) in New Britain 
(Jan. 14-17); New Ireland {idin. 18-Feb. 2): Danfu 
and vicinity, Namatanai, ascending Lelet Plateau 
(1000 m) from E. coast near Dalum and staying 
4 days at Lemkamin Aid Post, Kavieng and visit 
to the islands of Nusa and Nusalik offshore with 
B. N. ToREU; Manus (Feb. 2-8): Lorengau, by 
launch to Derimbat (N. coast) and inland to 
Kari (4(K)-500 m), and return to Derimbat and 

Collections. Herb. Lae: some 1700 nos in 
the NGF series (incl. 26201-27000, 28101-28500, 
32001-32200, 28501-28549; others jointly with E. 
E. Henty, C. E. Ridsdale, J. Buderus), of which 
c. 800 collected in New Britain. The few collections 
from the Malay Peninsula in Herb. Kep. (in FRI 

Special attention paid to Araliaceae and 

Later collections in Port Moresby, with UPNG 
nos, ranging from 500-1000, 2001-2200, 4001- 
4190, Dupl. distributed from Port Moresby 
to Lae, Leyden, Canberra, Kew and B.P. Bish. 

Froggatt, John Lewis 

Add: (1891, Sydney, Australia; x) was educated 
at Sydney University (B.Sc. 1913); temporary 
officer under Dept of Agriculture, N.S.W., 1913- 
18; appointed Entomologist, Queensland Dept of 
Agriculture and Stock, Brisbane, in October 1920, 
resigning August 1929, to take up a position as 



Cyclopaedia of collectors — Supplement II 


entomologist to the Mandated Territory, New 

Biographical data. In Bibliogr. Austr. 
Entom. 1775-1930, 1932. 

*Frohne, Gertrude 

collected plants in Leyre (Philippines) in August 
J957; numbered in the PNH series, Herb. Manila. 

Fruhstorfer, Hans 

(d. 1922, Munich, Germany). 

Collections. Add: Bryophytes from Celebes 
{1895), also in Herb. Eclinb. 

Biographical data. Add: The Entomolo- 
gist 55, 1922, p. 144; in Musgrave, Bibliogr. Austr. 
Entom. 1775-1930, 1932. 

*Fuchs, Hans Peter 

(1928, Basle, Switzerland; x) studied botany and 
geography at the University of Basle (1948-58), 
taking his Ph.D. (1958) on the taxonomy of the 
genus Isoetes. After a year in the National Her- 
barium of the Smithsonian Institution at Washing- 
ton, he joined the Bataafsche Internationale 
Petroleum Maatschappij, The Hague, and was 
transferred in 1961 to the Kon. Shell Exploratie 
en Produktie Laboratorium at Rijswijk as Research 
Palynologist. He participated in the Symposium 
on Ecological Research in Humid Tropics Vege- 
tation at Kuching, Sarawak, in July 1963,* and 
extended his stay to NW. Borneo (see below). 
Early 1967 he spent 4 months on the Pacific coast 
of Columbia, S. America. At present living at 
Trin-Dorf, near Chur, Switzerland. 

Rhododendron fuchsii H. Sleum., and Pandanus 
fuchsii H. St. John, etc., are named after him. 

Collecting localities. Malaysia. 1963.^ 
Sara>\ak: Mt Santubong (July 7), Semengoh F. R. 
and limestone hills at Bau (10). Brunei: Seria- 
Badas peat-swamp forest and Badas F. R. (12). 
Sabah: Mt Kinabalu, W. route (14-19) at Kam- 
baranga Relay Station and Paka Cave, Mt Kina- 
balu Low Peak; Penosok Plateau (21-25), partly 
with H. Sleumer, W. Meijer, and Mrs Collen- 
ette, at Mesilau Cave, Tenompok, etc.; Andulau 
F. R. (31. with J. Muller); Seria to Badas F. R. 
with D. O. J. Diederix (Aug. 1). Brunei: Lumut 
Hills (2). Sarawak: Long Palau, B. Tinjar (7); 
Lobok Pasar, B. Baram(9-10); Marudi F. R.(IO); 
Bt Subis. S. Niah (13); Bt Lambir, Miri (16); Bt 
Pasu, Tubau Valley (20-21). Sabah: Mt Kinabalu, 
E. route, partly with Mrs Collenette: Tenompok 
(27), Kundasan-Mcsilau Cave (28), to Janet's Halt 
(29), to Sheila's Plateau (30), Shangri La Valley, 
N. side of the mountain (31). Coking's Valley (N. 
side) (Sept. 1-6), Penosok Plateau (8); Mesilau 
Cavc-Kundasan (8j; Sheila's Plateau to Mesilau 
Cave (4-7, Mrs Collenette alone, with Fuchs 

Collections. A'o.y 21000-21682; first in Herb. 
Shell Lab. Rijswijk, now in private harbarium; 
dupl. distributed through Lcydcn to Leyden, Kcw, 
Sarawak, Arn. Arbor., Geneva, Canberra, Bogor, 
Wash., Copenh., .Slockh., Paris. 

Liter ature. (1) I-Or the field excursions of 
the UNESCCJ Symposium, sec Kuching Sym- 
posium 1963, 1965. p. 274-287 (trips 10 17 July 

*Fujioka, M. 

at the time Head of the Department of Agricul- 
ture, Tokyo Imp. Univ., collected in the Malav 
Peninsula, e.g. at Port Swettenham in yp-'V. 
Specimens in Herb. Fac. Agr. Tokyo Univ. 

*Fukuoka, Nobuyuki 

a member of the Kyoto Univer.sity Biological 
expedition to Ihaiiand and Malaysia, 1967. 

Itinerary. See sub K. Iwaisuki. 

Collections. Isl set in Herb. Kyoto, 2nd in 
For. Herb. Bangkok. 

Furtado, Cactano Xavier Dos Remedios 

Add: Pensioner, re-engaged Botanist, ceased to 
work for the Botanic Gardens, Singapore, in 
August 1960. He has continued to work, however, 
at the Botanic Gardens on an assignment for the 
Sarawak Government ; later supervising Colombo- 
plan Fellows. 

CoLLici iNG localities. Add: 1937. G. 
Baling. Kedaii (June 4). 

•Gabot, v., see Philippine National Herbarium. 

*Gadoh anak Umbai 

collected in the KL series (see there) of the 
PhytochemicalSurvey of the Federation of Malaya. 

*GaIau, see Forest Dcpt Sarawak. 

Galocngi, Karo-Karo Sinoelingga 
(d. Dec. 17. 1956). 

*(ialorc, Michael 
(1935, Wcdau. S\.. New CJuinca; x), Technical 




Flora Malesiana 

[ser. I, vol. 81 

Assistant with the Division of Botany at Lae, 
New Guinea; onwards of about 1972 Keeper of 
the Herbarium. 

Collecting localities. E. New Guinea, 
at least onwards of 1962. — 1967. With Ridsdale, 
SOEGENG, etc. on the Border Expedition in W. 
Papua. In the same year? islands in Milne Bay. 

Collections. In the NFG series, Herb. Lae. 
Also dupl. in Herb. Edinb., partly collected with 
P. J. B. Woods and Cruttwell. 

Gandrup, Jens Johannes Sorensen 

Biographical data. Add: Naturh. Tid. 8, 
1944, p. 46-47, portr. 

Gaudichaud-Beaupre, Charles 

Literature. (1) Add: For botanical drawings 
of the voyage of 'L'Uranie et La Physicieime' see 
Cl. Nissen, 'Die botanische Buchillustration' 
1951,vol. 2, p. 65. 

(3) Add: For botanical drawings of the voyage 
of 'La Bonite' see Cl. Nissen, 'Die botanische 
Buchillustration' 1951, vol. 2, p. 65-66. 

(5) Add: B. C. Stone: 'Gaudichaud's species of 
Pandanus in the atlas of the botany of the voyage 
of 'La Bonite' ' (Mai. Sc. 5, 1970, p. 14-19, pi. 22). 

Biographical data. Add: Adansonia ser. 
2, 11, 1971, p. 405-424 + portr. (new data on life 
and work). 

♦Gaynar, E. 

collected in the CF series (see Conservator of 
Forests in Cycl. FI. Mai. I, 1, 1950) in Selangor in 
1922. Probably no regular collector. 

*Geer, Louis de 

Collections. Herb. Stockholm: some 50 nos 
from Poentjak, W. Java (May 1965). 

*Geh Siew Yin, Miss 

collected orchids and other plants together with 
Miss Lee Wax Chin, and with Samsuri. 

Collecting localities. Malaysia. 1970. 
Pulau Langkawi (Dec. 3-6); Kaki Bukit, Perils (6); 
Maxwell Hill, Perak (7-10); Penang Hill, Penang 
ill).— 1972. P. Tioman, Bt Penak (Sept. 17-18). 

Gibbs, Lilian Suzette 

Literature. (2) Add: For the reconstruction 
of her route in the Arfak Mountains see W. Vink 
in Nova Guinea, Bot. 22, 1965, p. 485-486. 

*Gibot, A., see Forest Dept North Borneo. 

Gilliard, Ernest Thomas 

(d. Jan. 26, 1965, New York, U.S.A., after a 

Author of a paper on New Guinea (Nat. Geogr. 
Magaz. 1951, p. 669 seq.). 

Biographical data. Auk 83, 1966, p. 416- 
422, w. portr. & bibliogr. 

GilHIand, Hamish Boyd 

(d. June 1965, aged 54, ? Pietermaritzburg, S. 

In 1965 he took up a professorship at Pieter- 
maritzburg, S. Africa. 

Author of botanical papers.^ 


Collecting localities. Add: 1956. Sara- 
wak: Bako Nat. Park (May 16-22).— 1960. NE. 
New Guinea:* Chimbu and Daulo Pass. Borneo? 

Literature. (1) H. B. Gilliland: 'Plant 
Communities on Singapore Island' (Gard. Bull. 
Sing. 17, 1958, p. 82-90); 'Common Malayan 
Plants' (Singapore 1958); Flora of Malaya, vol. 3 
(grasses); and several other papers, partly on 
grasses, and Borneo plants. 

(2) Cf. Fl. Mai. Bull, no 18, 1963, p. 990. 

Biographical data. Nature 207, 1965, p. 
808; Gard. Bull. Sing. 22, 1967, p. 107-112, incl. 
portr. a. bibliogr. 

*Gillis, J. R., see Forestry Bureau, Manila. 

*GilIison, Andrew N. 

was transferred from the Department of 
Agriculture to the Department of Forests, 1964. 
Ecologist at Lae, New Guinea. 

His interests are mainly ecological, but he is 
currently (1967) teaching botany at the Forestry 
School, Bulolo. He is working on Melanesian 

Collecting localities. Pa/7Ma; Port Mores- 
by region, Louisiade Archip., Trobriand Is., 
d' Entrecasteaux Is., and Conflict group. — 1966. 
Accompanied part of the 'Leyden-Lae Doma 
Peaks Expedition'^ to the Southern Highland 
Distr. (see also Kalkman and Vink); later 
replaced by D. G. Frodin. He was not on the 
Andutakim. — 11967. Islands of Milne Bay Distr. 
with M. Galore and P. Eddowes. 

Collections. In the NGF series, Lae; dupl. 
distributed to other herbaria, incl. Herb. Edinb. 
His special interest is Diospyros. 

Literature. (1) A. N. Gillison: 'Plant 
succession in an irregularly fired grassland area — 
Doma Peak region, Papua' (J. Ecol. 57, 1969, p, 
415-428, 4 fig., 2 photogr.); 'Structure and 
Floristics of a montane grassland/forest transition, 
Doma Peaks region, Papua' (Blumea 18, 1970, p. 
71-86, photogr. 1-8, fig. 6). 

Gjellerup, Knud 

Literature. (7) Add: For annotations on 
the Arfak route (Apr. 19-May 16, 1912), see W. 
Vink in Nova Guinea, Bot. 22, 1965, p. 485, fig. 5. 

*Glasse Lindenbaum, Shirley Helen 

(1933, Melbourne, Australia; x), B.A. Mel- 
bourne, and 2 years post graduate work in 
anthropology at Sydney University. She made an 
anthropological field trip to the Eastern Highlands 
of Australian New Guinea, together with Dr R. 
M. Glasse in 1962-63, to study the social structure 
and ecology of the Fore, with special reference to 
'kuru', a neurological disorder suff"ered by the 
people. About 1966 she remarried, her new name 
being Lindenbaum, and lives in New York. 

Collecting localities. 1962-63. Austr. 
New Guinea: South Fore Census Division of the 
Okapa Sub-District, Eastern Highlands. 

Collections. NGF nos 15626-15672 in 
Herb. Lae; 321 items were identified in the field 
by Dr Womersley.^ 

Literature. (1) The full list is contained in a 
mimeographed paper entitled 'A Note on Fore 
Medicine and Sorcery'. 



Cyclopaedia of collectors — Supplement II 


♦Godlee, T. 

Forest Officer who collected some specimens on 
the Upper Clay River off Kerama River, Sepik, 
April J960; in Herb. Lae, Leyden, etc. 

Goebel, K. von 

Biographical data. Add: 'Ein deutsches 
Forschersleben in Briefen 1870-1932'. Heraus- 
gegeben von E. Bergdolt, Berlin 1940, 273 pp., 

Goering (or Goring), Philip Friedrich Wilhelm 

Collections. Add: Japanese orchids in 
Herb. v. Rosier, were described by G. Reichen- 
bach (Bot. Zeit. 3, 1845, p. 333-335). 

•Gonzales, Jose Salvador 

The orchid Dendrobium gonzalesii Quis. has 
been named after its fixst collectors, Mr & Mrs 


Graeffe, Eduard 

Add: Author of 'Samoa oder die Schifferinseln' 
(Hamb. Mus. Godeffroy 1873). 

Grassl, Carl Otto 

(1908, Menominee, Mich., U.S.A.; x). Add: 
A.B. and A.M. from the University of Michigan, 
1932 and 1933. 

Collecting localities. USDA Sugar Cane 
Expedition, 1957.^ Austr. New Guinea (Mar. 15- 
end of Apr.): between Laloki and Brown Rivers 
near Port Moresby, Warida R., Sepik R., Ramu 
R., Erap R., Markham R., Tauri R., Kaisenik 
near Wau, Snake R., Bulolo-Lae Rd, Gogol R., 
Madang, near Wewak, Bainyik etc. near Maprik 
(Sepik drainage), Sepik R., Nondugl, Purari 
drainage, Baiyer R., Kikori drainage; Dutch New 
Guinea (May): Manokwari, Wosi R., Anadi R., 
Bami R., Sorong, Ransiki, Anggi Lakes, Biak, 
Wissel Lakes, Merauke, HoUandia; Austr. New 
Guinea (fortnight in early June): Ramu R., 
Bainyik near Maprik (Sepik drainage), Sepik R.; 
New Britain: Keravat; Solomon Is (8 days, mid- 
June): Guadalcanal, Florida I., Buka I.; Austr. 
New Guinea (10 days): New Britain, near Keravat; 
Epo, near Mekeo; Popondetta. 

Collections.' Andropogoneae from New 
Guinea, specimens made from flowering living 
plants, will be deposited in the specialized her- 
barium at Canal Point, Florida. Earlier collections 
from Canada, and Michigan, in Herb. Un. 
Michigan; Andropogoneae from Centr. America 
and E. Africa in Nat. Herb. Wash, and in Florida. 

Large living collections of Saccharum from New 
Guinea in Canal Point, Florida. 

Literature. (1) J. N. Warner &. Carl O. 
Grvssl: 'The 1957 Sugar Cane Expedition to 
Melanesia' (Hawaiian Plant. Rec. 55, 1958, p. 
209-236, f. 1-14, App. A-G, incl. map). 

Cray, Edward Charles Gordon 

I^tcr Senior Forest Officer, stationed at Port 
Moresby, F'apua. He further collected on Brown 
River {1955), Mt Lawcs near Port Moresby (Apr. 
1956), and with E. J. H. Corner at Waitapc (Aug. 

Grcvenstuk, Anionic 
(d. 1973, Baambruggc, N.H.. Netherlands). 


Griffith, William 
Collections. Add: Aho ^u'pX.'m Herb. Edinb. 

Groenhart, Pieter 

(d. 1966, Leyden, Netherlands). 

Biographical data. F1. Mai. Bull, no 21, 
1966, p. 1371, 1377; Persoonia 4, 1966, p. 69-71, 
w. portr. a. bibliogr. 

Groot (van Embden), Cornelis de 

Biographical data. Add: Bijdr. Taal-, 
Land- en Volkenk. 14, 1958, facing p. 129. 

*Grubb, Peter John 

(1936, Ilford, England; x) was educated at 
Magdalene College, Cambridge University, B.A 
(1st Class Hons) 1957, Ph.D. 1961 . 

University Demonstrator (1961-64) and Lectur- 
er (1964- ). He made ecological expeditions in 
the Balkans, S. America, Australia and Solomon 
Is. (1963), and to Japan, Australia, New Zealand, 
and U.S.A. (1970-71). 

Collecting localities. 1970. Malaya: G. 
Ulu Kali, Cameron Highlands (with T. C. Whit- 
more); Jalang Mersing, Kluang, Johore, and G. 
Belumut (Sept. 20-25). E. New Guinea (Oct.-Dec. 
& Apr.-July 1971); Mt Kerigomna and Fatima 
Basin (Marafunga sawmill) in Eastern Highlands. 

Collections. Herb. Cambr. In Malaya 
collected for mineral analysis and anatomical 
studies. New Guinea collections numbered P. J. 
Grubb & P. J. Edwards, c. 380 fertile numbers 
(distributed to BM, Lae, Canb., Leyden) and c. 
20(X)-23(K) sterile and unicates kept in Cambr. 

Guillou, Elie Jean Francois le 

Biographical data. Add: in Bibl. Austr. 
Entom. 1775-1930 (R. Zool. Soc. N.S.W. 1932, 
p. 135). 

Gutierrez, Hermes G. 

Add: (1933, Laoag City, Luzon, P.I.; x) joined 
the Philippine National Herbarium (Bot. Dept of 
the National Museum) in 1954, and was later 
appointed Museum Researcher in Botany and 
subsequently Chief of the Botany Division. He 
was educated in Botany at the University of the 
Philippines (B.S.), the Arnold Arboretum (Harvard 
University) (1963-64), and the University of 
Santo Tomas (M.S., and working for Ph.D. in 
1968). Lately especially interested in Diptero- 

Collecting localities. Add: 1957. With 
G. E. Edano. — 1960. Hundred Islands, Pangasin- 
an (Algae); W. Java: Udjong Kulon, Mt Pang- 
garango,Tjibodas(with KosTERMANsand UNESCO 
Training Group). — 1961. Batan Is., Batanes and 
Babuyanes (with Quisumbing, see there, and R. 
DEL RosARio); Sierra Madrc Mts, Isabcia, Luzon 
(Apr. 17 - May 28).— 1962. Leytc: Mt Lubi, 
Dagami & Balinsasayao F. R., Abuyog; Malaya: 
Ulu Kelantan Limestone Hill (with H. Keng and 
UNESCO Training Group). — 1%3. Palawan: 
Lipuun Point, Quezon, with R. Espiritu. — 1965. 
Ethnobotanical collecting in W. Mindoro (Mar. 
8-20); trip in Basilan with E. J. Reynoso and Z. 
C. Fraga: with Dr Aaron Sharp & Dr Z. 
Iwatsuki on Mt Canlaon, E. Negros (May 25-29), 



Flora Malesiana 

[ser. T, vol. 8^ 

and in SE. Samar (May 30-June 28) with E. J. 
Reynoso; Mt Pulog, Benguet, Luzon. — 1966. 
With Reynoso in Mt Guiting-Guiting, Sihiivan I., 
Romhlon.— 1969. S. Samar (May 5-Junc 3)': Mt 
Concord, Hinabangan and Mt Apoy, Llorenle, 
with R. A. EspiRiTU and E. J. Reynoso. Mindanao, 
Davao: Mt Apo (Nov. I9-Dec. 20), with E. J. 
Reynoso and F. R. de la Cruz, K. Seto, and 



Others of the Osaka Museum of Natural History. — 
1970. S. Samar: Mt Sohoton (Mar.-Apr.) with 
D. A. Madulid, R. a. Espiritu, E. J. Reynoso 
and F. de la Cruz.^— 1973. TV. Luzon (April). 

Collections. Add: Appr. 1500 field numbers 
in 1968; dupl. distributed to Leyden, Kew, Am. 
Arbor., Sing., U.S. Nat. Mus., Bogor, etc. The 
collection from Mt Apo (1969) shared equally at 
the Osaka Mus. and Nat. Mus. Manila. 

Literature. (1)C/.F1. Mai. Bull. 770 24, 1969, 
p. 1786-1787. 

(2) Cf. ibid, no 25, 1970, p. 1892. 

Gwynne-Vaughan, David Thomas 

Collections. Add: The Curator of the 
Aberdeen University Herbarium recently (1973) 
came on a field note-book given to Prof. W. G. 
Craib by Dr O. Staff, at one time curator of the 
Kew Herbarium. It has been returned to Kew; it 
lists plants collected during the 1899 Skeat 
Expedition in the Malay Peninsula, nos 200-639 
with informative notes. 

*Haas, A. de 

collected Digitaria timorensis (Kunth) Balansa 


in 's Lands-fokkerij Padang Mangatas near 
Pajakombo (Sumatra West Coast) in Mar. 1922. 
In Herb. Bog. > 5. 

Hachisuka, Masauji 

Collections. Add: Evidently at least some 
phanerogams too. 

Literature. Add: (2) Cf. M. Hachisuka: 
'The Birds of the Phillippine Islands' (London 
1931). List of collected ferns (27) and phanerogams 
(19) on p. 71-73. 

Haenke, Thaddaus Peregrinus Xaverius 

Add: Among his MSS (in Madrid, fasc. 15-16) 
are short botanical notes, partly from the Philip- 
pines, which however, may partly originate from 
L. Nee.6 

Collections. Add: Part of the Lambert 
Herbarium collection (Mexico and Philippines), 
sold 1842, is now at Kew (cf. Taxon 19, 1970, 524). 

Literature. (1) Add: For itinerary in Alaska 
cf. Bot. Not. 1940, p. 297; for itinerary in Luzon 
cf. J. KuHNEL I.e. below sub Biogr. Data, p. 79. 

(2) Add: R. E. Holttum: 'A commentary on 
some type specimens of ferns in the Herbarium of 
K. B. Presl (Nov. Bot. Inst. Bot. Univ. Carol. Prag. 
1968, p. 3-57) (see also review by C. V. Morton 
in Amer. Fern J. 60, 1970, p. 119-123). 

(6) R. Gicklhorn: 'Unbekannte botanische und 
zoologische Forschungsergebnisse von Thaddaus 
Haenke' (Verh. Zool.-Bot. Ges. Wien 103/104, 
1964, p. 215); 'Neue Dokumente zu Beginn der 
Forschungsreisen von Thaddaus Haenke' (Phyton 
14, 1972, p. 295-308). 

Biographical data. Add: in Hemsley & 
Hooker, Biol. Centr.-Amer. 4, 1887, App. p. 119- 
120; Occ. Pap. Calif. Acad. Sc. no 20, 1943, p. 13; 
J. Kuhnel: Thaddeus Haenke, Leben und Wirken 
eines Forschers, Miinchen 1960, Veroff. Collegium 
Carolinum Bd G, incl. portr., bibliogr., maps, 
letters, etc.; Verh. Zool.-Bot. Ges. Wien 103/104, 
1964, p. IX'i-Tn^lfvg. 

Haenseler, Felix 

Add: (1767, Durrach, Bavaria, Germany; 1841, 
Malaga, Spain) was certainly not in Malacca 
himself. He was a correspondent of Mertens, 
Agardh, Webb, etc. He collected extensively in 
Spain and exchanged materials with botanists out- 
side Spain. The plants cited from Malacca must 
be cited either ex Herb. Haenseler, or more 
probably from Malaga! 

Biographical data. Bot. Zeit. 4, 1846, p. 

Hagen, B. 

Add : He collected in Sumatra at least up to 189L 

Hagerup, Olaf 

(d. 1961, Copenhagen, Denmark). He retired in 

Biographical data. Taxon 10, 1961, p. 25- 
26, 177-178, 201-204, portr., bibliogr.; Dansk Bot. 
Ark. 21, 1963, p. 63, 133-135 (bibliogr.). 

♦Hainsworth, Cecily Joan 

(1936, Perth, W. Australia; x) got linguistic 
training at the Summer Institute of Linguistics, 



Cyclopaedia of collectors — Supplement II 


Australia; one year linguistic and anthropological 
studies at the University of Western Australia. 
With a co-worker, Kathleen Johnson, Linguists 
with the Summer Institute of Linguistics, Ukar- 
umpa. Territory of New Guinea, for the Narak 
language, compiling a Dictionary. 

Collecting localities. In the 1960s. At 
Tabibuga in the Jimi River Valley of the Western 
Highlands of East Sew Guinea. 

Collections. Some 80 specimens in Herb. 
Lae. In 1968 continued collecting was planned. 
Plant collecting has been done to identify plants 
for the dictionary. 

* Hamad 

His name occurs on labels as Curtis' collector, 
e.g. in Langkawi Is. (July 1892). 

•Hamilton, Miss L. 

now Mrs Read at Lae; in 1956 stationed at 
Okapa, Eastern Highlands, New Guinea, as a 
Nutritionist with Public Health Department. 

Collections. About 230 nos (no good 
specimens); in Herb. Lae, and sent to various 
institutions for identification by specialists, to 
Leyden in 1958. Mainly useful plants from Papua 
gardens; partly with C nos, partly in the NGF 

♦Hanevelt, Miss A. D. 

collected plants in the Cameroon Highlands, 
Malaya, at 4000 ft, in 1937: Herb. Sing. 

♦Hardie, J. 

Collections. Java plants in Herb. Edinb. (on 
permanent loan from Glasgow Univ. Herb, since 

*Harland, William Aurelius 

(?. Scarborough, England; Sept. 12, 1858, 
Hongkong), a colonel surgeon, M.D. Edinburgh 
1845, first in China, and in Hongkong since 1848. 

Harlaiidia Hance and some other plants have 
been named in his honour. 

Collections. Herb. Kew: plants from China 
and Hongkong. In the Brit. Museum specimens 
also, including Cyperus conipactus from Singapore, 
collected in Maio (May) 1857. 

Biographical data. Biogr. Index Britten 
& Boulger, 2nd ed. by Rendle, 1931; Backer, 
Verkl. Woordenb. 1936. 

* Harrison, R. 

In the 1950s Cadet Agricultural Officer stationed 
at Nondugl, Territory of New Guinea. He made a 
small collection of weeds and pasture plants; in 
Herb. Lae. 

Hart, Alexander John 

CoLi kctions. Add: Dupl. in B. P. Bish. Mus, 
Herb. Honolulu. 

•Hartley, Thomas Gordon 

(1931, Beaumont, Icxas, U.S.A.; x) graduated 
in bf)tany (B.S. 1955, Wisconsin State University; 
M.S. J957. F'h.D. 1962. University of Iowa). 
From 1961-65 leader of C.S.I. R.O. Phytochcmical 
Survey of New Guinea; 1965-71 Associated 
Curator of the Arnold Arboretum, Harvard 


University, Cambridge, U.S.A.; 1971- present 
Senior Research Scientist, C.S.I. R.O. , Division 
of Land Research, Herbarium Australiense, 
Canberra, Australia. 

He revised the genera FUndersia and Zanth- 

Collecting localities. 1961. Territory of 
New Guinea: Morobe District (from Dec. 16), 


vicinity of Lae, in Markham Valley, Gurukor, 
Butibum R., Lae Botanic Gardens. — 1962. Same 
localities near Lae and Oomsis Creek, Lake Bed 
below Red Hill, beach near Malahang; S and N of 
Mumeng, Lake Wanum, above Busu R.; Papua: 
between Brown and Laloki R. (Feb. 20); Terr, of 
New Guinea: vicinity of Lae, a.o. at Yalu, Bulolo, 
Edie Creek Rd, Zcnag, Bewapi Creek, Burep R., 
Leron R., Bunga R., N and S of Garaina, Sopa 
near Garaina Singaua; Papua: Laloki R. (Sept. 5), 
2 miles E of Rouna (6), below Rouna Falls (7), 
Kakoda Rd ( 1 0), E of Rouna (II), Laloki R. E of 
Sogeri (11-12), E of Rouna (13), Brown R. (15). 
Kakoda Rd (17), Laloki R. 15 miles NE of Port 
Moresby (21), IJ miles E of Subitana (24), 
Musgrave R. (25); Terr, of New Guinea: same 
localities in vicinity of Lae and besides at Crooked 
Creek SW of Bulolo, Patep R., Lae-Bulolo Rd.— 
1963. Beach area NE of Lae, between Busu and 
Butibum Rivers, Mt Salawakct ( ? Saruwaged) 
(Jan. 19-26), localities near Lae, track between 
Tymne and Wagau (Mar. 12 18, 21). Kuali Creek 
(26-Apr. 8). S of Wau. Bulolo River Gorge. 
Kaisenik, Bulldog Track above Edie Creek, Mt 
Kaindi SW of Wau (May 8-9), vicinity of Lae; 
Kainantu Subdistr.. Eastern Highlands (July 3- 
Aug. J I); Kratkc Range S of Aiyra, Eastern 
Highlands (Aug. 12); near Lae, Morobe District; 



Flora Malesiana 

[ser. I, vol. 8^ 

Kainantu Subdistr., Eastern Highlands (Nov. 26- 
28); Morobe Distr., e.g. Mt Shungol (Dec. 10-17) 
5 miles SW of Wagau, partly in his series col- 
lected by C. D. Sayers (Dec. 16-18).— 1964. 
Morobe Distr. (Jan. 7, 8, 10, by C. D. Sayers) 
till Feb. 8 ; Papua: Mt Dickson, Goilala Subdistr. 
(Feb. 9-14); Terr, of New Guinea: near Lae (some 
numbers by Dzamo); Okapa Subdistr., Eastern 
Highlands (Sept. 24-30), Goroka Subdistr., 
Eastern Highlands (Oct. 9-14).— 1972. Morobe 
Distr.; Eastern Highlands District. 

Collections. 3750 TGH nos (9600-13305, 
13655-13699), partly collected by C. D. Sayers 
and a few by Dzamo; 1st set in Lae; dupl. in 
private herb., Canberra, Harv. Bot. Mus., Leyden, 
Gray Herb., Nat. Herb. Wash., Brisbane, and Kew. 

Other collections were made in the U.S.A., 
Puerto Rico, New South Wales & Australian 
Capital Territory. 

*Hasan Pukol 

collected on behalf of Forest Dept Brunei. 

*Hashim, A., see Forest Dept North Borneo. 

Hasselt, Johan Coenraad van 

Literature. (3) Under J. G. S. van Breda 
add: For particulars on the drawings see Cl. 
NissEN, 'Die botanische Buchillustration' 1951, 
vol. 2, p. 22 sub Breda. 

Hatusima, Sumihiko 

Add: made a joint expedition from Kagoshima 
University Japan and the National Science Museum 
at Manila, P.L in 1964.^ He was accompanied by 
M. Sato, a botany student of Kagoshima Uni- 
versity, and by the Philippine assistants F. Mer- 
CADo and A. Zabala. 

Collecting localities. 1964. N. Philip- 
pines: after a visit to Manila, collecting 8 days in 
Batan L^ and 4 in Camiguin (Nov.). In Batan I. 
collecting on Mt Iraya, along the coast of Song- 
song Bay, on the rocky cliff between fiasco and 
Mahatao, and in the valley behind Mahatao. 

Collections. 1054 nos Philippines, totalling 
2122 specimens. 

Literature. (1) S. Hatusima: *An Enumer- 
ation of the Plants of Batan Island, N. Philippines' 
(Mem. Fac. Agr. Kagoshima Univ. 5, 1966, p. 
13-64, 5 pi., 1 fig., map). 

H. Inoue: 'A small collection of liverworts from 
Isl. Batan, the Philippines' (Hikobia 4, 1965, p. 

T. Shin: 'A small Collection of Mosses from 
Isl. Batan, the Philippines' (Hikobia 4, 1965, p. 
263-271, 1 fig.). 

Biographical data. Portr. in Hatusima, 
Flora Ryukyus 1971, p. 69. 

Haviland, George Darby 

Collections. Add: Dupl. from A^. Borneo 
(1891-95) in Herb. Edinb. 

♦Heaslett, Dr E. A. 

In 1970 in the Geography Dept Univ. of 

Collecting localities. 1970. Malaya. He 
led the Herbarium staff of the Singapore Botanic 
Gardens on day trips to G. Panti (July 5); to the 


rivers Bantan (May 31), Juasseh (June 28, Sept. 6), 
and Panjang (July 19) in the Labis Distr., Johore; 
to Jalang Mersing, Kluang, Johore (Oct. 4). 

Collections. Herb. Sing. 

Literature. (1) Author of 'Gunong Chabang 
Tiga: a note on a small area of montane Ericace- 
ous forest in Johore' (Mai. Nat. J. 23, 1970, p. 
149-154, 1 fig.). 

*Hee, P., see Forest Dept North Borneo. 

*HeeI, Willem Albert van 

(1928, Leiden, Z.H., Netherlands; x), plant 
morphologist who attended the University at 
Leyden (Ph.D. 1966). In 1956 appointed at the 
Rijksherbarium. He made a study tour to Java, 
for collecting suitable material for his research. 

Author of morphological papers. 

Collecting localities. 1969. Java (Apr.- 
Aug.), mainly in Hort. Bog. 

Collections. Mostly pickled, but also some 
dried material in Herb. Leyden. 

Heide, Frits Ferdinand Rudolf 

(d. 1957). 

Biographical data. Add: DanskBot. Ark. 
21, 1963, p. 64, 145-146 (bibliogr.). 

*Heider, Karl Gustav 

(1935, Northampton, Mass., U.S.A.; x), anthro- 
pologist, educated at Harvard (B.A. 1956, M.A. 
1959) and Vienna (1957-58). Instructor in Anthro- 
pology, Harvard University 1965-66 (Ph.D. 1966).^ 
Assistant Professor at Brown University, Rhode 
Island, 1966 onwards. Attached to the Peabody 
Museum of Harvard Expedition as Cultural 
Anthropologist, making ethnographic studies in 
Irian (former Dutch New Guinea). 

Collecting localities. 1962. W. Irian: 
Grand Valley of the Balim, Central Highlands. 

Collections. 17 nos in Herb. Leyden, and in 
Herb. Harv. Univ. Cambr. 

Literature. (1) Author of an unpublished 
Ph.D. thesis: 'The Dugum Dani. A Papuan Culture 
in the West New Guinea Highlands' (1965). It 
would be revised as a book. 

Heinrich, Gerd 

Literature. (1) Add: 'Celebes, Selfsame Jagd 
durch seltsames Land' (Berlin 1943). 

♦Henderson, Douglas Mackay 

(1927, Blairgowrie, Scotland; x) attended 
Edinburgh University (1944-48); Scientific Officer 
in plant virology, Dept of Agriculture and Fisheries 
for Scotland, 1948-50; from 1951 on the staff of 
the Royal Botanic Garden, Edinburgh; in 1970 
Regius Keeper, succeeding Dr Fletcher. 

He made a world tour of Botanic Gardens and 
Herbaria in 1967. 

Author of an account of the bryophyte flora of 
Norfolk Island (Revue Bryologique et Lich6- 
nologique 37, 1970, p. 657-661, with H. T. 

Itinerary. 1967. Malaya: based in Kuala 
Lumpur (Aug. 13-20); Singapore (20-26); Java: 
Bogor (26-30), and Tjibodas. 

Collections. His tour was not a collecting 
trip, and the collections made (mostly rust fungi) 
are not numbered; in Herb. Edinb. 



Cyclopaedia of collectors — Supplement II 


Henrici, Henry Albert Baron 

(1782, Rome, Italy; 1838, Amsterdam, Nether- 
lands), officer in Austrian regiments from 1800- 
1805 and from January 1817 captain in the Dutch 
army; major in the D.E.I, from 1825.* 

Literature. (5) This is to replace partly 
incorrect or incomplete data in the Cycl. Fl. Mai. 
I, 1, 1950; cf. J. MacLean in Scient. Hist. 14, 
1972, p. 59, who found the data in Rijksarchief in 
The Hague. 

about mid- 1948 transferred to Buitenzorg at his 
own request with the possibility to continue his 
agricultural studies there. He passed his final 
examination at Wageningen in January 1955 and 
got an appointment with the Dutch Forest 
Administration in Limburg. 

Collecting localities. 79-^5-50. W.Java. 
Mostly from Buitenzorg (Bogor) and vicinity; G. 
Gedeh, G. Salak; between Buitenzorg and Serang; 
vicinity of Bandung. 

*Henty, E. E. (Ted) 

Animal Husbandry Officer, Department of 
Agriculture, E. New Guinea; in 1957 he joined the 
Division of Botany; in 1962 appointed Keeper of 
the Lae Herbarium. 

He is especially working on New Guinea grasses^ 

Collecting localities. Mandated Terri' 
tory of New Guinea. 1949-54. From Morobe, 
Eastern and Western Highlands Districts, mainly 
grasses. — 1959. Okapa, Kainantu subdistr.. Eastern 
Highlands Distr.— 1960, See sub R. F. Thorne.— 
1963. Mt Piora (Mar. 9-13), Kainantu subdistr., 
with S. Carlquist.= — 1964. Sagarai Valley, Milne 
Bay environs, Milne Bay Distr.; Western foothills, 
Finisterre Mts, Madang Distr., with D. Sayers. — ■ 
1965. Telefomin, Sepik Distr.; Rossel and other 
islands, Milne Bay Distr. — 1966. West New Britain 
(with D. Frodin); Aiome, Usino, Dump, 
Madang Distr.; Kui, Morobe Distr.; Lake Trist 
(6000 ft), Morobe Distr.; with J. S. Womersley in 
Ramu Valley.— 1967. W. Papua: foothills of the 
Star Mts with the Border Survey Party for a 
fortnight (see Ridsdale and Soegeng). — Probably 
at many more localities on many more dates. 

Collections. Herb. Lae: ±300 under his 
own numbers, mainly grasses and weeds (1949-54, 
and some NGF numbers), later in the NGF series. 
Dupl. distributed by Lae; also in Herb. Edinb. 

Literature. (1) Author of 'A Manual of the 
Grasses of New Guinea' (1969); 'Weeds of coffee 
in the Central Highlands' (Bot. Bull, no 4, Terr. 
Papua & N.G. 1970, p. 1-22, 21 fig.). 

(2) Cf. Fl. Mai. Bull, no 18, 1963, p. 989. 

Heringa, Jodocus 

Add to Cycl. Fl. Mai. I, 1, 1950 and Suppl. I.e. 
I, 5, 1958: (1844, Utrecht, Netherlands; 1907, 
Utrecht), military dispenser, educated at a 
military school at Utrecht. In 1865 or 1866 he was 
sent to the Dutch East Indies, being stationed at 
Padang in West Sumatra. His botanical interest 
focusscd mainly on grasses. During his term of 
office in Indonesia (till 1894) he was transferred 
many limes, but no collections from other places 
arc known to me. In 1896 he opened a chemist's 
shop at Utrecht. 

Collections. According to his son, P. K. 
Heringa, his father had a private herbarium 
which no longer exists. 

•Hermens, M. C. L. 

(1925, Ambij, L., Netherlands; x) studied 
tropical silviculture at Wageningen Agricultural 
College. His study was interrupted from 1947-50, 
when he was serving for his number in the army 
in the then Dutch East Indies. At first he was 
stationed in Celebes, subsequently at Batavia, and 


Collections. About 300 specimens, aiming 
at collecting at least 1 species, sometimes more, of 
each family occurring in Indonesia. Identified by 
Dr VAN Slooten, Endert, a.o. In 1951 presented 
to Herb. Wageningen. 

Herre, Albert William Christian Theodore 
(d. 1962, Santa Cruz, Cal., U.S.A.). 
Collections. Also dupl. in B. P. Bish. Mus. 

Herb. Honolulu. 

Biographical data. Mandrofio 16, 1961, p. 

102-103, 1 photogr.; Bryologist 65, 1962, p. 268- 

277, portr., bot. bibliogr. ; Circ. Div. Syst. Biol. 

Stanf. Mus. no 10, 1964, p. 1-20, portr. 

Hcurn, Willem Cornelis van 
(d. 1972, Wilp, GId., Netherlands). 

Hewitt, John 

(d. 1961, Grahamstown, S. Africa). 

BiociRAPHiCAL DATA. Add: S. Afr. J. Sc. 57, 
1961, p. 312. 

*Hfylij»crs, Petrus Cornelis 

( 193l,Gorinchem, Netherlands; x) was educated 
at Utreclit University, majoring in plant geography; 
Dr.'s thesis (1961, published 1963) on vegetation 
and soil of a white-sand savanna in Surinamc. 



Flora Malesiana 

[ser. I, vol. 8^ 

Since October 1961 with C.S.I.R.O., Div. of Land 
Research, as plant ecologist in the New Guinea 
section. He carried out natural resources recon- 
naissance surveys, and more specialized research 
into the vegetation of monsoonal Papua. ^ 

Collecting localities. Eastern half of 
New Guinea. Papua: vicinity of Port Moresby 
(Aug. 1963); Solomon Is.: Bougainville (July-Aug. 
1864); vicinity of Port Moresby (May-July 1965, 
Jan. 1966). Terr, of New Guinea: Sepik Distr. 
between Ambunti and Aitape (July-Sept. 1966). 
Papua: vicinity of Port Moresby (Apr., May, Aug. 
7967, June 1968). Terr, of New Guinea: Sepik Distr. 
(Aug. 1968). 

Collections. Herb. Canberra (CSIRO), in 
Aug. 1968 about 70 nos; numbers started with 
1001 , as he collected earlier in Suriname (1956-57). 

Literature. (1) Results of surveys published 
in the CSIRO Land Research Series. 

P. C. Heyligers: 'Observations on Themeda- 
Eucalyptus savannah in Papua' (Pacif. Sc. 20, 
1966, p. 477^89, 7 fig., 3 tab.); 'Analysis of the 
plant geography of the semi-deciduous scrub and 
forest and the Eucalypt savannah near Port 
Moresby' (Pacif. Sc. 26, 1972, p. 229-241, 2 fig., 
7 tab.). 

Heyne, Benjamin 

Add: He employed Indian draughtsmen during 
his years as Company's botanist on the Madras 
Establishment. He built up a large collection of 
flower paintings of which the India Office Library 
possesses, however, none. Many of his drawings 
passed to the second Lord Clive (Governor of 
Madras 1798-1803.)« 

Literature. (9) Cf M. Archer: 'Natural 
History Drawings in the India Office Library' 
(London 1962) p. 27, 28. 

*Higgings, Dr Joseph J. 

employed at the U.S. Plant Introduction 
Station, Glenn Dale, Maryland. He accompanied 
H. F. Winters (see there) on an expedition to New 
Guinea in 7970. 

*HiIl, Dr Ronald David 

(1935, Wellington, New Zealand; x) graduated 
M.A. from Victoria University of Wellington 
(1961); Drs. University of Singapore (1965). On 
the staff of the Geography Department, University 
of Singapore. 

Collecting localities. 7969. Singapore. 
Bukit Timah Nature Reserve, in Jungle Fall 
Valley; Singapore Botanic Gardens. — 7970. Bt 
Timah Nat. Res. (mid-Feb.-mid-June);^ Botanic 
Gardens, Singapore 80-year old secondary forest, 
Johore. — 7977. Planning to collect in fresh-water 
swamp forest. 

Collections. Up to 1971 > 370 specimens 
of tree species, exclusively in Herb. Sing., H 

Literature. (1) R. D. Hill: 'Soil moisture 
under forest, Bukit Timah Nature Reserve, 
Singapore' (Gard. Bull. Sing. 26, 1972, p. 85-93, 
3 fig., 7 tabl.). 

*Himsen, A. 

collected some 20 Graminae and Cyperaceae 
from grass plains at Yambi, Sepik Distr., Terr, of 
New Guinea, in Jan. 7959; Herb. Lae. 


Hinds, Richard Brinsley 

Literature. (3) Add: Drawings of the 
'Botany of H.M.S. Sulphur' by Miss Drake. 

Biographical data. Add: In Hemsley & 
Hooker, Biol. Centr.-Amer. 4, 1887, App. p. 128. 

*Hirano, Minoru 

(1910, Shizuoka, Japan; x) graduated from the 
Faculty of Science, Kyoto University, in 1937. 
Professor of Yoshida College, Kyoto University 
Specialist in fresh-water Algae. During a collecting 
trip in Borneo he was accompanied by M. Hotta 
(see there) and M. Matsubara (ethnology student). 

Collecting localities. See sub Hotta. 

Collections. Herb. Kyoto Univ., Hirano & 
Hotta nos 1-1656.^ Probably several (? mostly 
cryptogams) in his own series. Sarawak dupl. in 
Herb. Edinb. 

Literature. (1) K. Iwatsuki: 'Ferns of 
Borneo, collected by M. Hirano and M. Hotta'I 
(Acta Phytotax. Geobot. 21, 1965, p. 91-100, 2 
fig.); M. Tagawa: 'Ferns of Borneo, collected, 
etc.' (ibid. 21, 1965, p. 173-180; 22, 1967, p. 183- 

H.M.B,, see H. M. Burkill, not to be used in 

*Ho Coy Choke 

collected plants on Mt Kinabalu, Sabah, at 
least in March 796^; H nos in Herb. Kuala Lump. 
Possibly during the 2nd Roy. Soc. Expedition (see 

Hochreutiner, Benedict Pierre Georges 

(d. 1959, Geneva, Switzerland). 

Biographical data. Taxon 8, 1959, p. 81- 
83, portr.; Verb. Schweiz. Naturf. Ges. 139, 1959, 
p. 408-417, portr., bibliogr.; Candollea 17, 1959- 
61, p. 9-24, portr., bibliogr. 

Holstvoogd, Coenraad 

(d. 1967, Amersfoort, Netherlands). 

Holttum, Richard Eric 

Add: After a study tour to the U.S.A. etc. from 
Nov. 1959-May 1960, he paid a short visit to 
Bogor (Java) and Singapore.* In 1964 he was 
awarded the Gold Medal of the Linnean Society. 
He occasionally makes herbarium study tours 
round the world. 

Literature. (5) Cf Report in Fl. Mai. Bull. 
no 15, 1960, p. 743-746. 

Biographical data. Add: Amer. Fern J. 
59, 1969, p. 1-3. 

Home, Sir James Everard 

Collections. Add: Forbes was right in 
attributing Timor plants to Home. Cynometra 
cauliflora was collected at Coepang (May 1 , 1843). 

Hoogerwerf, Andries 

Add: In 1959 appointed Scientific Officer of the 
Stichting Agrarisch Onderzoek t.b.v. Ned. Nieuw- 
Guinea (SAONNG). In charge of the investigation 
of the damage done by vertebrates (mainly aquatic 
birds) to paddy in the fields, and stationed at the 



Cyclopaedia of collectors — Supplement II 


Rice Plantation Koerik, W of Merauke at the 
mouth of the Kumbe River, in Dutch South New 
Guinea; from October 1962 till April 1963 
stationed at the Agricultural Experimental Station 
at Manokwari (Vogelkop). Mid-1963 he settled at 
Bakkum, N.H., in the Netherlands; he occasionally 
visits Indonesia. 

Author of many ornithological publications and 
reports with information on the vegetation. 

Collecting localities. Add: 1951. Prin- 
seneiland-Krakatau (Strait Sunda) (Sept. -Oct. ).^ — 

1953. S. Borneo: Kumain, Pangkalanbaru (Apr. 
13-17); .V. Billiton pr. Peganin (Apr. 20-22).— 

1954. Bawean (May-June);* Kangean (Aug.- 
Sept.). — 1955. Strait Sunda and Udjong Kulon 
{W. Java) (June-July); Kariiuondjawa (Oct.- 
Nov.).— From 1950-56 several times in Udjong 
Kulon ( \V. Java),^ and neighbouring islands 
in Strait Sunda.— 1959-62. South Dutch New 
Guinea''. — 1971. E. Java: Baluran, Jang, Meru 
Betiri Nature Reserve (Oct.). 

Collections. Add: Herb. Bog.: Prinsenei- 
land-Krakatau collection (1951); and part of the 
other collections which are for the greater part in 
Herb. Leyden. 

Literature. Add: (5) A. Hoogerwerf: 
'8ste Versiag van tournee's naar het Wildreservaat 
Udjung-Kulon (21 November-9 December 1950)' 
(mimeographed, Bogor, Dec. 1950, 95 pp., 30 
photogr.); 'Versiag over in 1951 en 1952 naar het 
Natuurpark Udjung Kulon gemaakte dienst- 
tournee's' (mimeographed, Bogor, Apr. 1952, 19 
pp., 31 photogr., map); 'Udjung Kulon the land 
of the last Rhinoceros' (Leyden, Brill, 1970, 512 
pp., 155 ill., 3 maps). 

(6) A. Hoogerwerf: 'Versiag over de expedities 
naar het Natuurpark Pulau Panaitan (Prinsenei- 
land) in Straat Sunda van 30 Augustus-5 October 
1951' (mimeographed, Bogor, Nov. /Dec. 1951, 
143 pp., 32 photogr., map); 'Versiag over een naar 
de Krakatau-groep gemaakte dienstreis van 5-15 
October 1951' (mimeographed, Bogor, July/Aug. 
1952, 184 pp., 32 photogr.). 

(7) A. Hoogerwerf: 'Enkele gegevens over het 
vogelschadeonderzoek bij het Rijstproefbedrijf 
'Koembc' te Koerik (nabij Mcraukc) Zuid Nieuw- 
Guinea' (subtitle: 'Some particulars on a research 
on harmful birds in ricecrops in south New- 
Guinea') (mimeographed. Bull. Agr. Research Stat., 
Manokwari, Agr. ser. no 7, 1962, with English 

(8) A. Hoogerwerf: 'Notes on the Island of 
Bawean (Java .Sea) with special reference to the 
birds' (Nat. Hist. Bull. Siam Soc. 21, 1966, p. 
313-340, pi. 29-36; ibid. 22. 1967, p. 15-103, map). 

Hoogland, Ruurd Dirk 

Later Principal Research Officer C.S.I.R.O. 
Land Research; in 1968 he accepted the post of 
Senior Research Fellow in the Research School of 
Biological Sciences, Canberra. Early 1969 he 
visited Ceylon in connection with the Flora of 
Ceylon Project. 

Author of a paper on Mt Wilhclm.' 

Evodiella hooglandii Van or k Linoln has been 
named after him. 

CoLLEr-TlNf, i.Of:Al.lTiES. Add: Territory of 
New Guinea. 1959. Around Lac (Oct. -Nov.). — 
1960. Western Highlands (with R. Schoddf) 


(June-Sept.), Wabag area.— 1961. with P. J. 
Darbyshire in the Sepik Distr.: Aitape-Lumi 
area (July-Aug.). — 1962. Not in New Guinea. — 
1963. Northern Division, Papua, Safia-Pongamia 
area (hardly collected, except specimens of 
Torrenticola queenslandica July 25, Barigi R. near 
Biriri, no 8616).— 1964. With the 7th Archbold 
Expedition to Huon Peninsula in Terr, of New 
Guinea: Pindiu (Apr. 21-May 2, nos 8815-8852), 
Marba Creek (May 2-23, nos 8853-9033), Mt 
Rawlinson (June 3-July 4, nos 9034-9359), 
Mannasat (Cromwell Mts, July 20-Aug. 23, nos 
9361-9677; 9678-9685 collected en route), Mon- 
arauwe (Salawaket Range, 9500 ft, nos 9686-9785, 
Aug. 30-Sept. 7), Tempanpan (Salawaket Range, 
Sept. 7-Oct. 7, 11000 ft, nos 9786-10,013), en 
route to Kabwum (Oct. 7, nos 10,014-10,017).— 
1965. No collection in New Guinea. — 1966. 
Sepik Distr. with Craven; Hunstein Range and 
Mt Hunstein. — 1969. Mainly coll. Dilleniaceae 
and Saurauia in: Singapore (Jan. 16-19); Ceylon, 
India, Thailand; Penang and Kepong (May 23- 
27), Singapore (27-31); W. Java: Bogor and 
Tjibodas (31 -June 5).— 1973 (mainly Saurauia, 
some Dilleniaceae). Malaya (May 22-June 4): incl. 
Cameron Highlands (May 25-27), Eraser's Hill 
(29-30), UIu Gombak (31), Johore (June 3^); 
Singapore (4-8). Java (8-22), incl. Langkong (1 1), 
Jasinga (13). Cibodas (= Tjibodas) (14-18); 
Ciapus ( = Tjiapoes) Gorge (20), Telaga Warna 

Collections. =* About 12400 «05'. Also dupl. in 
Herb. Edinb. 

Literature. (1) Cf. Blumea Suppl. 4, 1958. 
p. 220-238, 6 photogr. 

(2) Mosses by E. B. Bartram in Brittonia 1 1 , 
1959, p. 86-98; Rev. Bryol. Lichen, n.s. 30, 1961, 
p. 185-207. 

B. L. VAN DER Linden: 'Evodiella, a new genus 
of Papuan Rutaceae' (Nova Guinea n.s. 10, 1959, 
p. 143-148, 1 fig.). 

*Hooley, Bruce Arthur 

(1931, Daylesford, Vict., Australia; x) gradu- 
ated with B. Chem E from the University of 
Melbourne in 1954 and joined the Summer 
Institute of Linguistics in 1958. He carried out 
linguistics research in the languages of New 
Guinea, especially the Buang language of Morobe 
District. In 1962 M.A. in linguistics from the 
University of Pennsylvania, and in 1968 engaged 
in completing work for a doctorate at the same 
university. He served as Chairman of the Lin- 
guistic Committee of the New Guinea Branch of 
the SIL and as principal of the Institute's Summer 
School at Brisbane; in I908 Associate Director of 
the New Guinea Branch of SIL. 

Author of linguistic papers. 

Collecting localities. In connection with 
the Buang language in Morobe District, Terr, oj 
New Guinea. 

Collections. About 160 specimens in Herb. 

*Hopc, Capt. 

see Cycl. Fl. Mai. I. I. 1950; identical? 

Colllctions. Plants from Malay Pcnin.iula 
in Herb. Edinb. (from Glasgow Herb., pres. 1888). 



Flora Malesiana 

[ser. I, vol. 8^ 

*Hope, G. (S.) 

of the Dept of Biogeography and Geomor- 
phology, Australian National University, Can- 
berra. He collected on Mt Wilhelm (1969-70), E. 
New Guinea; Herb. Univ. Canberra, ANU numbers 

A thesis on a vegetation history of Mt Wilhelm 
was due to be submitted, to Prof. D. Walker (see 
there), June 1972. 


In 1971 he accompanied an Australian ex- 
pedition to Mt Carstensz in W. New Guinea for 
botanical and palynological work. 

♦Hornabrook, Richard William 

(1925, Wellington, New Zealand; x), neurologist 
(M.D. New Zealand 1956), Director of the 
Institute of Human Biology, Port Moresby, 
Papua. Clinical Investigator, Kuru Research, 
Eastern Highlands New Guinea, 1963-65. 

Author of medical and entomological papers of 
Kuru etc. in various journals. 

Collecting localities. 1963-65. New 
Guinea: Eastern Highlands, especially in Okapa 

Collection. In Herb. Lae. 

Horsfield, Thomas 

Collections. Add: In the India Office is a 
volume of 337 folios which appears (according to 
Dr John Bastin) to be Bennett's (?) rearrange- 
ment of Horsfield's papers, giving Javanese and 
(or) Latin names of plants, . . . Catalogues of 
plants etc. The official description in the I.O. 
includes: localities where found. The lists are 
mainly in two handwritings, one that of Hors- 


field, the other that of a copyist (B. /./. July 1970). 
An 'Algemene naamlijst der Planten van Java' 
from about 1810 is in theRijksarchiefinTheHague 
(Arch. Binnenl. Zaken, afd. Onderwijs, exh. 8 
maart 1837 nr. 114). 

His collection of drawings in the India Office 
Library includes some of mosses from Java {nos 

Literature. (6) The cited English paper was 
not published in 1817, and never before 1848 in 
J. Ind. Arch, and East. Asia. Dr J. Bastin has 
drawn my attention to this error which was 
probably based on an advertisement in some 
copies of Raffles' History of Java. 

(9) Add: On the drawings see Cl. Nissen, 'Die 
botanische Buchillustration' 1951, vol. 2, p. 86. 

(1 1) C/. M. Archer: 'Natural History Drawings 
in the India Office Library' (London 1962) p. 47, 

Biographical data. Add: Portr. in Str. 
Times Annual for 1971, p. 60; J. B. Stanley: 
'Horsfield in the Indies' {ibid, for 1972, p. 85-88, 
portr., ill.). 

Hose, Charles 

Rightly: was born at Willian (not 'Witham') 
in Hertfordshire as a son of Rev. T. C. Hose (a 
brother of Bishop Hose), and died at Croydon, 
London (C. F. Cowan in lift. Feb. 12, 1968). 

*Hose, Edward Shaw 

(1871, Malacca; 1946 Normandy, Surrey, 
England), son of Bishop Hose, cousin of Charles 
Hose. In 1891 he entered the Malayan Civil 
Service, 1924-25 Colonial Secretary, Straits 
Settlements, Acting Governor of Singapore (1925); 
retired 1925. 

Collecting localities.^ 79/7. Malay 
Peninsula: Perak, G. Hijau, and Telok Anson, 
when stationed there; Sembilan Is (1918). 

Collections. Herb. Kew and Sing. 

Literature. (1) Cf. Burkill in Gard. Bull. 
Str. Settl. 4, 1927, nos 4-5. 

Biographical data. C. F. Cowan in lift. 
Feb. 12, 1968. 

*Hose, Ernest 

(1872, Roydon, Norfolk, England; x), younger 
brother of Charles Hose, to Sarawak as col- 
lector 1893-96, mainly in Baram District and 
Natunas in 1894. Inl896 he joined H. H. Everett, 
planting Cutch at Santubong; in 1901 joined the 
Borneo Co. and persuaded them to let him plant 
Hevea rubber. Started the first rubber plantations 
in Sarawak at Dahan Estate (1902), and at Sungei 
Tengah Estate (1907-19). Retired 1919. 

Dolichos hosei Craib was named after him and 
not after Rev. G. F. Hose. 

Collecting localities. 1894. Natuna Is- 
lands (July-Oct.): Bunguran, P. Laut, and some 
small islands. 

Collections. Mainly zoological, in B.M. 
(Nat. Hist.), and to Lord Rothschild. Plants 
from Natuna Is in Herb. Kew. 

Biographical data. C. F. Cowan in litt. 
Feb. 12, 1968. 

Hose, Ernest S. 
The entry in the Cycl. Fl. Mai. I, 1, 1950 is a 



Cyclopaedia of Collectors — Supplement II 


mixture and must be replaced by Ernest Hose 
and Edward Shaw Hose (see there). 

Hose, Rev. George Frederick 

Collections. Add: In 1940 his daughter 
Gertrlt) had a 'large' collection of (living?) 
ferns from Borneo and Malaya, made by her 
father, which she presented to the Botanic Gardens 
at Edinburgh (C. F. Cowan in litt. Feb. 12, 1968). 

Hose, Gertrud 

Add: (1883, Sarawak; x), from 1883-88 and 
1903-08 in Sarawak with her father Bishop Hose. 
from 1923-25 in Malaya with her brother Edw. S, 
Hose (C. F. Cowan in litt. Feb. 12, 1968). 

Hosseus, Carl Curt 

(d. 1950, Cordoba, Argentine). 

Biographical data. Add: 
bot. vol. 10, no 1, p. 32. 

Rev. Sudamer. 

•Hotta, Mitsuru 

(1937, Osaka, Japan; x) graduated in 1960 from 
the Agricultural Department University of Osaka 
Prefecture and was subsequently admitted to the 
graduate course of Kyoto University. Lecturer 
Biological Laboratory, Kobe Women's College, 
Aoyama, Suma-ku, Kobe-si. 

In 1960 he was a member of the Tonga and Fiji 
Expedition of Kyoto University; in 1963-64 he 
collected in Borneo with Professor M. Hirano 
(see there) of Osaka City University. 

Author of papers on Araceae. 

Collecting localities. Borneo} Brunei: 
Tutong (Oct. 6, 1963); Sarawak: Bintulu and 
Minah camp (Oct. 10-19, with Hirano), Ulu 
Minah and Sinonok (Oct. 23-28), Bintulu (Oct. 
31, Nov. 4), Ulu Kakus and Bt Keyan (Nov. 7-13, 
with Hirano), Bt Kana (18-22, with Hirano); 
Brunei: Seria and Bt Puan (Dec. 5-7), Merimbun 
(10-11), Bt Teraja (16-25), Brunei Town (Jan. 6 
and 10, 1964), Brunei Tembrong (Jan. 19-Feb. 2); 
Sarawak: Minah camp (Feb. 23-29), G. Mulu 
(Mar. 15-22). — 1968. Sabah, focussing special 
attention to the Kinabalu region in 7969. 

Collections. Herb. Kyoto Univ.: Hirano & 
Hotta nos 1-1656; Hotta nos 12300-15847. 
Many cryptogams,* but phanerogams' too. 
Duplicates partly distributed to Tokyo Nat. Sc. 
Mus. (Cyp., Gram.), New York Bot. Card. (Cyp.), 
Mveral families to Herb. Tokyo Univ. and many 
groups to Herb. For. Dept Sarawak and Brunei; 
Sarawak dupl. to Herb. Edinb. 

Literature. (I) M. Hotta: 'Itinerary of the 
Borneo (Notes on the Vegetation)' (Acta Phytotax. 
Geobot. 21, 1965. p. 153-160, maps 1-4). 

(2) Cf. literature sub Hirano. 

(3) M. Hotta: 'Notes on Schismatoglottidinae 
of Borneo' I (Mem. Coll. .So. Univ. Kyoto scr. B 32, 
1965, p. 19-30. 5 fig.); 'Notes on Borncan Plants' 
I-ir (Act. Phytotax. Geobot. 22, 1966. p. 1-10. 3 
fig.; ibid. 1967. p. 153-162. 7 fig.); 'Notes on the 
wild banana of Borneo' (J. Jap. Bot. 42. 1967, p. 
344-352. 1 fig., pi. 7). 

•Hou, Ding 

(1921, Hsinpkan. Kiangsi. China; x), botanist, 
B.S. in the National Chung-Cheng University, 
Kiangsi (1945); from 1945 47 A.ssistant in the 

same university; Assistant, Nat. Taiwan University 
1947-51; Research Assistant, subsequently Assis- 
tant Missouri Botanical Garden, St. Louis, 1951- 
55; Botanist Arnold Arboretum, Cambridge, 
Mass., 1955-56; Botanist Foundation Flora 
Malesiana 1956-60; since 1960 Scientific Officer 
Rijksherbarium Leyden. He made a study trip 
to the East in 1966, to familiarize himself with 
living Anacardiaceae. 

DING HOU, Mt Kinabalu, 1966 

Itinerary.' 1966. Singapore I. (Apr. 6-20), 
visiting Johorc (9), Kranji Nature Reserve (10), 
MacRitchie Reservoir Nature Reserve (14), Bt 
Timah Nat. Res. (16). Pasir Ris (17-18). N. 
Borneo (Sabah): Sandakan (21-27). visiting the 
nearby Leila For. Res., Badu Sapi (mangrove 
forest), Berhala For. Res.; to Lahad Datu (28), 
visiting the Eastern Lowlands (29-May 17), Mt 
Salim (29-30). Lahad Datu (30-May 3). P. Sakar 
(2), Kennedy Bay and the Silabukan For. Res. 
(3-4), Kennedy Bay to Kunak (Mostyn), on the 
way visiting P. Rohayan (4), Madai For. Res. (5-7) 
and return to Mostyn (7); to P. Tahauwan (9), to 
Semporna (10), Quoin Hill and Membalua For. 
Res. and Mt Andrassy (11-15); Tawau (15-17), 
making excursions to Tawau Hills and Hot 
Spring For. Res.; back at Sandakan (19 24), 
visiting Leila For. Res. (21). virgin forest near 
Long's Farm (21-22). Scpilok For. Res. (23); 
Jcsselton (25). Mt Kinabalu (26-Junc 9): National 
Park Headquarters vicinity (26-27), Tenompok 
(29), to the summit and back to Sosopodon 
(30-Junc 1), to Layan-layan Mountain hut (c. 
2600 m, 30), Panar Labah Mountain hut (c. 




Flora Malesiana 

[ser. I, vol. 8^ 

3300 m, 31), summit (c. 4200 m, June 1); Kunda- 
son (2), to Mesilau and staying in the bungalow 
of the Roy. See. Exp. (3-6), Sosopodon (7-9), 
Ranau (8), Jesselton (10-14), visiting Tuaran 
Agriculture Station and mangrove forests (13); 
Sarawak: Kuching (16-18), visiting Bau (18), 
Bintulu (19), Nyabau For. Res. (20-27), Bintulu 
(29), Segan For. Res., Nanga Sopulow (30-July 5), 
Bintulu (6), to Sibu (7), peat swamp forest (8), 
Bt Raya (9-11), back at Sibu (12), Bako National 
Park (14-17), Kuching (18-22), Arboretum 
Semengoh (20, 23-29); Kuching (30-Aug. 9), 
visiting Bau Limestone Hills (2), Samariang 
mangrove forest (5), Mt Medang (8); Malaya: 
Kuala Lumpur and to Kepong (10), to Kuala 
Kangsar, Perak, on the way to G. Bubu (12), G. 
Bubu(13-18);Kepong(19-31), visiting BtLagong, 
Ulu Gombah For. Res. (24), Telok For. Res., 
Klang (25), Batu Caves limestone hill (26), Kuala 
Selangor mangrove (28), Sg Meryak, Negri 
Sembilan (29), Batu Caves (31); to Kuantan, 
Pahang (Sept. 1), on the way visiting Kemasul 
For. Res. near Bentong, Menchli For. Res. (2), 
to Kuala Dungun, Trengganu, on the way visiting 
Balok For. Res. (3), Kuala Trengganu (4-5), 
visiting Bukit Bauk For. Res. and Jerangau State 
Land Forest; return to Kepong (6), to S. Patani, 
Kedah (8), G. Jerai (Kedah Peak, 9-11); P. Penang 
(12-15), Botanic Garden and Telok Bahang (13), 
Penang Hill For. Res. (14), Pandai Achech For. 
Res., Telok Bahang (15)& Cameron Highlands, 
Pahang (16-17); Kepong (18), Kuala Lumpur 
(19-24) and excursion to mangrove forest Klang 
and Kuala Selangor; Thailand (26-Oct. 3); back 
in Holland (4). 

Collections. About 750 nos, partly with 
material in spirit;^ some wood samples. In Herb. 
Leyden, numbering 115-869, including some 
cryptogams too. 

Literature. (1) Ding Hou: 'Report of a 
Study-trip on Anacardiaceae to Malaysia and 
Singapore in 1966' (stencilled, 23 pp., 3 maps 
giving routes followed); Fl. Mai. Bull, no 21, 1966, 
p. 1394-1395. 

(2) Ding Hou: 'Sarawakodendron, a new genus 
of Celastraceae' (Blumea 15, 1967, p. 139-143). 

Houttuyn, Martinus 

Add: The presence of sheets which have be- 
longed to the Houttuyn Herbarium has been 
proved beyond doubt. At least four sheets have 
been located in the Rijksherbarium Leyden, all 
collected by Thunberg in Java. Hoogland^ 
commented on the first. At least three others have 
been found as shown by Van Steenis & Veld- 


Literature. Add: (3) R. D. Hoogland: 'The 
genus Tetracera (Dilleniaceae) in the Eastern Old 
World' (Reinwardtia 2, 1953, 185-224, pi. 1, 
especially p. 207 and pi. 1). 

(4) C. G. G. J. VAN Steenis & J. F. Veldkamp: 
'Note on Houttuyn's Herbarium' (Blumea 21, 
1973, p. 151-152, with facsimile handwriting). 

Howison, James 

Add to Suppl. Cycl. Fl. Mai. I, 5, 1958: Author 
of 'Some Account of the Elastic Gum Vine of 
Prince of Wales' Island, etc' (As. Res. 5, 1797, 
p. 157-165). He certainly was the discoverer of the 


caoutchouc vine, but Roxburgh made his de- 
scription after material collected by Ch. Campbell 
in Sumatra, which was forwarded by Fleming. 

*Hughes, Cecil Graham 

(1913, Sydney, N.S.W., Australia; x) graduated 
B.Sc. in Agriculture from Sydney University in 
1934. Shortly after he joined the Bureau of Sugar 
Experiment Stations and has been working with 
that organization ever since. Associated with cane- 
breeding and diseases of sugar-cane. 

He joined the 1951 New Guinea Expedition in 
search of cane varieties. 

Itinerary, collections, etc. cf. sub J. H. Buzacott. 

*Hujah Kabun, see Forest Dept North Borneo. 

*Hulskanip, R. F., see Boswezen Nieuw Guinea. 

HZ, see H. Zollinger. 

His personal set was marked with these initials. 

Idenburg, Abraham Gerrit Adrianus 
(d. 1949). 

*Ijiri, Shoji 

(1913, Otaru city, Hokkaido, Japan; x), gradu- 
ated from the University of Tokyo (1936), specialist 
in Palaeontology and Odontology of Mammals; 
Dr. Sc. Tokyo (1949). 

Collecting localities. West New Guinea 
from March to June J943, with the Satake (sec 
there) party. 

Collections. About 700 sheets of flowering 
plants; in Herb. Nat. Sc. Mus. Tokyo. 

♦Iking, S. 

Plants from Java in Herb. Arn. Arbor. See 
Index Herb. Pt II (Regn. Veg. 86) 1972. 

*Ilias bin Paie 

collected in Sarawak from 1962 onwards. 
Collections. Herb. Sarawak; dupL in Herb. 


Add: Joseph C. (1893-x). See Index Herb. Pt II 
(Regn. Veg. 86) 1972. 

*Irmscher, J. 

is mentioned as the collector of plants in 
Borneo; specimens in New York and Paris (rec. 
1935). This record in Index Herb. Pt II (Regn. 
Veg. 86) 1972, seems doubtful to me. 


(d. Oct. 29, 1963, Bogor, Indonesia). 


Add: Frans Karel van (1884-x). 

Iterson, G. van 

(d. 1972, Wassenaar, Z. H., Netherlands). 

*Ives, David Wilson 

(1939, Waimate, New Zealand; x) has been 
educated at the University of Canterbury, Christ- 
church, New Zealand, B.Sc. (NZ), M. Agric. Sci. 
1st Hons (Cantaur.). From 1962-67 employed as 



Cyclopaedia of Collectors — Supplement II 


soil surveyor under U.N. Special Fund in Sabah, 
and by New Zealand Colombo Plan, in Pahang, 
both Malaysia. He worked in Indonesia on N.Z. 
Colombo Plan aid to Indonesia Beef Industry 
feasibility study on behalf of International Bank 
for Reconstruction and Development as part of 
team from consultant group, Asian New Zealand 
Development Consultants (A.N.Z.D.E.C.). 

Collecting localities. Indonesia. S\V. 
Celebes (Sulawesi Selatan): Pare Pare. Enrekang- 
Rappang and Wadjo, together with O. Roberts. — 
1971. Lesser Sunda Is. (June-July): Siimha, at 
Kabaroe Karendi, and near Kodi; Sumhawa, 
at Plampang. — 1973 (Feb. -Mar.). SE. Borneo 
(Kalimantan Selatan): near Tandjung, NE of 
Pleihari and along Sg. Tabanio; around Djilatan, 
Sebuhur, Djurong area SE from Pleihari. Centr. 
Sumatra: area E of Blang Bintang airport and N 
of Indrapuri; Padang Lawas area to E of Gunung 
tua and SW of Langga Pajung. 

Collections. Herb. Christ ch., specimens 
determ. by B. E. V. Parham and E. Edgar. 

Iwanggin, Gerrit Th. 
collected in the BW series (see there). 

*Iwatsuki, Kunio 

(b. 1934; x), bryologist, member of the Kyoto 
University Biological Expedition to Thailand 
(Aug. 7-bct. 10) and Malaysia, 1967. Other mem- 
bers were T. Shimizu, N. Kitagawa, N. Fukuoka, 
M. Hutch, J. Ashida, M. Tagawa, M. Hirano, 
S. Ueno. In Aug.-Sept. 1971 he accompanied an 
expedition to W. Sumatra. 

Itinerary.' 1967. Singapore and Bt Timah 
(Aug. 2); Malaya: Templer Park and Batu Cave 
(Aug. 5-6); Penang (Oct. 13-16); Cameron 
Highlands (18-21); Templer Park (25); Fraser's 
Hill (26), G. Gombo(c)k (28-29); Batu Cave (30), 
Gua Musang (Oct. 31-Nov. 2), G. Pulai and G. 
Panti in Johore (2-^)\ Bt Timah (6). 

Collections. 1st set in Herb. Kyoto, 2nd 
set in For. Herb. Bangkok. The Malayan specimens 
numbered with the prefix 'M' (Thailand with 'T'). 

Literature. (1) K. Iwatsuki: 'Biological 
expedition to Thailand and Malaysia' (Acta 
Phytotax. Geobot. 23, 1968, p. 127-128, 1 map). 

Biographical data. Portr. in Hatusima, 
Flora Ryukyus 1971, p. 71. 

Jaag, Otto 

Add: Biographical data. In Festschrift 
O. Jaag (Schweiz. Z. Hydrobiol. 22, 1960, p. 9-17, 
portr.); Wasser Abwasscr 111, 1970, p. 175-246, 
portr. (70 years old). 

Jack, William 

Literature. (7) Add: Dr J. Ba.stin kindly 
drew my attention to Don's Account of Lambert's 
Herbarium, Appendix p. 31, where it reads: "The 
late Dr. William Jack, who was attached to the 
suite of Sir Stamford RafUcs.Governor of Sumatra, 
has sent all the specimens described in the first 
volume of the 'Malayan Miscellanies' among 
which arc three splendid species, of that most 
remarkable genus of plants. Nepenthes." 

Biographical data. Add: J. Bomb. Nat. 
Hist. Soc. 56. 1960, p. 449-456. 


Jacobs, Marius 

In 1959 appointed, on the staff of Flora Malesi- 
ana Foundation, at Leyden; subsequently on the 
start' of the Rijksherbarium; Ph.D. Leyden (1965). 

In 1963 he joined a bio-archaeological ex- 
pedition in West Persia for some months. 

Cyclosorus jacobsii Holtt. has been named 
after him. 

gerrit th. iwanggin, 

noteworthy forest exploration ofHcer, Hollandia/ 

Manokwari, 1957-1961. 

Collecting localities. 1957. Add: W. 
Java: G. Gedch, above Tjibodas (Nov. 6); oc- 
casionally in Horl. Bog. — 1958. Hort. Bog.; 
Tjibodas (June 15). G. Pangiango (16), G. Gedeh 
(17), Tjibodas (18). Leaving Bogor (July 20), 
arriving in Sarawak, NW. Borneo, at Kuching 
(28); with J. A. R. Anderson to Mt Penrissen 
(Aug. 1-9; coll. nos 5000-5138); to Sungei Serin 
limestone hills (10-12, nos 5139-5183); to Belaga 
on the Rajang River. Segaham Range (Aug. 20- 
.Scpt. 7. nos 5184-5465); delta of the Rajang River 
near Sibu (Sept. 1 1 . nos 5466-5468); Scburuan Bau 
limestone hill near Kuching (14, nos 5469-5484); 
Bako National Park (16-17, nos 5485-5521); Mt 
Matang (19-21, nos 5522-5568); Brunei (arriving 
Sept. 25): with P. Shaw Ash ion to E. Brunei, 
Kuala Belalang (.Sept. 29-{)ct. 5), riverbeds (nos 
5569-75651); W. Brunei. Andulau Forest Reserve 
(8. nos 5652 5662), near Telamba bridge on the 
road Brunei-Kuala Belait (9, nos 5663-5695), 
Badas Forest Reserve (10, nos 5696-5698), Brunei 
town (10, no 5699); to N. Borneo (Oct. 12); 
Ranau (13, no 5100) and with D. L NiciioisoNto 
Mt Kinabalu (14 18, nos 5701-5792);=" leaving 
Ranau (20), and Singapore (28) for F-.urope. — 
1968. Philippines, Luzon. I. caving for Baguio with 
D. R. MiNDo/A and R. Fspiritu (Jan. 17), camp 



Flora Malesiana 

[ser. I, vol. 81 

at Babadak, SW. slope Mt Pulog* (c. 2400 m), and 
in the top area (from Jan. 28) at c. 2700 m; 
climbing summit on the 31st; return to Babadak 
(Feb. 7-10); Mt Tabayoc, camp at 2350m (Feb. 
14), visiting the summit (16), 2850m; return to 
Manila (22); Mar. 7 to lowland forest in the 
Sierra Madre mountain chain, ^ E. Luzon, col- 
lecting from sea-level up to about 1200 m NE and 
NW of Dingalan; camp on the road to Baler 
(Mar. 21-25). S. Sumatra, Prov. of Lampung:* 
above Giesting on the E. Slope of G. Tanggamus 
at 1100m alt. (base camp), collecting from Apr. 
25-May 4 (nos 8021-8274), reaching the summit 
on Apr. 25; primary forest NW of Kota Agung at 
350-450 m alt. (May 9-19; nos 8275-8515). 

Collections. Borneo collection in Leyden, 
Kew; resp. in Kuching, Brunei, or Sandakan (the 
territory where the collection was made); Geneva, 
Wash., Canberra, Berlin, Stockh., Manila, Ithaca. 
In Herb. Sing.: Borneo dupl. Annonaceae and 

Collections made in Hort. Bog., and on G. 
Gedeh (Nov. 6, 1957; June 1955) are without 
numbers, mostly for Bailey Hort. Ithaca, N.Y.; 
dupl. in Dehra Dun. 

Mosses from the Philippines (1968), about 600 
collections, will be numbered in a B series after 
study by A. Touw. 

Literature. (1) Replace by: M. Jacobs: 
'Contribution to the Botany of Mt Kerintji and 
adjacent area in West Central Sumatra' I (Ann. 
Bog. 3, 1958, p. 45-79, 24 photogr., 1 map). 

(2) Replace by: M. Jacobs: 'Botanical Recon- 
naissance of Nusa Barung and Blambangan, 
South East Java' (Blumea Suppl. 4, 1958, p. 68-86, 
map, 4 photogr.). 

(3) M. Jacobs: 'Plant Life on Mount Kinabalu' 
(Mai. Nat. J. 15, 1961, p. 134-139, map, photogr.). 

(4) Cf. Fl. Mai. Bull, no 23, 1969, p. 1686-1687. 

M. Jacobs: 'The Plant World on Luzon's high- 
est mountains' (Publ. Rijksherbarium 1972, 32 pp., 
16 photogr., 1 drawing, 1 map). 

(5) M. Jacobs & D. R. Me>jdoza: 'Opportun- 
ities and suggestions for botanical exploration 
work in the Sierra Madre Mountains, Luzon* 
(Rijksherbarium Leiden, 1969, folio, stencilled, 
14 pp., 1 map). 

(6) Cf. Fl. Mai. Bull, no 23, 1969, p. 1685; M. 
Jacobs: 'Botanical Exploration in the Lampung 
Province, Sumatra' (Reinwardtia 8, 1972, p. 345- 

Biographical data. Curr. vitae in Thesis 
Leyden 1965. 

*Jacovlev, Professor 

from Sovjet Russia, accompanied A. Dilmy 
(see there) to Bali in 1963, to study pioneer plants 
on Mt Agung. 

Jacquinot, Honore 

Add: Was admitted as a doctor in 1848, and 
practised at Nevers. Brother of the commander 
of La Zelee (see J. B. Hombron). 

Biographical data. In Bibl. Austr. Entom. 
1775-1930 (R. Zool. Soc. N.S.W. 1932, p. 62). 

♦James, P. W., see sub A. C. Jermy. 

Jannet, see Jaunet. 

* Jaswir, see Forest Dept North Borneo. 

* Jaunet, Victor 

whose name was erroneously written 'Jannet' 
(see Suppl. Cycl. Fl. Mai. I, 5, 1958) in Roxburgh 
Hort. Beng., was Director of the Governor's 
Garden at Ile-de-France (= Mauritius) and 
Bourbon.' As already assumed he distributed 
material from the garden to Calcutta, etc., from 
about 1809-1811. 

Literature. (1) Cf. Ann. Mus. Hist. Nat. 
Paris 16, 1810, p. 484, 485. 

*Jawanting, A., see Forest Dept North Borneo. 

Jensen, Hjalmar 

Biographical data. Add: Dansk Bot. 
Arch. 21, 1963, p. 64-65, 167 (bibliogr.). 

Jensma, J. 

Add: (d. 1965, 76 years old), former teacher 
at a secondary school at Leeuwarden (Fr.), 

Biographical data. Vakbl. Biol. 45, 1965, 
p. 81. 

* Jensma, Jan Rienik 

(1927, Vrouwenparochie, Fr., Netherlands; x) 
was educated in horticulture at Wageningen 
College from 1947-53, taking his Dr's degree in 
1957. In the first half of 1950 he served in the 
Dutch arm.y at Bogor. 

Collections. At the request of Prof. Venema 
he brought back one representative of each 
family, in the Bogor Botanic Gardens, assisted by 
a mantri, collected Nov. 1949-i\j\y 1950; in total 
130 nos. Herb. Wageningen. 

* Jermy, A. Clive 

(1932, Strumpshaw, Norfolk, England; x) 
graduated B.Sc. (Hons) in Botany at University 
College London in 1954; post-graduate research 
for Ph.D. in taxonomy at Leicester University; 
1957-58 Assistant-teacher at Gravesend, Kent, 
Secondary School, instead of military training for 
conscription. Since Oct. 1958 in charge of the 
Fern section of the British Museum (Natural 

Leader of the British Museum-Newcastle upon 
Tyne Expedition, 1964-65, to East New Guinea.^ 
Members : T. G. Walker, A. Eddy, P. W. James; 
C. D. Sayers and R. Pullen were both seconded 
to the expedition, at least in 1964 to the Finisterre 

Itinerary. British Museum-Newcastle upon 
Tyne Expedition, 1964-65.* 1964. E. New Guinea: 
for the Finisterre Mts see sub C. D. Sayers; 
lowland rain-forest around Lae (Dec. 12-Jan. 3) 
by the four U.K. members only. — 1965. Vagau 
and Herzog Mts, 3000-6000 ft (Jan.), in company 
with Sayers. Eastern Highlands: Kainantu; Feb. 
4 Eddy, Walker, and Jermy to Aiyura (coll. 
5000-7000 ft), Mt Elandora of the Kratke Mts 
(Feb. 11), 4 days in Okapa area. Jermy and Eddy 
to the Upper slopes of Mt Wilhelm, Bismarck Mts 
(Feb. 16-Mar. 5), from Lake Pindeunde. In the 
meantime Walker (see there) to New Britain. — 
1969 (end). Central Celebes and S. New Ireland. 



Cyclopaedia of Collectors — Supplement II 


Collections. The expedition concentrated 
mostly on ferns and other cryptogams; in total 
over 1000 phanerogams (pentuplicates) were 
collected. In Brit. Mus., Kew; representative sets 
to Lae and Canberra. 

Literature. (1) A. C. Jermy & C. D. Sayers: 
'An Expedition to the Finisterre Mountains of 
Eastern New Guinea' (J. R. Hort. Soc. 92, 1967, 
p. 117-128, fig. 54-55, 57-63, 1 map). 

A. C. Jermy: "British Museum (Nat. Hist.)- 
University of Newcastle upon Tyne Expedition' 
(32 pp., map, 2 pi., 9 photogr.). 

Jeswiet, Jacob 
(d. 1960, Bennekom, Gld., Netherlands). 

Jochems, Sarah Comelis Johannes 
(d. 1966). 

•Joenoes, see Forest Research Institute, Bogor. 

*Johns, R. J. 

M.Sc.; employed in the Division of Botany at 
Lae, Terr, of New Guinea, at present teacher at 
Bulolo Forestry College. 

He accompanied Dr B. O. van Zanten (see 
there) on a trip in E. New Guinea in Oct.-Nov. 

Joint author of a Check list of Mount Wilhelm 

He probably collected in both the NGF and 
LAE series. 

Literature. (1) R. J. Johns & P. F. Stevens: 
* Mount Wilhelm Flora: A Check List of Species' 
(Bot. Bull, no 6, 1971, 60 + 4 pp., 7 fig., 1 map). 

'Johnson (nee Garrard), Anne 

(1928, London, England; x) graduated B.Sc. 
(Special) at London (1949); Ph.D. 1951. Demon- 
strator in Biology, University of Malaya, Singa- 
pore, 1951-54; Librarian Singapore Botanic 
Gardens, 1954; 1954-64 Lecturer in Cryptogamic 
Botany, University of Malaya; 1964-66 Senior 
Lecturer, University of Singapore; 1966-68 
Professor of Botany, University of Malaya, Kuala 
Lumpur, now in the Biology Department, Nan- 
yang University, Singapore.' 

Collecting localities. 1951. Malaysia: 
Malay Peninsula, Singapore, Sarawak. — In 1967 
at Jambu Bongkok, Trengganu (Feb.-Mar.); G. 
Benom, Pahang (Apr.);' Langkawi Is (May). 

Collections. Mainly bryophytes and pterido- 
phytes, nos 1-4000 in Herb. Sing. Univ.; nos 4050- 
4378 in Herb. Univ. Malaya Kuala Lump. 

Literature. (1) Author of 'An account of 
the Malaysian Leucobryaccae Clumut puteh')' 
(Gard. Bull. Sing. 20, 1964, p. 315-360); 'Malayan 
Botany' (East. Univ. Press 1966); 'Bryophytes and 
Filmy Ferns from Gunong Bcnom' (Bull. Br. 
Mus. (Nat. Hist.) Zool. 23, no 2, 1972, p. 17-20); 

Biooraphical data. Who's who in Ma- 
laysia and Singapore 1967. 

'Johnson, Lawrence Alexander 

0925, Cheltenham, near Sydney, Australia; x) 
graduated from the University of Sydney (1948); 
botanist at the National Herbarium of New South 
Wales since 1948 (from 1962-63 Liaison Officer at 

Kew); 1972- Director of the Sydney Botanic 
Gardens and the N.S.W. National Herbarium. 

He attended the opening of the Lae Herbarium 
in New Guinea. 

Collecting localities. Between 1962-63. 
Some cultivated Casuarina spp. in Singapore. — 
1965. East New Guinea: Port Moresby area (Apr. 
6), Sogeri area (7); Wau, Bulolo, Mt Kaindi (8-9); 
Bulolo-Lae (10), Lae (11-12), Aiyura-Kainaniu 
area in Eastern Highlands (13-14). 


Collections. In Nat. Herb. N.S.W.: c. 25 
nos from New Guinea. He collected some herbarium 
and cytological material of selected groups of 

Liter ATURE. (I) Cf. Pac. Sc. Ass. Inf. Bull. 19. 
no 4, 1967, p. 9. 

Some results incorporated in B. G. Briggs & 
L. A. S. Johnson: 'The Status and relationships of 
the Australasian species of Typha' (Contr. N.S.W. 
Nat. Herb. 4). 

'Johnston, A. M. 

collected BucUlleja sp. no 75 in the Cameron 
Highlands, Pahang (Jan. 27, 1952); in Herb. Sing. 
Possibly Anne Johnson (see there). 

*Jones, William 

Plants from the Philippines (coil. 1908-09) in 
Herb. Chicago Field Mus. Sec Index Herb. Pt II 
(Rcgn. Vcg. 86) 1972. 

Jong, Jan Kornclis dc 
(d. 1972, Hilvcrsum, Netherlands). 




Flora Malesiana 

[ser. I, vol. 8^ 

Biographical data. Vakbl. Biol. 52, 1972, 
p. 342. 


collected 18 varieties of sugar-cane on Tabar J. 
and New Ireland in 1921. He was sponsored by 
the Colonial Sugar Ketining Co of Australia.* 
Whether any herbarium specimens were made is 
unknown to me. 

Literature. (1) C/. Hawaiian Plant. Rec. 55, 
1958, p. 213; Cane Growers' Quart. Bull. Brisbane 
15, 1951, p. 40. 

*Juliano, Bienvinido O. 

collected a Scleria on the College Campus, 
Laguna, Luzon (Sept. 1952). 

*Juliano, Jose Buencamino 

(1900-x). Plants from the Philippines in Herb. 
Stanf. and Rocky Mts Herb. Un. Wyoming. See 
Index Herb. Pt II (Regn. Veg. 86) 1972. 

*Junus Dali, see Forest Research Institute, Bogor. 

*Kadim bin Abdul 

erroneously mentioned as Kadir bin Abdul 
(see Suppl. Cycl. Fl. Mai. I, 5, 1958). 

*Kadim bin Tassim 

collected with J. Sinclair in A'^. Borneo (June- 
July 1957); with J. Carrick & I. Enoch in 
Sarawak: Bako National Park (Apr. 18-May 11, 
1959). In Malaya with Mahmud bin Awang in 
Pahang (Mar.-Apr. 1959), with Mohd Nur in 
Johore (June-July 7959), with Mrs Allen (Aug. 
1959), with R. K. Dentan in Perak (Feb. and Apr. 
7962). He must have collected in several other 

Collections. Herb. Sing.; dupl. elsewhere, 
a.o. Herb. Edinb. 

Kadir bin Abdul, rightly: Kadim bin Abdul. 

Kaempfer, Engelbert 

Biographical data. Add: Cl. Nissen: 'Die 
botanische Buchillustration' 1951, p. 89-91; K. 
Meier-Lemgo: 'Die Briefe Engelbert Kaempfers 
(1651-1716)' (Abh. math.-naturw. Kl. Akad. Wiss. 
Mainz no 6, 1965, p. 265-314). 

*Kairo, A. 

was attached to the Forestry School, Bulolo, 
East New Guinea, in the 1960s. 

Collections. From E. New Guinea, in Herb. 
Lae; dupl. elsewhere, also in Herb. Edinb. 

Kalkman, Cornelis 

Add: He left New Guinea in Sept. 1959, and 
was appointed staff member of the Rijksherbarium, 
Leyden; Ph.D. Leydcn (1965); since September 
1972 Director, and Professor at Leyden University. 

Dimorphanthera kalkmanii Sleum. was named 
after him. 

Itinerary & Collecting localities. 
Add: 1957. W. New Guinea: Kelobo I., Strait Sele 
and Rufei, Sorong (Nov.); S. New Guinea: 
Merauke, Lampu Satu (Dec.).— 1958. W. New 
Guinea: surroundings of Manokwari: near 
Teminabuan (Beriat), Wersar (Apr. 18-May 5); 


P. Adi (May 31-June 1); Fak-Fak (June 21-24); 
Kebar Valley. — 1959. S. New Guinea: Muju 
region (Jibi, Opka, Biakatem, Awemko, Umkubun 
Iwurmot) (Mar. 8-31 ; numbered in the BW series). 
Centr. New Guinea: Star Mountains Expedition 
(leader: L. D. Brongersma, sec there) (Apr. 16- 
Aug. 30)M Sibil Valley, Orion Mts (Ok Tenma, 
Tenmasigin, Along), Junction of Bon and Minam 
R., Mt Antares. — 1966. Centr. New Guinea: 
Expedition to Doma Peaks and Hindenburg 
Range^, with Vink, Gillison, and Frodin: South. 
Highlands Distr., Tari Subdistr. (Tigibi, Ibiwara, 
Mt Kerewa, Mt Ne, Mt Ambua, Lei R.) (June 5- 
Aug. 30); W. Sepik Distr., Telefomin Subdistr., 
Hindenburg Range (Mt Amdutakin, and between 
Feramin and Telefomin) (Sept. 11-18). 

Collections. Add: BWhoj 6201-6500, 8501- 
8550; Star Mts Exp., own numbers 4001-4560 
(partly with Tissing, Nicolas, Van Zanten, De 
Wilde), Exp. Doma Peaks etc. private nos 4601- 
5324. In total about 2060 nos. Private numbers: 1st 
set in Leyden, 2nd at Lae, 3rd Canberra, etc. 

Literature. (1) Cf. reports in Tijd. K. N. A. 
G. 76, 1959, p. 98-103 w. map, 211-213, 323-326; 
C. Kalkman: 'Description of vegetation types in 
the Star Mts Region, West New Guinea' (Nova 
Guinea, Bot. no 15, 1963, p. lAl-iei). 

(2) C. Kalkman & W. Vink: 'Botanical 
Exploration of the Doma Peaks and adjacent 
mountains in the Southern Highlands District, 
Terr, of Papua, Austr. New Guinea' (Report for 
1966, Neth. Found. Advanc. Trop. Res. WOTRO, 
p. 50-52, 2 photogr.); 'Botanical Exploration in 
the Doma Peaks, New Guinea' (Blumea 18, 1970, 
p. 87-135, fig. 1 (map)-5, photogr. 1-25). 

Biographical data. Curr. Vitae in Thesis 
Leyden 1965. 

Kalshoven, Louis George Edmund 

(d. 1970, Netherlands). 

Biographical data. Vakbl. Biol. 50, 1970, 
p. 144. 

*Kaniis, see Forest Dept North Borneo. 

"'Kandilis, see Forest Dept North Borneo. 

Kanehira, Ryozo 

Literature. (2) Add: For sketch map of the 
route in the Arfak Mts and itinerary see W. Vink 
in Nova Guinea, Bot. 22, 1965, p. 488. 

(4) Add: P. van Royen: 'Publication Dates of 
and Index to Taxa in Reports by R. Kanehira on 
Plants collected during the Kanehira-Hatusima 
1940 Collection in New Guinea' (Bot. Mag. Tokyo 
86, 1973, p. 143-154). 

*Kanis, Andries 

(1934, Alphen a/d Rijn, Netherlands; x), 
botanist, educated at Leyden University, where he 
took his Ph.D. in I968i. In August 1965 he entered 
the service of the Dutch Ministry of Foreign Affairs 
and was stationed as a botanist in the Forest 
Department of Sabah (N. Borneo), Malaysia; from 
Jan. 1966 acting as Head of the Sandakan Her- 
barium; in Oct. 1966 he returned to Leyden. 
April 1969 he left for Canberra, Australia, for the 
time being on a contract in the employ of C.S.I.R.O. 



Cyclopaedia of Collectors — Supplement II 


En route he visited Ceylon in connection with the 
Colombo Plan. 

Collecting localities. Malaysia. 1965. 
Sabah (N. Borneo): with W. Meijer to Mt Kina- 
balu (Sept. 2-28).— 1966. Sabah: Tour along the 
W. coast, visiting regions of Kota Belud, Kudat, 
Jesselton, Tenom, Keningau, and Ranau (Feb. 3- 
Mar. 10); to the E. coast with D. Hou in the 
regions of Lahad Datu Kunak, Semporna, Tawau 
(Apr. 28-May 18); Mt Kinabalu tour with D. Hou 
(May 25-June 11); Sarawak: field work in Ulu 
Bakong, Baram (one month), continued in the 
Lambir Hills, Miri (2 months), Nyabau and 
Segan Forest Reserves, Bintulu (2 months); Mt 
Matang, Semengoh Arboretum, and Bako National 
Park (Aug.). When returning to Leyden in 
October he paid short visits to Brunei, Kuching 
(Sarawak), Singapore (Bt Timah), and Kepong in 
the Malay Peninsula (8 days making trips to Bt 
Lagong, Telok F. R., Batu Caves, Eraser's Hill), 
and Bangkok. — 1969. Papua: Owen Stanley Range, 
also Mt Wadimana (E. summit Simpson massif). — 
1970. Papua: Astrolabe Range and near Efogi on 
the Kokoda Trail (4 weeks after ANZAAS 

Collections. Herb Sandakan: c. 250 nos, 
collected in the SAN series, often together with 
Sabahan collectors; dupl. mainly in Kew and 
Leyden. A small, private series, numbered B. 1-22, 
contains mosses from Sabah (in Herb. Leyden) 
and Ochnaceae from Sarawak (Herb. Leyden and 
Kuching). Papua plants will be in Canberra. 

Literature. (1) On 'A revision of the 
Ochnaceae of the Indo-Pacific area' (Blumea 16, 
1968, p. 1-83). 

Biographical data. Curriculum vitae in 
his thesis (preprint of Blumea, see Liter. 1), Leyden 

*Kapis bin Sisiron, see Forest Dept North Borneo- 

*Kardin, M. Kosim 

graduate biology student of the Bandung 
Technological Institute, assisted J. R. Flenley 
and R. J. Morley (see those) in their investigations 
in Indonesia in 1972. 

^Kassim, Rajab 
collected at Gua Batu, Selangor (Aug. 17, 1962). 

Kastalsky, G. 

(fl. 1826-29) was mentioned as a collector in 
Luzon (cf. Cycl. Fl. Mai. [, 1, 1950). Physician 
and Naturalist on the sloop 'Moller' (Capt. 
Stanjukovitch) which accompanied the 'Sen- 
jawin' (sec K. H. Mertens), 1826-29. 

Collections. Herh. Leningrad (cf. Hult^;n 
in Bot. Not. 1940, p- 300); some in Kcw; Herh. 
Oxford: from Manila (Luzon) and the Pacific 
islands (with Herb. Fielding). 

*Katayama, Tadao C. 

n932, Konosu. Saitama, Japan; x). agriculturist, 
educated at Kagoshima University (1952-56), and 
taking his Master's and Doctor's course at Kyoto 
University (1956 61); Dr. of Agriculture (1963); 
Researcher, National Institute of Cicnclics (1961- 
67); A.ssistant Professor of Agriculture at Kago- 
shima University from 1967. He was sent to the 


Philippines and New Guinea in January 1961, and 
to Borneo and Java in March 1963 for collecting 
wild and cultivated rice, with grants from the 
Rockefeller Foundation. 

Collecting localities.^ 196L Philippines. 
Mindanao: Zamboanga; \V. Negros: Hinigaran; 
Palawan: Bangcodo; Sorsogon: Juban; Legaspi; 
Mindanao: Muswan. W. Irian: Opeco. Baad and 
Wajaw in Koembe area; Merauke; Territory of 
Papua: Madang. — 1963. W. Java: Bogor, Depok, 
Bidaratjina. Kalimantan (Borneo): Bandjermasin, 
Belandean, Handilmanarap; N. Borneo: Bandau, 
Langlon, Malalap, Keningau, Pangi; Brunei: 
Lemunin, Labi; Sarawak: Limbang, Hiller and 
Ulu near Simanggang, Miri, Seriau, Lundu, Bunsit 

Collections. Numerous strains of 7 species 
were collected for his institute; possibly part of 
them were made into herbarium specimens. 

Literature. (1) T. C. Katayama: 'Wild 
Oryza species of the Philippines, New Guinea, 
Borneo, and Java (Seiken Ziho, Rep. Kihara Inst. 
Biol. Res. Yokohama no 15, Dec. 1963, p. 35-46; 
list of collecting localities I.e. p. 36-39); 'Scientific 
Report of the Rice-Collection-Trip to the Philip- 
pines, New Guinea, Borneo and Java' (Mem. Fac. 
Agr. Kagoshima Univ. 6, 1968, p. 89-134; 
extensive diaries on p. 112-115, 130-133); 'Botan- 
ical Studies in the genus Oryza' I (ibid. 7, 1969, 
p. 89-117, diagr., fig. 1-163, incl. map). 

*Kate, ten 

collected some interesting mountain plants in 
Central Timor on March 13, 1891, viz Drosera, 
Vaccinium timorense, Habenaria, and Diuris 
fryana Ridl., on G. Lakan or Reedtz Thott Mt, 
c. 2000 m alt. ?Exclusively in Herb. Leyden 
(labels written by Boerlage). 

♦Katik, Paul 

(1940, Kemeng, Terr, of New Guinea; x), field 
assistant in Division of Botany at Lae. He partici- 
pated in numerous explorations in various parts 
of New Guinea, including in the N. district of 
Papua at loma (2 weeks, 11966), and in New 
Ireland (with M.J. E. Coode). 

Collections. Herb. Lae; dupl. also in Herh. 

*Kaul, Robert Bruce 

(1935, Faribault, Minnesota, U.S.A.; x). Ph.D. 
Univ. of Minnesota (1964); Associate Professor of 
Botany, Univ. of Nebraska in Lincoln, U.S.A.; 
his special interest is aquatic monocotyledons. 

He collected in Sabah together with Abbe, and Mi.uiR, as a lield research assistant 
to Prof. Ahhf, with an NSF grant. Hesidcs the 
tropics, Europe, the Near East, and Japan was 

He published on the morphology, ontogeny, 
phylogcny, evolution, etc., specially in connection 
with aquatic plants. 

IriNiKARV. 1961. India (Sept. 21-Oct. 6); 
Ceylon: Colonibo (Oct. 23); Malaya: Singapore, 
Cameron Highlands, I'enang, Eraser's Hill, Kuala 
Lumpur (Oct. 31 Jan. 21, 1962).— 1962. Sabah: 
Mt Kinabalu (I'cb. 2 23); Philippines: Luzon: 
Baguio, Quezon Prov. (Mar. 3 17). 



Flora Malesiana 

[ser. I, vol. 81 

Collections. Fagaceae for Prof. Abbe and 
material for his thesis. In Herb. Minnesota, nos 
1706-2124, mainly aquatic monocotyledons; dupl. 
in Herb. Lae and Herb. Edinb. 

♦Keck, V. 

Collections. Herb. Gottingen: from S. 
America (1878-79) and Singapore (1878). Euro- 
pean plants in Geneva. See Index Herb. Pt II 
(Regn. Veg. 86) 1972. 


Keir, Walter 

Add: He was in England in August 1700 (cf. 
letter to Petiver in Sloane MSS 4063, folio 41). 

Keith, Henry George 

Later for F.A.O. at Benghazi, Libya. 

♦Kellers, Henry Carsten 

(1874-x). Philippine plants (1929) in U.S. Nat. 
Herb. Wash. See Index Herb. Pt II (Regn. Veg. 
86) 1972. 

♦KeUman, Martyn Carden 

(1941, Barbados, West Indies; x), B.Sc. McGill 
University (1962), M.S. University of Wisconsin 
(1963), Ph.D. Australian National University 
(1968); Assistant Professor of Geography in Simon 
Eraser University, Burnaby, British Columbia. He 
joined an Arctic expedition of McGill University 
to Axel Herberg Island (1961), visited the Philip- 
pines twice (see below),^ and British Honduras in 

Itinerary. Philippines (Jan.-Mar. 1964, Nov. 
1964-Nov. 1965),^ collecting in Coronon Valley 
and Gumate Distr. (Barrio Tungkalan) on the E. 
slopes of Mt Apo, Mindanao.^ — E. New Guinea 
under D. Walker. 

Collections. Herb. Canberra, numbered in 
the ANU series of the University, viz 1500-1729 
(complete), 1730-1999 (partial), 3000-3442 (partial). 

Literature. (1) Author of: 'Some environ- 
mental components, of Shifting Cultivation in 
Mindanao' (J. Trop. Geogr. 28, 1969, p. 40-56); 
'Secondary Plant Succession in Tropical Montane 
Mindanao' (Canberra, Austr. Univ. Press, 1970). 


♦Kelly, Raymond C. 

of the Department of Anthropology, University 
of Michigan, Ann Arbor, U.S.A. He collected 
some 30 plants in Mt Hagen area, Territory of 
Papua and New Guinea, probably in 1968. 

Collections. In Herb. Leyden, received via 
Van Zanten, Groningen. 

♦Keng, Hsuan 

(1923, Kiangsu, China; x), B.Agr. Nat. Chung- 
Chen Univ. Kiangsu, China (1947); M.Sc. Nat. 
Taiwan Univ. Taiwan, China (1949); Ph.D. Univ. of 
Calif., Berkeley, U.S.A. (1959). Teaching Assistant 
Dept of Bot., Nat. Taiwan Univ., 1950-53; 
Lecturer in Botany, Dept of Bot., Nat. Taiwan 
Univ., 1953-57; Research Fellow, Morris Ar- 
boretum, Univ. of Pennsylvania, 1959-60; Lecturer 
in Botany, Univ. of Malaya, Singapore, and later 
Univ. of Singapore, 1960-66; Senior Lecturer in 
Botany, Univ. of Singapore, 1966 onwards. 

Author of numerous taxonomical and morpho- 
logical papers and some books. At present working 
on Labiatae and Theaceae. 

Collecting localities. On and around 
Singapore; with J. A. R. Anderson in Sarawak 
(NfV. Borneo), Baram Distr. (July 1961); UNESCO 
Limestone Expedition (Ulu Kelantan, Malaya) 
(Aug. 1962); early 7965 exploration trip to the 
Langkawi Is, with some students. 

Collections. In Herb. Dept Bot. Univ. Sing.; 
some dupl. in Herb. Sing. 

♦KEP nos 

a series of the Forest Research Institute at 
Kepong, Malaya. As it reached the six figures by 
1965, a new series with the prefix FRI was begun 
in 1966; the first number is FRI (KX)1. The junior 
staff continued (at least still in 1967) with the KEP 

Kern, Johannes Hendrikus 

Hon. Dr at Nijmegen University (1970). 

Biographical data. Gorteria 4, 1968, p. 
69-72; Blumea 17, 1969, p. 1-3; Fl. Mai. Bull, no 
25, 1971, p. 1910 (Hon. Dr); Jaarb. Versl. & 
Meded. Kon. Ned. Bot. Ver. over 1970, 1971, 
p. 95-99. 

Kerr, Arthur Francis George 

Itinerary. Add: 1932. From Mergui by 
steamer to Rangoon, arriving June 8; leaving (9) 
for Penang (arrival 12); proceeding to Pulau 
Langkawi (16), collecting there and leaving (21); 
back at Penang (22), sailing for Europe on the 23rd. 

Biographical data. Add: Blumea 11, 1963 
p. 427-493, portr., itin., bibliogr. 

♦Kerr, Harlan 

collected a Cyperacea in Cotabato Prov., 
Mindanao, Nov. 1956. 

Kerr, William 

Data at Kew make it clear that he visited Java 
in 750-^, not in 1805. 

Keysser, Christian 

Collections. Add: Duplicates bought by 
the Herb. Brit. Mus. (1923). 



Cyclopaedia of Collectors — Supplement II 


Kiah bin Hadji Mohamed Salleh 

Add: In 1937 he collected with Moysey; from 
1949-55 with J. Sinclair; in 1957 with Chew 
Wee Lek. At the end of the latter year he retired 
after 3 7 It. years of service. 

Collections. Herb. Sing.; dupl. Herb. 
Edinb. etc. 

*Kickx f., Jean 

(1803-1864). In the Index Herb. Pt II (Regn. 
Veg. 86) 1972, he is cited as the collector of plants 
in Borneo, Java, and Sumatra. To my knowledge 
he did not visit these islands, and the plants will 
have been in his private herbarium which is 
preserved in Herb. Caen. 

*Kilang, see Forest Dept North Borneo. 

•Kingston, James 

District Agricultural Officer, who has collected 
herbarium specimens in various parts of E. New 
Guinea, including New Britain in the 1950s. 

In 1968 living at Buderim, Queensland, Australia. 

Collections. Herb. Lae, >143 nos. 

*Kinted, V., see Forest Dept North Borneo. 

*KjelInian, Frans Reinhold 

(1846, Bromo, Torso, Sweden; 1907, Uppsala, 
Sweden) studied botany at Uppsala University 
(Ph.D. 1872); from 1883 Professorof Botany there. 
He made botanical explorations in Spitsbergen 
(1872-73), and Nova Zembla (1875), and ac- 
companied the Vega Expedition (see under 

He is the author of numerous publications, 
mainly dealing with Algae. 

Kjellmania J. Reinke has been named after him. 

He collected phanerogams near Alaska,^ and 
he might have collected others in the Malaysian 
region in the 'Vega' (1878-79). The botanical 
collections of that expedition are in Herb. Uppsala 
and dupl. in Stockholm. 

Literature. (1) Cf. E. Hult^n in Bot. Not. 
1940, p. 303. 

Biographical data. Svensk Bot. Tidskr. 
1907, p. 276-285; Svensk Vet. Ak. Arsbok f. 1908, 
p. 279-300; Ymer 1907, p. 286-292; Krok, Bibl. 
Bot. Suec. 1918, p. 349-353, incl. bibliogr. 

K.K.S.S., stands for Kostermans, Kuswata, 
Soegeng, Soepadmo. 


a series of the Phytochemical Survey of the 
Federation of Malaya, was started in 1957 by 
Millard. It has been resumed by Dr J. Carrick, 
Field Pharmacologist, with number 2332. Some 
confusion has arisen because of the three numbers 
attached to the old scries (KL, PC'S, and a herb, 
number). The first number, KL, is the only one 
that is continuous. 

•Kleckham, F. 

collected a few plants on Mt Strong {Papua), 
end August 1963; Herb. Lae. 

Kleinhoonte, Anthonia 
(d. 1960. Netherlands). 


Biographical data. Vakbl. Biol. 41, 1961, 
p. 3-6. 

Kleiweg de Zwaan, Johannes Pieter 
(d. 1971, Blaricum, N.H., Netherlands). 

Kloss, Cecil Boden 

Collections. Add: The Christmas I. plants 
(pres. 1923) were probably made by Dyak collectors 
((/. Raffles Mus. Bull. 18, 1947). 


*Kochummen, Kizhakkedathu Mathai 

(1931, Kerala, India; x), B.Sc. from the Uni- 
versity of Travancore in 1951, came to Malaya in 
Dec. 1952; from 1953 onwards in the Botanical 
Division of the Forest Research Institute at 
Kepong, since Dec. 1957 Assistant Forest Botanist. 
Author of several papers, mostly on forest trees; 
co-author of the Tree Flora of Malaya. He left 
the Forest Institute to attend the 1967/68 Diploma 
course in Taxonomy at Edinburgh, under the 
Colombo Plan. Mid- 1968 he passed his examin- 

Collections. He collected some thousands 
of plants throughout Malaya; material in Herb. 
Kepong, numbered in one of the successive scries 
(KF-P and FRI); dupl. distributed from Kepong. 
After his return from Europe he resumed collecting. 

•Kock, Father Cornclis Josephus Lconardus 

(1907, Breda, Netherlands; x) took his doctoral 
examination in social geography at Utrecht 



Flora Malesiana 

[ser. I, vol. 8^ 

University (1937); teacher in secondary schools in 
the Netherlands (at present Canisius College, 
Nijmegen); Professor of Geography at Djok- 
jakarta, Indonesia, in the 1950s. 

Collecting localities. Java: Djakarta 
(June 1955), Bedugiil {Bali, June 1957), G. Gedeh 
(June 1953), Kali Urang (Aug. and Oct. 1958, 


Jan. 1959), Niisa Bandng) and G. Kidui (Feb. 
1958), Ungaran (June 1961), Djokja (Nov. 1958, 
Jan. and Feb. 1959). Without known year at: 
Angke, Bogor, Tjilintjing R., Rawa Tembaga, 
Depok, Bandung, G. Prahau, Lembang, Taman- 
djaja, and Muntilan. Also in Sumatra. 

Collections. In private collection; mostly 
common (vascular) plants. In all about 9000 
specimens (260 families, c. 3000 species) of which 
c. 4000 from Java and a few from Sumatra and 
Bali. Some identified at Leyden, many at Bogor. 

*K0ie, Mogens Engell 

(1911, Copenhagen, Denmark; x) was educated 
at the University of Copenhagen (Mag. Sc. botany 
1936); Assistant at the Botanical Laboratory of 
the University of Copenhagen 1938; Dr. phil. 1951 ; 
head of the Ecological Department 1956; Pro- 
fessor in Botanical Ecology 1967.^ Member of the 
'3rd Danish Central Asian Expedition', 1948-49;^ 
UNESCO Science Research specialist in Peru, 
1956-57; from Jan. 8-Apr. 28, 1962, he partici- 
pated in the Danish 'Noona Dan Expedition' (see 
there for Itinerary, Collections, etc.). 

Literature. (1) List of publications inDansk 
Bot. Ark. 21, 1963, p. 188-189. 

(2) K0IE & Rechinger: 'Symbolae Afganicae' 
(in Kongl. Dansk. Vidsk. Selsk. Skr.). 

Biographical data. Dansk Bot. Ark. 21, 
1963, p. 33-34. 

*Kokawa, Shohei 
(b. 1927) of Osaka City University spent 5 


months (Oct. y96S-Mar. 1969) in many areas of 
the state Sabah (N. Borneo), with Ando and Hotta 
(see there). Special attention was given to Tawau 
Hills and Silabukan in the south and to Kinabalu 
and Trus Madi on the west coast. 

Kokkelink, M. Ch. 

Add: Author of 'Wij vochten in het bos' (with 
biography and portr.). 

*Kondo, Yoshio 

(1910, Kaeleku, Isl. of Maui, Hawaiian Is; x) 
malacologist, educated at the University of 
Hawaii (B.A. 1940, M.A. 1947), taking his Ph.D. 
at Harvard University in 1955. Previous to his 
university studies he was employed as a ship 
steward, electrician, fisherman, diesel engineer, and 
finally as an Assistant in Malacology in the B. P. 
Bishop Museum. He accompanied expeditions in 
Micronesia (1936), Fiji (1938), in the Trust 
Territories (1949, 1952), and the Philippine-Hawaii 
Expedition (1957), mainly collecting land mollusca. 

Itinerary. 1957. Philippine-Hawaii Expe- 
dition: leaving Honolulu (Jan. 7), arriving at 
Manila (9), and via Zamboanga (staying 23-26) to 
the Sulu Archipelago: Siasi I., at Laminusa etc. 
(26-Feb. 7); Jolo I. (8-9); Tawitawi I. and Sibutu 
I. (10-22); Taganak (22-24, 27); Sandakan (A^. 
Borneo, 25-27); Cagayan Sulu (27-28); Palawan: 
Balabac (Mar. 1-5), Ursula I. (6), Puerto Princesa 
(7, 9), Babuing (8); Negros Oriental (11-21); 
Bohol (23-24); Cebu (21-23); Calamangan, off 
Leyte (24-25); Leyte: Polompon (25) and nearby 
Gigantanga I. (26-27); Capul (or Tapul) I. off 
Samar (27); Naranjo I.: Medio and Rosa (28); 
Destacado I. (28-29); Masbate, Deagan I. and 
Homoraon islet (30); Ticao: Bagababoy (31); 
Burias: Tinalisayan (Apr. 1); Mindoro: Puerto 
Galera and Golo I. (3); back at Manila (4). 

Collections. No botanical field note books 
were kept; in most instances the material was 
flown back to the Philippine Nat. Museum for 
numbering and drying there. In Herb. Manila and 
in B. P. Bish. Mus. Herb. Honolulu. 

*Koningen, H. 

(1940, Amstelveen, Netherlands; x). When being 
in the army as a soldier, he made a small collection 
in New Guinea. Later gardener at Amstelveen. 

Collections. 7 plants at Remu near Sorong 
(Jan. 1961) and near Ifar (Cyclops Mts, July 1961), 
W. New Guinea. In Herb. Leyden. 

*KooIs, J. P., see J. J. P. E. de Wilde. 

*Kooy, Father Cornells Willem 

(1917, Hoorn, N.H., Holland; x), a Roman 
Catholic Missionary educated at Steyl and 
Tetringen (1942-46), from 1949 stationed in 
Indonesian Timor, stationed successively at 
Manufui (Beboki) from 1949-50, at Manumean 
(Beboki) from 1951-56, at Soe (S. Central Timor) 
from 1956-62, and at Nikiniki (S. Central Timor) 
from 1962 onwards. 

Collecting localities. Timor, from 1964 
onwards: in the districts of Kupang, Amanuban, 
Amanatun, Molo, Miomafo, Insane, and North 



Cyclopaedia of Collectors — Supplement II 


Collections. He collected 560 numbers of 
plants in the southern central part of the island 
during 1964-69 for Herb. Leyden; in 1972 some 
150 new numbers were received. 

Koppel, Cornelis van de 

(d. 1963 or 1964, ?Bussum, Netherlands). 

Biographical data. Add: Versi. Treub 
Mij. Mei 1963-Mei 1964. 

Koster, Christiaan 

He left New Guinea October 1962; in 1969 he 
tried to get an employment in Surinam. 

Collecting localities. Add: 1957. Kebar 
Valley, P. Adi.— 195^. W. part of Arfak Plain 
(Sidei, Wariki) (Julv-Aug.), Kebar Valley (Oct.- 
Nov.).— 1959. Sarmi Hinterland (Bodei) (Jan.- 
Feb.).— 1960.1 starting for Mt Krabo (Arfak Mts) 
from Maruni (Oct. 31), collecting mainly on the 
crest (900-1060 m) (Nov. 1-8); leaving Mt Krabo 
and returning to Manokwari (9). — 1961. Arfak 
Plain (Woseggi-Masni) (Feb.), Japen I. (Soember- 
baba, Woda, Aisao) (June-July), Andai area near 
Manokwari (Oemboei, Andai, Oesiabi, Beri-creek, 
Maroeni, Warmare) (Nov. 10-Dec. 17).— 1962. 
Wandammen Peninsula (Wondiwoi Mts, Wasior) 
(Feb. 24-Mar. 16), Arfak Mts with Chr. Ver- 
steegh (see there) (May 16-28),* Andai area 
(Warmare) (June 20-26), Meos Noem (July 31- 
Aug. 11). — From 1957-62 in the surroundings of 

Literature. (1) Ch. Koster: 'Verslag van 
een tournee naar het Krabo-gebergte' (Manokwari, 
3 pp., typewritten); cf. also W. Vink in Nova 
Guinea, Bot. 22, 1965, p. 492, fig. 6. 

(2) Cf. W. Vink in Nova Guinea, Bot. 22, 1965, 
p. 493. 

Kostermans, Andre Joseph Guillaume Henri 

Add: In summer 1962 in Europe, in 1967 in 
Europe and the U.S.A., in 1971-73 he worked in 
Europe, viz at Leyden, Aarhus, Kew, Paris, and 

Collecting localities. 1952. Suppi. Cycl. 
Fl. Mai. L 5, p. (59)b 3rd line from top. Peak of 
Balikpapan, add: G. Beratus. — 1958. W. Java 
and Prinseneiland; Bali: Bedugul (June). — 1960. 
W. Java: Tandjung Lajar, P. Peutjang, and 
Udjung Kulon (July); Tjibodas (Sept. 13).— 1961. 
Leaving Tandjong Priok (Mar. 27) in the 'Samu- 
dera' with 2 'mantri's' of the Herbarium, 3 students 
(Sujarta, PRfJANTO, Wirawan), to Siimhawa:* 
touching Bandjcrmasin (30), Surabaya (Apr. 1- , 
paying a visit to the Bot. Garden at Purwodadi); 
arrival at Sumbawa Bcsar (7), Badas; making 
collections at: 8 km E of Sumbawa Besar (8-9); 
Scmongkat, 17 km W of Sumbawa Bcsar (1 1-12); 
Batudulang on the slope of the Batulantch, W of 
Scmongkat (12-18), from c. 700-summit (1200 m); 
trail to Punih. 600 m (19); between Scmongkat and 
Batudulang (20-21); in mountain forest from 
Batudulang to Brangbossang, 700 -900 m alt. 
(22 25); bivouac at Brangbossang. 800 900 m alt. 
(26). and from 800 1200 m (26 28); N of Batu- 
dulang, Mt Samperolat, 500-800 m (Apr. 29 May 
6) in drier forest; to Pusu on the NW. side of the 
mountains, 500 m alt. (May 7-11); back to 
Fiatudulang and .Scmongkat (12), 200-500 m alt.; 


Apr. 27 they were joined by 2 Russians, Dr A. 
Fedorov (Curator of the Leningrad Herb.) and 
Dr Kabanov of the Forestry Service, accompanied 
by KusvvATA (see there) of Herb. Bog. SiV. Java: 
near (Lake) Raw ah Danau (July 19-20). In the 
same year for the 2nd time to Sumbawa:* leaving 
in the 'Samudera' (Oct. 14) with several others, 
including Mr Ma.mad, assistant of the Bot. Div. 
of the Forest Research Institute at Bogor; Oct. 19- 
Nov. 8 was spent in the field, collecting in the same 
places as earlier in the year in order to get fertile 
material of those species which were not in flower 
or fruit the first time. iV. Java, Bantam: lower W. 
slope of Mt Hondje (Dec. 28-Jan. 2, 7962).— 1963. 
Kalimantan (former Dutch Borneo): leaving Bogor 
(Sept. 12) for a tour to the Berau region (NE.): 
from Tandjong Redeb to Long Lanuk on the Sg. 
Kelai; base bivouac at the foot of Mt Njapa 
(limestone), where he fell ill and returned to Tg 
Redeb, in the meantime Mr Sim leading field 
operations; later in lowland forest N of Teluk 
Bajur and in swamp forest along Sg. Sega (col- 
lecting over 700 field nos); on Nov. 14 back at 
Bogor. — 1964. W.Java: Mt Pangrango (Feb.); Mt 
Pajung and Udjung Kulon Reserve (Dec). — 1965. 
Flores. With N. Wirawan, leaving Bogor (Mar. 
20); Waingapu (Siimba), Ende, to Mborong (S. 
Manggarai); base camp at Kisol, exploring the 
Mt Ndeki complex; Wae Rana and explorations 
southwards to the coast; Ruteng, exploring the 
forest up to 2400 m; swamp forest W. of Mborong; 
Wirawan to Rana (= lake) Mese near Sita 
mountains above Ruteng, and further to the 
North, to Reo; June 2 back at Bogor. In Dec. a 
short trip to Central Sumatra was made in com- 
pany of Mr Jahn (a German forester) and a team 
of Japanese foresters. — 1966. E.xpedition to W. 
Irian (for collecting plants causing permanent or 
temporary sterility in women): SiV. Celebes: 
Bantimurung near Makassar (July 20); Irian: 
surroundings and in the valleys of Sukarnapura 
( ^ Hollandia) (July 25-Aug. 15); vicinity of Lake 
Sentani and Sukarnapura (Aug. 16); Sukarnapura 
(17); Dosai, E of Sukarnapura (18); Abepura near 
Sukarnapura (19-20), Sukarnapura (21); Dosai 
(22-25); Baliem valley, surroundings of Wamena 
and Wellesey up to 2500 m (26-Sept. 3); P. Biak: 
Bosnik (Sept. 9); Parieri (9-13); P. Morotai (16). 

Collections. Add: The Russians (guests of 
the Council for Sciences) who joined the Sumbawa 
expedition (I96I) would get a set of the collection, 
nos 18001-18865; the expedition was financed by 
the Council. In Flores (1965) 1300 nos were 
collected (about 6 dupl. sets); in [V. Irian & Biak 
(1966) ] 000 no.i. 

The numbers IIO(X)-12000 were given to living 
plants; in Hort. Bog. 

Lithrature. (2) Add: For sketch map of his 
route in the Arfak Mts, and comments on labels 
etc., sec W. Vink, Nova Guinea, Bot. 22, 1965, p 
488-489. fig. 3. 

(4) Cf. Fl. Mai. Bull, no 16, 1961, p. 802; A. J 
G. H. Kostermans: 'Notes on the Vegetation of 
W. Sumbawa (Indonesia)' (Symp. Icol. Res 
Humid Trop. Vcgct. Kuching 1963; UNE-.SCO Sc 
Coop. Off. SE. Asia 1965, p. 15-25. 2 fig.). 

*Koyama, Hiroshigc 
(b. 1937), a member of the Kyoto University 



Flora Malesiana 

[ser. I, vol. 8» 

Biological Expedition to Thailand and Malaya, 
1967, who paid special attention to cytotaxonoray. 

Itinerary. See sub K. Iwatsuki. 

Collections. 1st set in Kyoto Herb.; 2nd set 
in For. Herb. Bangkok. 

*Kraepelin, Emil 

(1856, Neustrelitz, Germany; 1926, Munich, 
Germany) studied medicine in Wiirzburg and 
Leipzig, specializing in neurology and psychiatry, 
and working in Leipzig, Munich, and Dresden. 
In 1886 Professor of psychiatry at Dorpat Uni- 
versity, from 1891-1903 at Heidelberg, and from 
1903 in Munich. 

Having a large interest in nature, he often 
accompanied his elder brother Karl (see there) 
on tours in the alps, the Mediterranean, etc., and 
in 1904 to the tropics (see below). Whereas Karl 
made the zoological collections, Emil took care of 
plant collecting, at least in 1904. 

Though his publications exclusively deal with 
his discipline, he wrote a diary (letters to his wife) 
during his tour to the East, which was preserved.^ 

Itinerary.^ 1904. By ship (Dec. 23, 1903) 
from Genoa to Ceylon (staying Jan. 10-Feb. 4), 
paying a visit to S. India (Jan. 29-Feb. 1); pro- 
ceeding to Singapore (staying Feb. 13-19): 
Botanic Garden, Bt Timah (15), Serangoon R.; 
West & Central Java {¥ eh. 21-Apr. 1): Buitenzorg 
Botanic Garden, Tjampea (Mar. 11), Borobudur 
(17), G. Papandajan (22), Situ Bagendit (23), 
Tjibodas (24-29), Tjibadak and G. Gede (26), G. 
Pangrango (27), Tjiborrom (= Tjibeureum) (29), 
Buitenzorg (30) ; sailing Apr. 2, making short stays 
in Penang and Colombo, reaching Genoa on Apr. 

Collections. Herb. Munich (see Liter. 1, p. 

Literature. (1) H. Weidner: 'Karl und Emil 
Kraepelin's Studienreise nach Indien und Java im 
Jahre 1904. Aus dem Brieftagebuch von Emil 
Kraepelin ausgewahlt, eingeleitet und mit Anmer- 
kungen versehen' (Abh. Verh. Naturw. Ver. Hamb. 
16, 1972, p. 19-72, 12 Abb., 2 Karten). 

Biographical data. Cf. Liter. 1, p. 21-25, 
72, w. portr. 

♦Kraepelin, Karl 

(1848, Neustrelitz, Germany; 1915, ?Hamburg, 
Germany) studied mathematics and natural 
sciences at Gottingen and Leipsic; from 1873-78 
teacher in natural history in Leipsic, and subse- 
quently in Hamburg; from 1887 professor and 
since 1889 Director of the Natural History 
Museum in Hamburg. In later years his mainly 
zoological interest turned to ecology. He exten- 
sively travelled, often accompanied by his younger 
brother Emil (see there). 

Author of numerous zoological publications.^ 

Itinerary & Collections. See under E. 
Kraepelin. Botanical collections were presented 
to the Museum in Hamburg. As far as can be 
surmized from his brother's diary, botanical 
collections in the East (1904) were made by Emil. 

Literature. (1) Also of 'Naturstudien in 
fernen Zonen. Plaudereien in der Dammerstunde' 
(Leipzig & Berlin 1911). It contains little infor- 
mation on the voyage to the East. 

Biographical data. Abh. Verh. Naturw. 
Ver. Hamb. 16, 1972, p. 21-25, w. portr. &bibliogr. 
(p. 71-72); I.e. p. 72. 

Kreulen, D. J. W. 

= Dirk Jan Willem. 


= Krischtofovich, African Nikolaevic (1885- 

""Kurata, Shigeo 

of the Insectivorous Plants Society, Dept of 
Biology, Nippon Dental College, Fujimi, Tokyo, 
Japan, made two expeditions to the Philippines in 
search for Nepenthes.^ 

Collecting localities. Philippines: 1965. 
Mindanao (Surigao del Sur: E. slope Mt Legaspi, 
Camp David, and near Carrascal Bay) and Luzon 
(Mountain Prov., 3 weeks in Aug.). — 1967-68. 
Mindoro (Dec.-Feb.): Mt Kidayap. 

Literature. (1) S. Kurata & M. Toyoshima: 
'Philippine species of Nepenthes' (Gard. Bull. 
Sing. 26, 1972, p. 155-158). 

*Kuripin, see Forest Dept North Borneo. 

Kurz, Wilhelm Sulpiz 

Biographical data. Add: Ind. Forester 4, 
1878, p. 1-4. 

*Kuswata Kartawinata, E. 

(1936, Java, Indonesia; x), B.Sc. of the Tjiawi 
Academy of Biology (1959); since then Assistant 
Botanist of the Herbarium Bogoriense. He left for 
the University of Hawaii in February 1965 to 
major in ecology, taking his Ph.D. In 1973 
temporarily working for BIOTROP at Bogor. 

Collecting localities. 1958. Bali and 
Java (see sub Soepadmo). — 1959. W. Ceram: 
village Gemba, Kairatu (June 1-21); Waiselang, 
Kairatu (22-26); Amboina: Mt Salahutu (July 5- 
20), and elsewhere (July 22-Oct. 8); W. Java: 
Bogor and Tjibodas. — 1960. Together with 
Kostermans at Udjungkulon (Aug. 13-21, Oct. 
13-14).^— 1961. W. Sumbawa: Batulante (May 1- 
4), Olat Sekampil (13), Semongkat Atas (13-14), 
Batugong, W of Sumbawa Besar (23); \V. Java: 
Udjungkulon (June 15-17). 

Collections. Partly made together with 
Soepadmo (see there), and nos 1-242 ( W. Ceram), 
243-305 (Ambon), 85-290 (tV. Sumbawa), etc.; 
in Herb. Bog. 

Literature. (1)E. Kuswata Kartawinata: 
'Notes on the Vegetation of Peutjang Island (SW. 
Java) (Symp. Ecol. Res. Humid Trop. Veget. 
Kuching 1963; UNESCO Sc. Coop. Off. SE. 
Asia 1965, p. 26-30). 

Kuyper, H. P. 

= Hendrik Pieter. 

Kuyper, Jan 

(d. 1969, Groningen, Netherlands). 
Biographical data. Jaarb. Versl. & Meded. 
Kon. Ned. Bot. Ver. over 1970, 1971, p. 5. 

*Kwok, S. 

and Philippine collectors, collected 844 plants; 




Cyclopaedia of Collectors — Supplement II 


Herb. Lingn. Un. Canton. See Index Herb. Pt II 
(Regn. Veg. 86) 1972. 

*Laan, A. C. van der 

former employee of the Rijksherbarium at 
Leyden, collected some plants at Port Swettenham, 
Malaya, in April 1962; 8 nos in Herb. Leyden. 

LAE nos 

After no 50.000 the series letters NGF (New- 
Guinea Forces, later New Guinea Forests) were 
changed into LAE. 

Lahaie, Felix de 

Collections. Add: Some plants labelled 
'Java', virtually came from Mauritius. 

*Lajangah, see Forest Dept North Borneo. 

Lam, Herman Johannes 

retired in 1962. 

Collecting localities. Add: 1962. 
Malaya: during the UNESCO limestone Expedition 
in Ulu Kelantan (July 24-Aug. 18). 

Biographical data. Blumea Suppl. 4, 
1958 (Jubilee vol.), p. 1-16, bibliogr., portr. 

* Lancaster, C. R. 

an Army officer; he collected some plants in the 
Cameron Highlands, summit of G. Batu Brinchang 
at 6666 ft, Malaya (Mar. 1-3, 1958); Herb. Leyden. 

*Landingin, G., see Philippine National Herbar- 

Landon, Sylvanus 

settled at Banjar = Bandjermasin, SE. Borneo 
in 1700 (cf. letter from James Cuninghame to 
Petiver, Sloane MSS 3321, folio 52, British 
Museum). The collecting locality *Doole' (see sub 
S. Landman in Cycl. Fl. Mai. I, 1, 1950, and sub 
S. Landon in Suppl. Cycl. ibid. I, 5, 1958, is 
rightly the oldest Portuguese settlement in Solar, 
Lesser Sunda Islands. 

Lanc-PooIe, Charles Edward 

fd. 1970, Sydney, Australia). 

Biographical data. Commonwealth For. 
Rev. 50, 1971, no 143, 3 pp., portr. 

'Lang, William Henry 

(1874, Groombridge, Sussex, England; 1960, 
Milnthorpe, Westmorland, England), M.S., B.Sc, 
D.Sc. (1900) at Glasgow, Senior Assistant to 
Professor of Botany, subsequently Lecturer, Bot. 
Dcpt. University of Glasgow, 1890-1908; Pro- 
fessor of Cryptogamic Botany, Manchester 
University, 1908-1940. One of his interests was 

He paid a short visit to Malaya when he went 
to Ceylon, early in this century (1900). 

Collections. In Herb. Glasgow: small 
collection of ferns from Malaya: dupl. presumably 
in Manchester fProf. R. E. Holttum /./. June 24, 
1958), but according to Prof. D. \\. Valentine no 
ferns in Manchester, but a lot of bryophyte 

Biographk:al data. DoRhLiR, Botanikcr- 
Adrcssbuch 2nd cd. 1 W2, p. 110; C. W. Wardlaw 
contributed to Nat. Diet. Biogr. (1962). 

*Langdon, Raymond Forbes Newton 

(1916-x) collected in A'^m' Guinea and Queens- 
land (1940-56); Herb. Univ. Brisbane. Fungi only? 

*Larsen, Egon 

(1928, Denmark; accidentally killed in vehicle 
accident April 1969, Celebes (Sulawesi), Indonesia) 
took his M.Sc. in Forestry in Copenhagen; 
Scientific Officer, Forest Research Institute, New 
Zealand Forest Service, 1954-62); in the Seeds 
Section of the Forest Research Institute, Canberra, 
Australia, 1962-69. 


He undertook seed collection expeditions to 
U.S.A. (1956), Mexico (1961), Timor and Flores 
(1968). and finally to Celebes (1969), especially for 
collecting Eucalyptus. 

He was a member of the FAO Panel of Experts 
on I-orcst Gene Resources. 

Collecting localities. 196H. Lesser Sunda 
Islands (July-Aug.): Timor, both Portuguese and 
Indonesian parts: NW. part of Portui^ucsc Timor, 
collecting along the road from Dili to Maubissc 
( luriscai) and from Dili t(^ Ermcra (both from sca- 
Icvcl to 5(K)0 ft); by road from Dili to Kuixing; by 
air to /'lores, travelling by car from Lndch to 
I.arantuka.— /%9. SiV. Celebes. South and west 
of Palopo (Apr.). 

Collections. Eucalypts in I'Rl Herb., 




Flora Malesiana 

[ser. I, vol. 8^ 

Canberra. In Celebes also specimens of Casuarina 
sumatrana were collected. 

(c/. Suppi. Cycl. Fl. Mai. I, 5, 1958) was not an 
employee of the Forestry Service, but a collector 
in the BW series. 

Latif, Soetan Mahmoed 

Literature. (1) Add: 'Two orchids of 
Indonesia' (Amer. Orchid Soc. Bull. 35, 1966, p. 
907-909, 2 fig.). 

♦Laverock, W. S. 

M.A., B.Sc, a former Keeper of the Department 
of Botany in the Public Museums, Liverpool, 

Plants from Malaya have been in Herb. Mus. 
Liver p. (see Handbook and Guide to the Herbarium 
Collections in the Public Museums Liverpool 1935, 
p. 71); destroyed during World War II. 

Laycock, John 

(d. Dec. 3, 1960, aged 73). 

Layosa, Helen S. 

Add: Instructor of Agronomy; in the 1960s 
Assistant Professor at Los Baiios College of 
Agriculture. She took her Ph.D. in Florida. 

*Lazaro, J., see Forestry Bureau, Manila. 

*Lazo, P. P., see Philippine National Herbarium. 

*Leach, G. 

of the Div. of Botany, Lae, New Guinea, 
collected with R. Pullen (see there) on Mt Suckling 
in 7972. 

Leano, Forestry Bureau, Manila, read: E. C. 

Lecomte, Henri 

Biographical data. Add: Adansonia 2, 
1962, p. 147-159, portr. 

Ledoux, Jacques Alphonse 

Add: (1880, Liverpool, England; Apr. 1, 1961, 
Johore Bahru, Malaya). From his youth he had an 
interest in plants. He volunteered for service in the 
war in S. Africa, remaining in that country until 
1906, when he went to Malaya. From tin-mining 
in Johore he soon turned to rubber-planting in the 
Kota Tinggi District, where he remained for the 
rest of his life. 

Biographical data. Gard. Bull. Sing. 18, 
1961, p. 328-329, portr. 

*Ledyard, E. M. 

Plants from the Philippines (1909-10) in Herb. 
Ann Arbor. (300). 

*Lee Koh Hat, see Forest Dept North Borneo. 

*Lee Wai Chin, see Miss Geh Siew Yin. 

Leefers, Cornelus Leonardus 

(1909, Soerabaja, Java; 1962, Ibadan, Nigeria), 
educated at Wageningen Agricultural College. Tea 


planter (1938-41); in the Dutch East Indian Army 
(1941-42), P.O.W. in Japan, 1942-45; in the army 
again, 1945-47; from 1948 in the Forestry Service 
of the Dutch East Indies and Indonesia. From 
1952-60 Conservator of Forests (Opperhoutvester) 
and Inspector of the Service in West New Guinea, 
finally stationed in Biak. Repatriated Sept. 1960. 

*Leeuwen, Arie van 

(1915, Meerkerk, Z.H., Netherlands; x), edu- 
cated at Wageningen Agricultural College. In 
September 1956 he went to Dutch New Guinea 
(Hollandia) as Inspector of the Veterinary Depart- 

Collecting localities. 1957. W. New 
Guinea: Kebar Valley (Oct. 25); Kabarai, IVaigeo(u) 
(Oct.-Nov.); Kofiau I., W of New Guinea (Nov. 
22).— 1958. W. New Guinea: Baliem Valley (Mar.). 

Collections. He collected on behalf of the 
Forestry Service; Herb. Manokwari and Leyden. 

*Leigh, P. 

collected sugar-cane varieties along the banks 
of the Laloki River near Port Moresby (c/. Cane 
Growers' Quart. Bull. Brisbane 15, 1951, p. 42). 

*Lele(a)n, Yakas 

(1937, Aregena, Terr, of New Guinea; x), a 
good field assistant at Lae, who collected with 
others in the NGF series. 

*Lennox, Colin Gordon 

Honolulu, was sponsored by the Hawaiian 
Sugar Planters' Association for the collecting of 
seeds of wild relatives of the sugar cane in 1937.^ 
He was accompanied by C. E. Pemberton (see 
there for collecting localities etc.). No herbarium 
collections were made. 

Literature. (1) C. G. Lennox: 'Sugar cane 
collecting in New Guinea during 1937' (Hawaiian 
Plant. Rec. 42, 1938, p. 235-246). 

Biographical data. Amer. Men of Science 

Leschenault de la Tour, Louis Theodore 

In the Library of the Rijksherbarium, Leyden, 
there is a bound copy of a handwritten MS titled 
"Liste des Plantes de I'lsle de Java observees par 
M. Lechenault de Latour". It contains descriptions 
and notes in French of 693 species and is dated 
"Samarang, 5.7bre, 1806". This is obviously not an 
original MS but a copy made by a clerk of "Het 
Marineschool te Semarang" which did also work 
for Norona, and where the plates of Louis 
Deschamps were copied. 

Several of his papers were translated in English 
for Colonel Colin Mackenzie by his Dutch 
translator L. R. Burke. It forms parts of a whole 
body of MSS which were collected in Java during 
the first couple of years of the British Adminis- 
tration (now in India Office Library, London). In 
one volume Burke has inscribed in his own hand: 
"Some of the Leschenault Memoirs were trans- 
lated into Dutch for Mr M. Engelhart by whom 
Mr Leschenault was employed" (Mackenzie 
collection, private, vol. 37, 'Collection on the 
Natural History of Java', 296 folios of which 194 
by Leschenault; Dr John Bastin /./. July 1970). 



Cyclopaedia of Collectors — Supplement II 


Itinerary. Additional to the data mentioned 
in Cycl. Fl. Mai I, 1, 1950, p. 321a the following 
can be derived from the MS mentioned above: 
Many plants from the vicinity of Soerabaja, 
Grissee (together with the Resident 'Karel Van 
Naersen'), Madoera I. (nos 203-400, Sumenep and 
islet 'Mimdang'hine' before the coast), Banju- 
wangi (nos 401-574), climbing Mt Idjen with 
Commander 'Vikerman' (nos 575-663), Westcoast 
of Bali (rto^ 575-581). He visited also Mt 'Onorang' 
near Semarang (Mt Ungaran). The final few 
numbers 688-693 are from the environs of 
"Melam" which I cannot locate. He was obviously 
acquainted with, or even met, Horsfield. 

Literature. (11) Add: 'Dessins de plantes 
de Java' (fol., cart. ; 55 plants of which the numbers 
correspond with those of his journal and her- 

Lesson, Pierre Adolphe 

Literature. (4) Add: For particulars on the 
botanical drawings see Cl. Nissen, 'Die botanische 
Buchillustration' 1951, vol. 2, p. 53 sub Dumont 

Biographical data. In Bibl. Austr. Entom. 
1775-1930 (R. Zool. Soc. N.S.W. 1932, p. 199). 

*Levinge, Harry Corbyn 

(1831-1896). His large private herbarium is in 
Dublin, but plants from Malava are recorded at 
Kew Herb. See Index Herb. Pt II (Regn. Veg. 86) 
1972. Probably the Malayan specimens have been 
presented to him. 

♦Lewis, S. H., see S. R. J. White. 

*Loatoc, A. 

Philippine plants in Herb. Ann Arbor. 
Index Herb. Pt II (Regn. Veg. 86) 1972. 


*Loeters, Johannes Jacobus 

(1913, Didam, Netherlands; x), Roman- 
Catholic priest. Following his theological edu- 
cation (at Teteringen) he attended the Agricultural 
School (Koloniale Landbouwschool) at Deventer. 
Since 1945 stationed at Nita, Flores (Lesser Sunda 
Is.), teaching at an agricultural school. 


•Lichanco, J. 

Philippine plants in Herb. Ann Arbor. 
Index Herb. Pt II (Regn. Veg. 86) 1972. 


♦Lichtenthaler, Dr H. K. 

botanist of the Bio-Organic Chemistry Group, 
1557 Life Sciences Building, Berkeley (Calif.), 
planned to visit Bogor in Jan. J964, to collect 
orchids, ferns, Araceae, etc. on his return to 
Heidelberg, Germany, by way of the Far East. 

Lillez (or Lilies?), Leonardo 

mentioned under Lii.les in Cycl. Fl. Mai. I, 
1, 1950, evidently collected several plants for the 
Bureau of Agriculture at Manila in March 1909. 

Lindenbaum, see Glasse, S. H. 


Add the initials H. W. 

Also plants from NE. New Guinea (New Britain). 
See Index Herb. Pt II (Regn. Veg. 86) 1972. 

Llanos, Antonio 

When revisiting Madrid in 1958, Dr E. Qu\- 
suMBiNfi studied his collection and selected 
Icctotypcs for Herb. Manila. 

•Llobrera, C. 

Philippine plants in Herb. Ithaca. Sec Index 
Herb. Pt II (Kcgn. Veg. «6) 1972. 



Collecting localities. Lesser Sunda Is. 
since 1952, mainly in Flores (Badjawa, Maomere, 
Nita, Ruteng, Hokeng, Geli Mutu (30 March 1962) 
Larantuka, etc., but also on some nearby islets, 
viz: P. Paloe (N of Flores), P. Lomblen (Apr. 
1962), P. Solor (May 3, 1962). 

Collections. He owns a private herbarium 
of c. 1000 numbers (with 1 or 2 dupl.) of Flores 
plants (end 1967: nos 1001-2230). For private use. 
Some dupl. were sent to Leyden Herb. 

The numbers S 50 and S 57 leg. Schouten 
(July 4, 796/). 

L5rzing, J. A. 

Literature. (6) Reference must read: 
'Sumatra' instead of 'Sumatra Post'. 

*Loh Hoy-Shing 

Assistant of the Forest Research Institute, 
Kepong, Malaya. 

Collecting localities. 1969. Malaya. 
Selangor, Gading F. R. with Chan.— /97(^. Gua 
Pcningal, Pahang (July). 

Collections. In Herb. FRI Kepong. 

Loher, August 

Collecting localities. Add: Plant speci- 
mens collected in April 1906 arc often labelled 
'Rio Anipalit' in Ri/al Province. This locality 
refers lo a large stream Dampalit (the ragal()g 
name for river) on the N. side of Mt Makiling in 
Laguna prov. (not Ri/al). 



Flora Malesiana 

[ser. I, vol. 8* 

*Lomibao, Beningno A. 

collected for and with William L. Stern (see 

Loreja, Miss Milagros 

Add: In 1966 in Stanford, U.S.A., taking 
courses in botany. 

♦Lorenzo, H., see Boswezen Nieuw Guinea. 

Loria, Lamberto 

Itinerary. Cycl. Fl. Mai. I, 1, 1950, p. 330a, 
1st line from above, after 'Goodenough I.' add: 
(Jan. 13-18). 

Lorrain, W. B. 

Collections. Add: Also ferns in Herb. 
Edinb., a.o. with Thomas Moore's fern herbarium, 
Gourlei herbarium, and Kew dupl. According to 
Index Herb. Pt II (Regn. Veg. 86) 1972, a Herb. 
Lorraine is in Inst. Bot. Nancy. It is not known to 
me whether one and the same person is involved. 

Lotsy, Johannes Paulus 

Biographical data. Add: in Dictionary of 
Scientific Biography, New York, vol. 8, 1973. 

Low, Henry Stuart, see Stuart Henry Low, 

Low, Hugh 

Correct: Trained to take over the Clapton 
Nursery, but then sent to the Orient for the 
collection of plants and seeds for his father 
(Hugh Low Sr)'s nursery at Clapton. He reached 
Singapore on Nov. 23, 1844, disembarked on the 
25th. From 18-22 Dec. he met his brother Stuart 
there, and together they had one day's collecting.' 

On Jan. 1845 he arrived at Kuching, Sarawak, 
made friends with Rajah James Brooke, and with 
few breaks remained in Borneo for 28 years. 
Appointed Colonial Secretary of Labuan under 
Brooke in 1847, arrived there 1848, and stayed 
until 1876 (with leave in England in 1861 and 
1876). Resident of Perak 1877-88 (introducing the 
first Hevea rubber to Malaya, and later to Singa- 
pore and Java) ; sent to Brunei to negotiate treaty 
with the sultan in 1889; retired to England 1889. 

Itinerary. Add: He stayed for 30 months in 
Sarawak, being back in England in Oct. 75-^7. 

Collections. He also made entomological 
collections, among which some remarkable 
butterfly collections. 

Literature. (3) Add: Cf. H. Low, Sarawak 
etc. (London 1848), p. 350-406. 

(7) C. F. Cowan: 'Sir Hugh Low, G.C.M.G. 
(1824-1905)' (J. Soc. Bibhogr. Nat. Hist. 4, 1968, 
p. 327-343). 

Biographical data. J. Soc. Bibliogr. Nat. 
Hist. 4, 1968, p. 327-343, w. portr. from 1848. 

Low, James 

Collections. Also dupl. in Herb. Edinb., 
incl. ferns with Herb. Greville. 

Low, Stuart Henry 

correct for Henry Stuart Low {cf. J. Soc. 
Bibliogr. Nat. Hist. 4, 1968, p. 328). 2nd son of 
Hugh Low (1793-1863), Nurseryman of Clapton 


Nursery, was privately educated; joined the 
merchant navy. Returned to Clapton in 1845, 
assisting his father and taking over the Nursery on 
his death in 1863. 

He grew many plants sent home from Borneo 
by brother Hugh. The Dendrobium microglaphys 
Rchb. /. cited from Borneo (Stuart Low 1865), 
mentioned in the Cyclopaedia, was certainly 
grown at Clapton and not collected by himself. 

Biographical data. C. F. Cowan in lift. 
Feb. 12, 1968. 

*Luning, Bjorn Erik Gustav 

(1929, Stockholm, Sweden; x). Organic Chemist 
(Dr hab. 1960) at the University at Stockholm; 
since 1967 Associate Professor; specialist on the 
alkaloids of Orchidaceae. He made expeditions to 
Panama (1964) and New Guinea (1967).^ 

Itinerary. 1967. Papua and New Guinea 
(June-July): Lae area, Wau area, Wagau area, 
Wampit area, Mt Hagen area. 

Collections. Large amounts of one sjjecies 
for chemical investigation, and reference plants to 
the live collection of the Institute of Organic 
Chemistry at Stockholm University in connection 
with alkaloid screening of species (c. 250 N.G. 
species). At least some plants in Herb. Lae. 

Literature. (1) Articles on the journeys in 
Orchid Review. Author of many phytochemical 
papers on orchids. 

*Luerssen, C. 

probably Christian (1843-1916), the well-known 
pteridologist. In the Index Herb. Pt II (Regn. Veg. 
86) 1972 he is recorded as a collector in Banka in 
Polynesia (r. 1880). Luerssen was professor of 
botany at Koningsbergen from 1888-1910, and he 
is the author of some papers on Polynesian ferns, 
but to my knowledge he did not collect in Poly- 
nesia or Banka himself. 

*Lugtmeyer, H. W., see Boswezen Nieuw Guinea. 
The right initials, instead of C. (in Suppl. Cycl. 
Fl. Mai. I, 5, 1958). 

*Luitjes, J., see Boswezen Nieuw Guinea. 

*Lunau, see Forest Dept North Borneo. 

Lundquist, Erik 

Biographical data. Add: In St. Bergman: 
'Vildar och Paradisfoglar' 1950, p. 41-47, portr. 
following p. 32. 

McAdam, James Bannister 
(d. early 1959, Brisbane, Queensl., Australia). 

McClure, Floyd Alonzo 

(rightly: 'Mc' and not 'Mac', b. 1897; d. 1970, 
Bethesda, Maryland, U.S.A.). Well-known expert 
on bamboos. 

Biographical data. Amer. Men of Science, 
Phys. & Biol. Sc. Uth ed. 1966; Taxon 20, 1971, 
p. 777-784, portr., bibliogr. 

MacGillivray, John 

Literature. (2) Add: M. Bassett: 'Behind 
the Picture. H.M.S. Rattlesnake's Australia and 
New Guinea Cruise' (Oxford Univ. Press Mel- 
bourne 1966, with 2 maps). 



Cyclopaedia of Collectors — Supplement II 


Biographical data. Add: Portr. in H.M. 
Whittell, 'The Literature of Austral. Birds' 1954, 
pi. 26, opposite p. 116, and see pt 2, I.e., p. 465-466. 

MacGregor, Richard Crittenden 
Collections. At the end add: Herb. Edinb. 

Mcintosh, Donald Henry 

Add: Author of 'The effect of man on the 
forests of the Highlands of Eastern New Guinea' 
(Symp. Humid Trop. Veget. Unesco, Goroka 
1960, 1963, p. 123-126). 

*MacKaughan, Dr Harold 

Head Department of Linguistics of Hawaii 
University. He evidently visited the Philippines, 
where he collected plants of Lanao del Sur 
{Mindanao) with Maranao names in July 1964. 
They were identified by D. Mendoza.^ 

Literature. (1) H. P. MacKaughan &B. A. 
Macaraya: 'Maranao plant names' (Oceanic. 
Linguist. 4, 1965, 112 pp.). 

McKee, H. S. 

Add: joined the C.N.R.S. at Noumea, New 
Caledonia (Fl. Mai. Bull, no 20, 1965, p. 1241). 

♦MacLagan, Dr 

Collections. In Herb. Edinb. from India and 
Malay Peninsula {cf. Index Coll. Edinb. Herb. 
1970, p. 106). 

MacTier, W. F. 

His ferns were collected in 1865 (see Cycl. FI. 
Mai. I, 1, 1950 under Mactier). 

♦McVeagh, David 

Technical Officer with the Forest Department, 
collected few spjecimens in the NGF series in 
Waitape area, Centr. Division, E. New Guinea, in 
1959. See also E. C. McVeagh. Herb. Lae. 

•McVeagh, E. C. 

? Same as above and initials possibly wrongly 
read. Said to have collected in the NGF series 
near Oriomo River, E. New Guinea, June 1960. 

•MacVean, Dr Donald N. 

one of the staff members working under Dr D. 
Walker (see there). He collects plants in E. New 
Guinea. In May 1967 he had just left for Mt 
Wilhelm;' he also collected on Mt Giluwe, S. 
Highlands, and in 7969 he accompanied Dr B. O. 
VAN Zanten (see there) to E. New Guinea (June- 

Collections. In Herb. Univ. Canberra, 
numbered in the ANU scries (Australian National 

Literature. (1) L. K. Wade &. D. N. 
McVean: 'Mt Wilhclm studies I. The alpine and 
subalpinc vegetation' (Res. School Pacif. Stud. 
Dept. Biogcogr. &. Gcomorph. Publ. BG/I, 1969, 
325 pp.) (Austr. Nat. Univ. Press). 

*Madani, L., see Forest Dept North Borneo. 

'Madulid, Domingo Allado 

(1946, I>aoag, Ilocos Norte, Philippines; x), 
U.S. (Botany) University of the Philippines (1967), 


M.S. (Botany) University of Sto. Tomas (1971); 
Museum Researcher in the Philippine National 
Herbarium, Manila. 

Collecting localities. Philippines. 1968. 
Luzon: Mt Data National Park, Mt Province 
(Nov.; regeneration of oaks). — 1969. E. Mindoro: 
Mt Iglit Game Refuge and Bird Sanctuary (May- 
June).— 7970. S. Samar: Mt Sohoton (Mar.-Apr.; 
ecology), with H. G. Gutierrez. — 7977. Luzon: 
Bo. Guina-ang, Bontoc, Mt Prov. (ethnobot. coll. 
with Dr L. Reid; Apr.). S. Samar: Guinma- 
ayuhan, Balangiga (May). Luzon: Ilocos region 
(I. Norte & La Union), for sand-binding grasses. 


Collections. //(T^.A/a/i/Zfl (Phil. Nat. Herb.), 
c. 2000 flow, plants, 200 ferns, 200 Algae, Furtgi 
and mosses. The Samar duplicates will be distrib- 
uted to foreign herbaria. 

•Mahmud, S. 

collected with Samsuri in 1970-71, in Selangor 
and Perak. 

•Mahmud bin Awang, see sub Kadim bin Tassim. 

*Mahuze, B., see Boswezen Nieuw Guinea. 

*Maidin, S. Mohammed, sec Forest Dept North 

Mangold, Rudolf Paul 

Add: lor some annotations on the collecting 
localities in the Arfak Mts sec W. Vink in Nova 
Guinea, Bot. 22, 1965, p. 491-492, lig. 5. 



Flora Malesiana 

[ser. I, vol. 81 

•Manner, Harley Ichiro 

(1941, Honolulu, Hawaii, U.S.A.; x), geog- 
rapher, B.A. University of Hawaii, currently 
(1968) working on M.A. and Ph.D. in Geography 
at Hawaii University. He accompanied Dr John 
M. Street (see there) as Research Assistant to 
New Guinea, to gather data on morphology, 
structure, and floristics of the savannah and forest 
vegetation. Publications will follow. 

Collecting localities. Terr, of New 
Guinea (Mar. 7967-Feb. 1968): in the vicinity of 
Kupeng, Kompiai, Koinambe and Tsuenkai of the 
Jimi Valley, Western Highlands District. 

Collections. Nos 1-599 in Herb. Leyden, 
Lae, and in Hawaii Geogr. Dept. 

♦Marchette, Nyven John 

(1928, Murphys, California, U.S.A.; x) was 
educated in zoology at the University of California 
(A.B. 1950, M.A. 1953) and took his Ph.D. in 
Bacteriology at the University of Utah in 1960. 
Assistant Research Microbiologist, Hooper Foun- 
dation, University of California Medical Center, 
San Francisco (Cal.) from 1961 onwards. In 
1962-64 associated with the Institute for Medical 
Research Kuala Lumpur, working on the ecology 
of ricketsial diseases in Malaya; from 1965-67 
associated with the Dept of Bacteriology, Fac. of 
Medicine, Univ. of Malaya, Kuala Lumpur, for 
the investigation of the ecology of arthropod-borne 
viruses of Malaya. On two occasions in 1963 the 
Institute for Medical Research sponsored ex- 
peditions which spent a total of about 3 weeks on 
G. Jerai in Kedah {Malay Peninsula), viz. in March 
and September. 

Collecting localities. Malaya. 1963. G. 
Jerai (Aug.-Sept.),^ Pulau Aur.^ — 1964. Taman 

Collections. The specimens were not col- 
lected with any herbarium, some are still in the 
Faculty of Medicine at Kuala Lumpur. 

Literature. (1) N. J. Marchette: 'Notes on 
the flora of Gunong Jerai (Kedah Peak)' (Mai. 
Nat. J. 19, 1965, p. 129-137; with History of 
Exploration and List of species collected). 

F. L. Dunn: 'Notes on the ferns of Gunong 
Jerai' (Mai. Nat. J. 20, 1967, p. 131-138). 

(2) N. J. Marchette: 'Notes on some Flowering 
Plants of Pulau Aur' (Mai. Nat. J. 18, 1964, p. 50- 
59; annotated list of records). 

*Martin, Adam Menzies 

(1940, Lanark, Scotland; x), Foreman, Royal 
Botanic Garden, Edinburgh, accompanied B. L. 
Burtt (see there) on a trip to Sarawak in 1967. 
He was mainly in charge of the collecting of living 
Gesneriaceae and Zingiberaceae. 

Collections. Herb. Edinb. 

*Martin, Ricarte, see Philippine National Her- 

•Mason, L. Maurcie 

(1912, Fincham, King's Lynn, England; x), 
farmer and amateur gardener, who made extensive 
travels for the collecting of living plants, mostly 
orchids, in the tropics.* 

Collecting localities. 1952. Malaya. — 
1954. NE. New Guinea (Mand. Territory) : at Lae, 


Nondugl, and Mt Hagen. — 1958. Central Malaya: 
mostly around Kedah Peak; in spring N. Borneo: 
Segama, Mt Silon, Darvel Bay, Lahad Datu; 
Sarawak: Kuching area. 

Collections. Living plants only, of which 
specimens have been offered to Kew Gardens, 
mostly orchids and Begonias. Occasionally 
material is preserved in spirits; Herb. Kew. 

Literature. (1) L. M. Mason: 'The Travels 
of a Plant collector' (J. R. Hort. Soc. 94, 1969. 
p. 481-489). 

Matie, Miss 

Instructor of Botany in Manila. Possibly 
identical with the collector Oeny Mati (see Suppl. 
Cycl. Fl. Mai. I, 5, 1958). 

Matthew, Charles Geekie 

Collections. At the end add: Herb. Edinb. 


Unknown collector of that name. MS list of 
plants from Surinam and Amboina in Oxford 
Univ. (in MS 174). 

Mayr, Ernst (Walter) 

Add: In 1953 besides appointed Alex. Agassiz 
Professor of Zoology in the Museum of Com- 
parative Zoology, Harvard University. 

Literature. (1) Add: For annotations on his 
itinerary in the Arfak Mts see W. Vink in Nova 
Guinea, Bot. 22, 1965, p. 487, fig. 5. 

Biographical data. In H. M. Whittell: 
'The Literature of Austral. Birds' 1954, pt 2, p. 

Mearns, Edgar Alexander 

Collecting localities. Add: Sulu Archi- 
pelago: Dammi I. (Jan. 4, 1906), Tawitawi I., 
Tataan I. (Jan. 8, 1906). 

M.E.D.P., see M. E. D. Poore; not to be used in 


•Meer, Jikke Henriette Hermina van der 

(1893, Amsterdam, Netherlands; x), phyto- 
pathologist, who studied and took her Ph.D. 
(1925) with honours in the University of Amster- 
dam. From 1919-32 she was on the staff of the 
Phytopathological Institute of the Agricultural 
College at Wageningen. As a fellow of the Inter- 
national Education board she made a study tour 
to the Dutch East Indies from Nov. 1927-Jan. 
1929, till August 1928 stationed at the Deli Experi- 
ment Station at Medan. From 1932-36 on half- 
pay; in 1936 married to Dr Ir J. H. J. Van De 
Laar. At present living at Spaubeek (L.), Nether- 

Collecting localities. Nov. 7927-July 
7929 in Sumatra East Coast: garden Deli Experi- 
ment Station; at Brastagi; Aug.-Sept. 1928 in 
Java, where she visited all experiment stations, 
collecting on G. Gedeh-Pangrango, etc. 

Collections. 162 nos of material for phyto- 
pathological study in Herb. Wageningen, col- 
lected for educational purposes of students of 
tropical agriculture. Also material from Java 
preserved in formalin. 



Cyclopaedia of Collectors — Supplement II 


Meer, Pieter van der 

Collections. Add: Occasionally he pre- 
sented specimens to Herb. Leyden. 

Meer Mohr, Johannes Carolus van der 
(d. 1969, Barchem, G., Netherlands). 

*Megata, Moritane 

(1917, Kyoto, Japan; 1945, Ley te I., Philippines) 
graduated in horticulture. Faculty of Agriculture, 
Kyoto University, 1942. He served as a lieutenant 
during World War II. His main interest was 

Collections. He collected some plants in 
Java, Celebes (Manado), and at ? Tjimoelon in 
1940. In Herb. Kyoto Fac. Sci. living plants from 
Java too. 

Mendoza, Demetrio R. 

Add: he is engaged in the preparation of a Flora 
of Banaue (N. Luzon) with R. M. del Rosario, 
with whom he also wrote 'Philippine aquatic 
flowering plants and ferns' (Manila 1967). 

In 1969 he retired as Curator of the Philippine 
Herbarium. He is now connected with the faculty 
of the Dept of Forestry, Araneta University. 

Collecting localities. Add: 1961. Luzon: 
since June 1 in cutting and logging areas of the 
Nasipit Lumber Co. in Agusan Prov., and on the 
mountains in Agusan. — 1964. Palawan (Apr.- 
June). — 1965. Quezon Prov., Luzon (June 1-28). — 

1967. Luzon: Dalton Pass in Nueva Vizcaya Prov. 
(Mar. 3-18); Banaue,^ Ifugao Prov. (May 11- 
June 11) with R. M. del Rosario. 

Collections. Add: Herb. Kew. 
Literature. (1) Cf. Fl. Mai. Bull, no 22, 

1968, p. 1538-1539. 

Biographical data. Portr. in Philip. Orch. 
Rev. 7, 1959, p. 35. 

*Meiior, v., see Philippine National Herbarium. 

Menzies, Archibald 

Collections. See this Supplement under D. 
Nelson. Herbarium specimens often signed 
'A.M.' on the back of small sheets. 

Literature. (2) Add: For itinerary in Alaska 
sec Bot. Not. 1940, p. 297. 

Biographical data. Last line (Suppl. 
Cycl. FI. Mai. I, 5, 1958) add: and in Occ. Pap. 
Calif. Ac. .Sc. no 20, 1943, facing p. 15; Trans. R. 
Soc. New Zeal. 88, 1960, p. 63-64. 

Meiriil, Elmer Drew 

Rightly: d. at Forest Hills. 

Collecting localities. Add: The locality 
'.Separacion' or '.Separation Point', not found on 
atlases, is situated in E. Palawan at about one 
third of its length (from the south); it is a small 
cape in Island Bay. 

Biographical data. Add: J. Wash. Ac. 
Sc. 46. 1956, p. 267-268; Philip. J. .Sc. 85, 1956. 
p. 181-188, portr.; Taxon 6, 1957. p. 89-101, 
portr.; Biogr. Mem. Nat. Ac. Sc. U.S. 32. 1958. 
p. 273-333, portr., chronol. bibliogr.; Dedication 
in F-l. Mai. I, 6, 1972, p. (7) (16), portr. 

Merritt, Mclvin Ixroy 

Collections. Add: Philippine dupl., coll. 
with H, M. Curran, in Herb. Ldinb. 

Mertens, Karl Heinrich 

Literature. (1) Add; 
Not. 1940, p. 299. 

For Alaska see Bot. 

*Merton, (L.) F. (H.) 

collected at Gua Batu, Selangor (Dec. 22, 7960), 
in Pahang, Malaya, with Chew Wee Lek, 7962, 
and with Poore, 


His plants from E. Java were collected about 

*Metzner, Joachim K. M. 

(1940, Berlin, Germany; x) took his 'Diplom 
Volkswirt' (economist) at Heidelberg University, 
Sudasien Institut (1966), and subsequently read 
at Cornell University (1967/68). He spent 15 
months in 1969-70 in Portuguese Timor to study 
the ethnography and vegetation, collecting voucher 
specimens. He worked in the Baucau-Viqueque 
area. A Ph.D. thesis is forthcoming. 

Collections. Herb. Leyden: 270 nos; private 
dupl. collection. 

Meyer, Adolf Bernhard 

Itinerary. Add: 1871-72. Philippines.*" 
Literature. (4a) 'On the Population' cf. 

Meyer in Peterm. Geogr. Mitt. 20, 1874, p. 17-19, 

tbls; and I.e. p. 19-22. 

Meyer, Robert 

cannot possibly have been born in 1891; in ? 

Meijer, Willem 

Add to Suppl. Cycl. Fl. Mai. I, 5, 1958: From 
May 1959 in the employ of the Forest Dept North 
Borneo, stationed at Sandakan. In 1962/63 he 
made a round-the-world trip, visiting herbaria; 
in 1966 with European leave again. In 1968 
appointed Visiting Associate (later Associate) 
Professor in the University of Kentucky, Lexing- 
ton, U.S.A. 

Author of a botanical guide to Mt Kinabalu 
(Sandakan 1963, mimeogr.). papers on Diptero- 
carpaceae, etc. His investigations in Sumatra 
resulted in some other papers.'* 

Collecting localities. Add: 1959. From 
May 20 in N. Borneo (Sahah):^" regions of Tawau 
(end of June-end of July), Scpilok F. R., W. coast 
regions (Padas Gorge and surroundings of 
Kinabalu); lower slopes of Kinabalu near Ranau 
and above Bundu Tahan and Tenompok (Oct. 27- 
Nov. 9); Darvcl Bay near Lahad Datu (Nov. 15- 
iy)._1960-6L Several times in the Kinabalu 
area.'* and short trip to Sarawak and Brunei 
(1 96 1). —1962. r:arly in the year (Feb.) a 3-wccks 
tour on Mt Kinabalu with Prof. E. Aiiiii-, R. Kaul, 
and Mr & Mrs Bogi i: of Minnesota (sec those). 
Malaya: E. Johore to Kuala Trcngganu (May 27- 
Junc 5). with K. M. Koc hummin. N. Borneo: Mt 
Andrassy and near Ouoin Hill. Tawau; Mt 
'Irusmadi in Ikaufort Dislr.— 1963. Mt Kinabalu 
and Mcsilau R. (July) with Sieumer a.o.; in 
various other places in Sabah. — 1964. In Jan. 
accompanying H. f". Moore in Sabah. — 1966. 



Meijer's collectors 

Flora Malesiana 

[ser. I, vol. 8^ 

Collections. Add: The Borneo plants 
numbered in the SAN series; those from Malaya 
in the KEP series; also dupl. in Herb. Edinb. 

Literature. (7) Add: 'Rottans van Nunukan' 
(Pengg. Alam 36, 1957, p. 51-64, 5 fig.). 

(10) Add: W. Meijer: 'On the Flora of Mount 
Sago near Pajakumbuh, Central Sumatra' (Pengg. 
Alam 38, 1959, p. 2-11, fig. 1-3; ibid. 40, 1961, 
p. 3-13, 3 fig.). 


(14) W. Meijer: 'Notes on the Ecology of Sawah 
Plants in the region of Pajakumbuh' (Res. Pamphl. 
no 2 Bot. Dept Fac. Agr. Pajakumbuh 1957, 
mimeogr., 7 pp. folio); 'Some Notes on the Flora 
of the Taram-Harau Region etc' (ibid, no 3, 1958, 
mimeogr. 5 pp. folio). 

(15) W. Meijer: 'Notes on the Flora of Sepilok 
Forest Reserve 15 miles N of Sandakan' (Forest 
Dept, Sandakan 1960, 12 + 19 pp. folio, stencilled). 

(16) W. Meijer: 'Plant life in Kinabalu National 
Park' (Mai. Nat. J. 24, 1971, p. 184-189, pi. 61-62). 

Meijer's collectors, W. 

in Sabah; a.o. Kapis bin Sisiron and Kilang. 

*Michael Anak, see Forest Dept North Borneo, 

Micholitz, W. 

Collecting in Borneo (see Cycl. Fl. Mai. I, 1, 
1950) took place in 1903-04. 

*Miki, Dr Shigeru 

(1901, Kagawa Pref., Japan; x) graduated at 
Dept Bot. Fac. Science, Kyoto University, 1925. 
For a long time Professor of Botany (paleobotany) 
of Osaka City University, later of Mukogawa 
Women's College. 

Collecting localities. DuringWorldWar 
II in New Guinea (Manokwari, Momi, etc., 1943), 
and in Borneo (Danau Saloh, Feb. 1945). 


Collections. About 2000 specimens from 
New Guinea in Kyoto Herb. Fac. Science (specimens 
inserted in the general herbarium by family); at 
least partly also in Herb. Bog. (> 10 nos). 

Biographical data. Portr. in Hatusima, 
Flora Ryukyus 1971, p. 63. 

*Mikil, C, see Forest Dept North Borneo. 

Miiitante, Miss P. J. 
geologist, Ph.D. from Stanford (Cal.). 

♦Millar, Andree Norma (nee Manners-Sinclair) 

(1916, Paris, France; x) studied arts at Wood- 
ford House, Sussex, and then Auckland University 
and came with her husband, John William Millar 
(1912, Sydney, N.S.W.; 1966, Lae, Terr, of New 
Guinea), a marine engineer, to the gold fields of 
Bulolo, Morobe District, New Guinea, in 1947. 
Life in the Highlands drew their attention to 
plant collecting as a hobby, starting with living 
ferns, then orchids, and finally other plants as 
well. In the meantime she taught in the European 
school at Bulolo, and through the interest of the 
Forestry Office there they began collecting both 
live and herbarium specimens for Lae. In 1956 
she joined the staff of the Lae Botanic Gardens, 
combining horticultural with field work, starting 
as Technical Assistant and leaving the Gardens in 
January 1971 as Curator. At that time she became 
Curator of the Gardens of the University of Papua 
and New Guinea, Port Moresby. She hopes to 
create a purely native botanic garden. 

Author of orchid and garden articles for various 
overseas magazines, and of a gardening book. 

Collecting localities. Territory of New 
Guinea. 1949-53. Bulolo and surroundings, mostly 
living ferns and orchids. 1954. Accompanying 
WoMERSLEY in Jimi Valley, W. Highlands. — 1955. 
Joining the Forestry Botanic Party following the 
old Muboi trail from Wau, through the Range and 
down to Skin Diwai. — 1961. With D. H. Nicolson 
(see there).— 1962-64. With C. D. Sayers.— 1963. 
With P. Van Royen etc. (see there). Between 
1956-71 she accompanied field work and was 3 
times in Mt Wilhelm area, twice on Mt Kerigomna, 
on Kar Kar I., Long I., and in the Gogol Valley. 
Besides collecting extensively from Lae in the 
Morobe Distr. to the W. Highlands, ontheKokoda 
Trail, and by boat to islands in the Huon Gulf. 
Field botanist on a trip by the Dept of Agriculture 
in Papua, following D'Albertis' route from Daru 
to Kiunga, travelling up the Fly, Strickland, and 
Herbert Rivers and on Lake Murray en route. 
Leading a party down the Sepik River from the 
Yellow R. and through the Murik Lakes and up 
the coast again to Aitape, collecting at all islands 
on the way. Further in Bougainville {Solomon Is.), 
New Ireland and on the W. coast of New Britain 
(Bismarck Archipelago). — 1971 — . Papua. Under- 
taking extensive collecting in the Central District. 

Collections. Herb. Lae, partly together with 
her husband and others, numbered in the NGF 
series until October 1970. Later collections in 
Herb. Univ. Papua New Guinea, starting with no 
UPNG 1001. 

Mitchell, A. S. 
(d. 3 Oct. 1959, Singapore). 



Cyclopaedia of Collectors — Supplement II 


Biographical data. Mai. For. 22, 1959, 
p. 259. 

♦Mitchell Brian Anderson 

(1930, England; x) graduated B.Sc. in Forestry 
with 2A Hons in Botany; 1952-53 Military Service; 
in 1954 one year post graduate scholarship with 
British Nature Conservancy; 1954-65 Silvicultural 
Ecologist with the Malayan Forest Service, Ke- 
pong. From 1965 Forestry Officer with the 
Commonwealth Forest Research Institute, in 
charge of the Southern Regional Station at Mt 
Gambler, South Australia, engaged on pine 
nutrition research. 

Collecting localities. 1964-65. Malay 
Peninsula. — 196S. Christmas Island (Ind. Ocean), 
from June-Aug. general over the whole island. 

Collections. In Kepong Herb.: small col- 
lection from the Malay Peninsula, and 52 numbers 
from Christmas /.; a dupl. set from Christmas I. 
will be sent to Leyden. 

Mochtar bin Musa 

(see Suppl. Cycl. Fl. Mai. I, 5, 1958) is identical 
with Mantri Muchtar of Herbarium Bogor. He 
is recorded by Kostermans {in litt.) to have 
collected 60 field numbers in Komodo {Lesser 
Sunda Is.) in Oct. -Nov. J96I. He accompanied 
W. J. J. 6. DE Wilde and Mrs B. de Wilde on 
their expedition to the Leuser complex, in 1972. 
In S. Lampongs (S. Sumatra) he collected 115 nos 
(Jan.-Feb. 1972). 

♦Moll, Victor Wilhelm 

(1930, Salatiga, Java, Indonesia; x) was educated 
at the Forestry School at Buitenzorg. From 
1953-58 Assistant on a rubber plantation in 
Sumatra (East Coast). In the latter year he went 
to Holland, and from May 1959-1963 'Bosarchi- 
tect' in the Forestry Service in Dutch New Guinea. 

Collecting localities. fV. New Guinea. 
J 959. Biak I. (Siabes), Kebar Valley (Andjai) 
(.Sept. 4-11), Biak I. (Mansforbo, Son, Parieri, 
Soendei) (Nov. 6-Dec. \).—1960. Oransbari (Feb. 
10); Oemboei, near Andai, Manokwari (May 11- 
19); with W. ViNK (see there) to the Terr, of Papua 
& New Guinea {June-}u\y); Oransbari (Nov. 18). — 
1961. P. Adi (Feb. 24-Apr. 9); Warsamson Valley, 
E of Sorong (July 16-Aug. 24); Kebar Valley 
(Aifat R.) (Nov. 21-Dec. \\).—1962. Babo Distr., 
Sjuga-Wagura area (Armina, Kwafa) (Apr. 19- 
May 20); Supiori I. — From 1959-63 in the sur- 
roundings of Manokwari. 

Collections. Herb. Manokwari: about 1100 
nos; dupl. at Leyden; numbered in the BW series. 

♦Momot, sec Boswczen Nieuw Guinea. 

Moore, Chas. 

(sec Cycl. Fl. Mai. I, 1, 1950) collected also 
Cyperaceae in Duke of York I. 

•MfM>re, Jr. Harold Emery 

{V>\1, Wmthrop, Mass., U.S.A.; x) was edu- 
cated m Massachuscts (B.S. 1939), Harvard (M.A. 
1940; Ph.D. biol. 1942). From 1947-48 Technical 
A.^sistant Asa Gray Herbarium (Harvard), 1948- 
5 1 Assistant Professor of Botany Bailey 1 lortorium . 


Cornell, subsequently Associate Professor (1951- 
60) and Professor and Director of the Bailey 
Hortorium, N.Y. He collected palms on a world 
tour in 1963-64. 

Collecting localities.^ 1963. Malaya 
(early Dec.-29th): Singapore and Bukit Timah F. 
R., Johore and Mersing F. R., a Forest Reserve 
on the road to Kluang (17 miles from Jemaluang), 
Kuala Lumpur, G. Takun; departure for Borneo 
(30), Sarawak, tour on Mt Matang (31).— 796-/. 
Vicinity of Kuching, several times to Mt Matang, 
4-day trip to Baku National Park (Jan. 2-6), 
limestone hills near Bau (Jan. 8), Bintulu (9-10), 
Nyabau (11); overnight stop at Brunei and pro- 
ceeding to Sabah {N. Borneo) (Jan. 14-26, with 
Dr W. Meuer): Kebon China, Mt Walker, 
Sepilok F. R. (15), Uchung Tanjong trail (16), 
Sandakan, Jesselton (19), Kimanis F. R., Beaiifort 
Hill (20), Tenom, stop at Rayoh, slopes above Sg. 
Masanoi, slopes of Bt Tenom (in Crocker Ridge 
Reserve), N. end of Sabah (25), Kota Belud, 
Maruda Bay, Jesselton. — Terr, of New Guinea: 
Lae (arriving Mar. 1), road to Buiolo and Wau, 
road to Goroka, Bupu Village (2500 ft) (3^), 
vicinity of Lae, Buiolo (10), Bulolo-Watut Divide 
(11), Wau (12-13), by plane to Wagau (14), Lae 
proceeding to the Solomon Is. and New Caledonia, 
Fiji, etc. 

Collections. In Herb. Bailey Hon.: 75 nos 
Sarawak, 11 from Sabah.* Duplicates will be 

Literature. (1) H. E. Moore Jr: 'Palm 
hunting around the World. II. Malaya and 
Sarawak' (Principes 9, 1965, p. 103-117, fig. 89- 
107); ibid. III. 'Sabah and Australia' {I.e. p. 138- 
152, fig. 125-148); ibid. V. 'New Guinea to the 
New Hebrides' {ibid. 10, 1966, p. 64-85, fig. 1-29); 
'New Palms from the Pacific' {I.e. p. 85-99, fig. 
1-7; incl. a Cyrtostachys from Papua). 

(2) H. E. Moore Jr «fe W. Meijer: 'A new 
species of Arenga from Borneo' (Principes 9, 
1965, p. 100-103, fig. 86-88). 

Biographical data. Americ. Men of Sc. 
10th ed. 1961, p. 2842. 

Moraux-Baas Becking, Louise Hermine 

Rightly: (b. Brummen, Netherlands; 1969, 
Renkum, Gld., Netherlands). 

♦Morgan, Tom R. 

A soils officer of the Papua and New Guinea 
Department of Agriculture. In 1968 living at 
Surrey Hills, Vict., Australia. 

He made a few collections in E. New Guinea, 
mainly herbaceous plants, in the 1950s; in Herb. 

♦Morley, R. J. 

Research Assistant at the Geography Depart- 
ment, University of Hull, England, accompanied 
J. R. Flenlxy on a tour to W. Java and Sumatra 
in \')12. 

Collecting localities. 1972. Sumatra 
and yy. Java (sec under Flkni ev). After Flenlev's 
departure he continued to Malaysia, and visited 
Tasck Bcra in Malaya, and Mt Kinabalu in Sabah 
(N. Borneo). He returned to Britain on Nov. 4. 

Collections. With Flenliv (sec there) and 
M. K. Kardin, and alone nos 1-48 {Sabah), and 



Flora Malesiana 

[ser, I, vol. 81 

49-98 {Malaya). In Herb. Hull Univ., etc. (see 

*Morris, (Sir) W. 

Collections. Singapore plants in Herb. 
Edinb. (purch. 1895 with Herb. F. Henderson). 

Mortensen, Ole Theodor Jensen 

Biographical data. Add: Dansk Bot. Ark. 
21, 1963, p. 68. 

*Mortier, C. M., see Boswezen Nieuw Guinea. 

Moszkowski, Max 

Collections. Add: New Guinea dupl. (1910) 
in Herb. Edinb. 

Moulton, John Coney 

Collections. Add: Plants from Kinabalu, 
N. Borneo (1913), and from Sarawak in Herb. 

Mousset, J. P. 

Collections. Add: Java plants (1906) in 
Herb. Edinb. 

Moysey, Louis (or Lewis) 

Collections. Add: Plants Mai. Peninsula 
(1937) with KiAH bin Salleh also in Herb. Edinb. 

M.S., see Moh. Shah 

Specimens preferably to be cited with his full- 

*Muas, A., see Forest Dept Sarawak. 

*Muchtar, see Mochtar bin Musa. 

*Muin bin Chai, see Forest Dept North Borneo. 

*Mujin, M. A., see Forest Dept North Borneo. 

*MulIer, Jan 

(1921, Rotterdam, Netherlands; x) was educated 
at the College for Tropical Agriculture, Deventer, 
Netherlands; from 1946-67 Palynologist of the 
Royal Dutch Shell in Venezuela and Borneo; from 
1967 Palynologist at the Rijksherbarium, Leyden. 

Collecting localities. Malaysia. 1962- 
64. In Brunei, P. Labuan and P. Mullankassan, 
estuaries of Brunei and Tutong rivers. 

Collections. Specimens from 22 marked 
trees, nos 1-22, in Herb. Leyden. Mangrove plants.^ 

Literature. (1) J. Muller & S. Y. Hou-Liu: 
'Hybrids and Chromosomes in the genus Son- 
neratia (Sonneratiaceae)' (Blumea 14, 1966, p. 337- 

Mundo, A. del 

Add: Assistant Professor of Botany, Agricul- 
tural College, Central Luzon. 

*Murata, G. 

accompanied a Japanese expedition to Sumatra 
(Aug. 4-Sept. 20, 1971); other members: Prof. 
M. HiRANO (algologist), Dr K. Iwatsuki (bryol- 
ogist). Dr J. Dransfield (see there) from Bogor 
joined the expedition. 

As he is the author of papers on phanerogams. 

it was probably he who was responsible for the 
collecting of higher plants. 

Itinerary. 1971. N. Sumatra: Sibolangit (Aug. 
11-12) and Toba Lake, Sikundur Nature Reserve 
near Tandjungpura (13-15), Gajolands (18-29), 
climbing a.o. G. Kemiri; Takengon area (early 

Collections. Herb. Kyoto! 

* Murdoch 

probably a collector of H. N. Ridley. He 
collected in Selangor in 1908; in Herb. Sing. 

*Narciso Jr, P., see Philippine National Herbarium 


Collecting localities. Add: 1967. Apr. 
10 with SoEGENG to New Guinea (Irian) to take part 
in the border expedition with Australian New 

Nee, Luis 

Literature. (1) Add: For itinerary in Alaska 
cf. Bot. Not. 1940, p. 297. 

Biographical data. Add: In Hemsley & 
Hooker, Biol. Centr.-Amer. 4, 1887, p. 119. 

N.E.G.C., see Cruttwell, N. E. G. 
preferably to be cited with his full name. 

Nelson, David 

Collections. Add: According to Dr New- 
combe his botanical specimens of Cook's last 
voyage seem to have become mixed with those of 
Menzies (cf. Arch. Brit. Columbia, Mem. no 4, 
1923; n.v.). 

Literature. (1) Add: For itinerary in Alaska 
cf. Bot. Not. 1940, p. 296. 

*Neubauer, Johann (Hans) Franz 

(1911, Vienna, Austria; x), biologist, educated 
at Vienna University (Ph.D. 1934); from 1948-51 
Expert Advisor to the Ministry of Agriculture, 
Kabul Afghanistan, and teacher at the Faculty of 
Science; from 1952-62 Professor of Botany, 
University of Indonesia, Bandung; in 1962-63 in 
East Pakistan, and from 1963-65 in the Faculty 
of Science at Kabul University once more. From 
May 1965 teacher and custodian in the Faculty of 
Science at Giessen University. 

Author of numerous papers on general biology, 
morphology, pharmacognosy. 

Collecting localities. Between 1952-62. 
S. Sumatra: from Kroe via Martapura to Tandjong 
Karang (Aug. 1954), beach at Ulu. Java: Nusa 
Kambangan (Apr. 13, 1955), Dago, Batavia, 
Kuningan, Tjiater near Bandung, Pasir Djati 
(Jan. 24-28, 1961), Tjisolok (Wijnkoopsbaai), 
Kadipaten, Bromo, Idjen, Dieng, Pasir Putih (E 
of Pasuruan) on the beach. Bali (1959 and 1961) : 
Lake Batur and Bratan. 

Collections. His Afghan collection is put at 
the disposal of Prof. Rechinger at Vienna. 
Indonesian collections mainly in Herb. Bandung 
Univ., but also in Bot. Lab. Padjadjaran Univ. 
Bandung, and less in Herb. Bog.; some dupl. in 
Herb. Leyden (rec. Mar. 1966). 

The numbers are mostly those by which the 
specimens are kept in Bandung herbaria. In cases 




Cyclopaedia of Collectors — Supplement II 


where those numbers got lost on transportation to 

Europe, the plants were numbered above 5000, as 

the Afghanistan collection amounted to near 50)00. 

Biographical data. Who is who in Austria. 

New Guinea Forests (formerly Forces) 
Add: Michael Galore*, A. E. H. Ross*, R. 

RiDGWELL*, etc. After no 50.000 the series letters 

were changed into LAE. 
Collections. Add: Also dupl. in Herb. 



♦Newell, Thomas Kenneth 

(1939, New York City, U.S.A.; x) graduated 
from the New York State Agricultural and 
Technical Institute (1962), B.Sc. from California 
State Polytechnic College in Ornamental Horticul- 
ture (1964), and subsequently a graduate student 
at the University of Hawaii, majoring in plant 

He was working on a thesis on the genus 
Joinvillea, and made a trip to the Malesian region 
to collect material and to make observations on 
pollination, dissemination, and ecology of that 


Sabah: Sandakan (Aug. 5), Jcssclton (9), vicinity 
of Sabah National Park and along the road 
between the Park and the town of Tamparuli (II- 
16); Sirtfiapore (20); .Songkla in Thailand (28); 
Malay Peninsula: G. Hijau Claiping) (31), G. 
Brinchang (Cameron flighlands) (Sept. 2-4), 
Fravcr's Hill (7); F'acific: Solomon Is., liji, Samoa. 
Collections. Mainly Joinvillea,^ but also a 


few Flagellaria and Hangitana; nos 141-175 (176- 
255 from Pacific islands). In Herb. B. P. Bish. Mus.; 
dupl. will be sent to Sandakan, Paris, Leyden, 
Florence, and Kew. 

Literature. (1) Th. K. Newell: 'A Study 
of the Genus Joinvillea' (J. Am. Arbor. 50, 1969, 
p. 527-555, 6 fig., 1 tab.). 

*Newton, Ken. G. 

An Agricultural Officer in Australian New 
Guinea: in 1968 stationed at Samarai, Papua. He 
collected a few specimens in the Sepik District in 
1957; in Herb. Lae. 

*Ng, Francis S. P. 

(1940, Kuala Lumpur, Malaysia; x) graduated 
1964 B.Sc. (Hons) in Botany at the University of 
Tasmania; joined the Forest Research Institute, 
Kepong, as Forest Botanist in June 1964. He is 
writing chapters for a Tree Flora of Malaya, and 
makes expeditions to various parts of Malaya to 
collect material. He went on a Colombo Plan 
Scholarship to Oxford University for three years 
from October 1968; D.Sc. 1971. At present Forest 
Botanist at Kepong, and co-author of the Tree 
Flora of Malaya.^ 

Collecting localities. Malay Peninsula: 
in various parts. In 7966 a.o. in Johore, Penang, 
Selangor, Malacca, Perak (Tapah Hills, Gopeng, 
Slim R., Pahang (G. Tahan); in 7967 in Negri 
Sembilan, Johore (Mersing), Ulu Kelantan, Lower 
Perak, Sg. Chior and Siput in Perak; in 7965 in 
the Cameron Highlands and Perak (Kledang 
Saiong, and Bubu F. R.). 

Collections. Herb. FRI Kepong: dupl. at 
Kew, Leyden, Sing., Arn. Arbor., Sandakan, 
Kuching. Medio 1968 about 2200 nos. In the FRI 
series and 100 early collections in KEP series. 

Literature. (1) Author of 'A Guide to the 
Arboreta at Kepong and Changlun' (Mai. For. 
28, 1965, p. 289-306, map); Tn search of Combreto- 
carpus' (I.e. 29, 1966, p. 32-33); 'Allantospermum 
in Bubu Forest Reserve' {I.e. 31, 1968, p. 165-169). 

Ngadiman bin Ha(d)ji Ismail 

(d. Sept. 1958). 

Biographical data. Card. Bull. Sing. 17, 
1959, p. 337-338, portr. 

NGF nos, see New Guinea Forests. 

Nicholson, Donald Ian 

Add: He retired from the Sandakan Herbarium 
in 1965; subsequently he joined the Forest Depart- 
ment in North Queensland, at Cairns, Australia. 

* Nicolas 

1st lieutenant, accompanied the 'Star Mountains 
Expedition' to Central New Guinea in 1959 (sec 
under C. Kai kman, in whose series some plants 
have been collected). He took part in the trip to 
Ml Antares. 

•Nicolson, Dan H. 

(1933, Kansas City, Mo., U.S.A.; x). botanist, 
educated at Cirinncll College (Iowa; A.M. 1955), 
Stanford University (M.H.A. 1957). and Cornell 
University (N.Y.; M..Sc. 195'>); Teaching Assistant 
at C\irncll, V)5') V)W. His main interest is 



Flora Malesiana 

[ser. I, vol. 8^ 

Araceae on which he planned to write his thesis. 
For the study of this family he made a trip to 
Malesia in 1960-61 and subsequently visited 
several European herbaria. Ph.D. 1964 Cornell 
University, and subsequently on the staff of the 
Smithsonian Institute. 

Collecting localities. Philippines, stay- 
ing at the College of Agriculture, Laguna, Sept. 
y960-Jan. 1961. Luzon: to summit Mt Makiling 
(Sept. 18, I960); Mindanao (22-29): Davao, 


Maparat, Compostella, Mampising, Apocon School, 
base Mt Apo to 1200 m (25), Agusan, Esperanza, 
Sibagat, Sianib (29); Camiguin I. (Oct. 2); Min- 
danao (2-5): Misamis Or., Binoni to Mambajao, 
Bukidnon, Musuan, Lake Apo; Luzon: Coll. Agric. 
(8), Molawin Creek (20), Sorsogon and Bulusan 
National Park (24-27), Mt Mayon to 700 m (28); 
Catanduanes, center of island {29); Luzon: Laguna, 
Coll. of Forestry (26), Quezon National Park (27); 
Samar: Bagacay Barrio (Dec. 1); Leyte: Putok 
River (3); Negros Oriental: Mt Canlaon to ICXX) m 
(6); Luzon: Laguna, Coll. of Agr. (14-17), to 
summit Mt Makiling (18); Coll. of Forestry (Jan. 
8, 1961). — Java: stay at Kebun Raya Indonesia, 
Bogor (Jan.-May 1961): Tjiseeng (Jan. 28), Hort. 
Bog. (Feb. 2), Djasinga (7), Tjampea (16), G. 
Gedeh (between 1700 and 2700 m, 17-18), Hort. 
Bog. (24-26), Tjibodas (Garden and along way to 
Tjib(e)ur(e)um, 1400-1600 m) (Mar. 2-3), Hort. 
Bog. (Mar. 14-Apr. 1), Pelabuhan Ratu (2), Hort. 
Bog. (3-23), Palabuhan Ratu (24), Hort. Bog. (25- 
May 10); Singapore I.: Hort. Sing, (to June 14); 
Malaya: KJang Gates Reservoir, Selangor (17); 
Bot. Gard. Penang (19-21); Perak: Taiping and 
vicinity (22-25); Singapore L: Hill, Bt Timah, etc. 
(July 4-15); Selangor: Klang Gates (16), Gunting 
Simpang F. R. (17), Batu Cave Ravine and Telok 
F. R., Klang (18), to Eraser's Hill (19); Cameron 


Highlands in Pahang (20-23), summit G. Brin- 
chang (21), G. Beremban (22), G. Jasar (23); 
Johore (28); Singapore: Bot. Gard. (Aug. 1-2). 
NW. Borneo, Sarawak: Semengkok F. R. (3), 
Matang F. R. (5), Bau (5, coll. Edward Taylor, 
1 no: 6), Bako National Park (7-8), Setapok F. R. 
(9); Singapore L: Bt Timah etc. (13). By way of 
Australia (coll. a few numbers) to Terr, of New 
Guinea: Morobe Distr., N of Lae (Sept. 21); 
Western Highland Distr. (26, partly coll. with 
Millar), Lae Bot. Gard. (30-Oct. 1, 10-11), Lake 
Wanum (W of Lae, Oct. 1, in NGF series), 
Bumbu River (8); Papua: Centr. Distr., Brown 
River F. R. (12), Sogeri Plateau (13), headwaters 
Kemp Welch R. (14), Sogeri Rubber Estate; Terr, 
of New Guinea: Malambe track up Bussu R. (17), 
Lae Bot. Gard. and Patep Creek (20), Edie Creek 
Road and near Wau (21), Bulolo (23, 25), Oomsia 
Creek (25), Bot. Gard. Lae (26), Markham R. 
(26); New Britain: Rabaul and Kerevat (28-29); 
Bougainville (30-Nov. 2): Arawa Plantation (3()- 
31, Nov. 1), Kupei (Nov. 1); Terr, of New Guinea: 
Morobe Distr. (5) ; New Britain (9), road to airport 
(9-10); Terr, of New Guinea: Morobe Distr., Lae 
Bot. Gard. (12), Madang (17), Wewak (17); 
Dutch New Guinea: Lake Sentani (17), Biak I. 
(17), Manokwari and Faninda (18), Manggoapi 
(19), Tafelberg F. R. (20), Agric. Station (21), 
Maripi (21), Maroemi (22), Biak (24). 

Collections. Dupl. from Malesia in Herb. 
Ley den. 

*NieI, Joannes Petrus van 

(1930, The Hague, Netherlands; x) was educated 
at the Rijkstuinbouwschool Boskoop (tropical 
section; 1947-50); rubber planter in E. Sumatra 
(1951-56), and Australian New Guinea (1957-60). 
From 1960 onwards Assistant Palynologist, later 
Palynologist in the employ of the B.I.P.M. (Oil Cy) 
respectively at The Hague, in Argentine, from 
1964-1971 at Seria (Brunei), at present living in 

Collecting localities. Mostly in Brunei, 
NW. Borneo, 1964 onwards. 

Collections. Herb. Leyden: nos 3377-4415 
(in July 1969); no dupl. From 1951-60 he collected 
molluscs for the Amsterdam Zoological Museum. 
He makes pollen slides, of which dupl. sets are at 
Pondichery (Inst. Fran^ais), India, and Herb. Bog. 

Nielsen, Finer Steemann 

(under Steemann Nielsen in Cycl. & Suppl. 
Cycl. Fl. Mai. I, 1 & 5) also joined the Danish 
'Deep-sea Galathea Expedition', 1950-52. 

In 1944 appointed Professor of Botany in the 
Pharmaceutical College in Copenhagen; since 1962 
Lecturer in Oceanography at the University. 

Biographical data. Dansk Bot. Ark. 21, 
1963, p. 42, w. portr. ; in A. Hansen, Den Danske 
bot. litt. 1940-1959, 1963, p. 227-231 (bibliogr.). 

Nieuwenhuis, Anton Willem 

Literature. (1) Add: Peterm. Geogr. Mitt. 
44, 1898, p. 9-13, map. 

*Niimura, Taro 

(1917, Shimosuvva-machi, Nagano Pref., Japan; 
1951, Tokyo, Japan), graduated from Suwa 



Cyclopaedia of Collectors— Supplement II 


Middle School in 1935; Laboratory worker of the 
National Science Museum at Tokyo, 1938-51. He 
accompanied Y. Satake (see there) to New 

Collecting localities. IV. New Guinea, 
March to June 1943. 

Collections. About 700 sheets of flowering 
plants and ferns ; Herb. Nat. Sc. Mus. Tokyo. 

*Nomura, Noboru 

(1905, Niihama, Kyoto Pref., Japan; x) gradu- 
ated at Kyoto Normal School; teacher at an 
elementary school in Kyoto. 

Collections. Collected in Johore {Malaya) 
(Aug. 1936); in Herb. Kyoto. 

*Noona Dan Expedition 

from the University of Copenhagen, 1961-62.^ 
In total 19 persons of different disciplines partici- 

Itinerary.^ Noona Dan Expedition, 1961-62. 
1961, The 'Noona Dan' arriving in Mindanao on 
Aug. 1 ; Palawan: lowland near Brooke's Point 
(2 months), collecting from Uring-Uring (Aug. 13- 
26), Penigisan (Sept. 1-2), Tagimbung (5-13), 
Penigisan (15-21), Makagwa (21-25), Uring-Uring 
(29-Oct. 1); Ursula I. (Oct. 2); Balabac I. (4-12); 
Tawi-Tawi: Tarawakan (19-27); Luzon: Mt St. 
Thomas (Nov. 5); Tawi-Tawi: Tarawakan (13-17), 
Lapid-Lapid (17), Batu-Batu (18), Lapid-Lapid 
(20-22), Batu-Batu (26); Bongao 1. (26); Papahag 
I. (26); St. Cruz /., Zamboanga, Mindanao (Dec. 
11); Sapamoro, NE. of Zamboanga (16-20). — 
1%2. Bismarck Archipelago: New Ireland, Kavieng 
(Jan. 12), the nearby Nago I. (13), Mussau (16- 
Feb. 18); Manus (20), Dyaul (Mar. 2-10), New 
Hanover (Mar.); New Ireland: Kalili (Apr. 8-9), 
Lelet (Apr.); New Britain: Yalom (May 22-23); 
Manus (June 22-25); Hermit Is.: Luf I. {11); New 
Britain: Cape Hoskins (July 5-10); Credner I. 
(17); New Britain: Matupi (20), Vulcan Point (23), 
Rabaul (24); Solomon Is. (Aug. 18-30). 

Collections.* Herb. Copenhagen: c. 3300 
nos; during the last part of the expedition attention 
was specially focussed on the collecting of Fungi. 

For collectors see under S.-E. Sandermann 
Olsen, M. E. K01E, H. DissiNG, S. F. Christian- 
sen, and T. L. Wolff. 

Unfortunately hardly any duplicate material 
was collected; the labelling was rather un- 
satisfactory for a period after S.-E. Sandermann 
Olsln left the expedition. 

Literature. (1) T. Wolff: 'The Noona Dan 
Expedition' (Nature 198, 1963, p. 1044-1045); in 
Noona Dan Paper no 19. 

Cf. also: 'Chronological List of Localities in 
Noona Dan Paper No. 55' (Dansk Bot. Ark. 25, 
no 2, 1967, p. 18-19 and maps 1-7). 

(2) 'Botanical Report of the Danish Noona Dan 
r.xpcdition 1961-62 to The Philippines, The 
Hisrnarck Archipelago, and The Solomon Islands' 
(Noona Dan Paper no 55 in Dansk Bot. Ark. 25, 
no 2, 1967, 88 pp.. 27 fig.). For the list of numbers 
and localities sec p. 19-20. 

B. O. van Zanten: 'Mosses of the Danish 
Noona Dan Expedition to the Philippines, 
Bismarck Archipelago and the British Solomon 
Islands'(J. HalloriBot. Lab. 31, 1968, p. 135-151). 


*Noor bin Jumaat, Mohd 

= NuR BIN Jumaat (see there). In 7970 he 
collected with Dr E. A. Heaslett in Johore, with 
T. C. Whitmore in Taman Negara, Pahang, and 
with Dr R. Hill in Johore, all Malaya. 

*Nooteboom, Hans Peter 

(1934, Waingapoe, Sumba, Dutch East Indies; 
x), botanist, taking his M.Sc. at Leyden University 
in 1961 ; employed at the Laboratory of Experi- 
mental Plant Taxonomy at Leyden. He is working 

J. p. van niel 

on a thesis on the taxonomy and phytochemistry 
of the Symplocaceae. Earlier he worked on 
Papilionaceae and Simaroubaccae. In 1969 and 
1970 he made explorations in Malcsia and Thai- 
land for supplementary collections of Symplocos, 
sponsored by WOTRO. 

Collecting localities. 1969. N. Thailand 
(Jan. 6-27); IV. Java: Tjibodas and Mt Pangrango 
(Feb. 1 1-15); Sabah (former N. Borneo): G. Alab 
(19-27). Mcndulong (Mar. 3). Sg. Muyaya (G. 
Lumaku (4-7), Malaman (9), Tambunaii (15-16), 
Ingaran (18), G. Trusmadi (20 21). Repatriated 
after an unfortunate fall, returning in the next 
year.- 1970. .SV/ranv/A; arrival at Kuchinj^t Mar. I); 
.Sahah: to Kola Kinabahi (3). and rcsthousc 
Kinabalu National Park (1500 m) (4); Mt Kina- 
balu (5 13), also on F'cnibukan Ridge; Kota 
Kinabalu (14), Sandakaii (Id). Sarawak: with P. 
C MAI in the environs of Hario (25 29); to G. 
Miirud (Mar. 30 Apr. 4); ascent (J. Murud (4- 
11); return at Baiio (13) and collecting in the 



Flora Malesiana 

[ser. I, vol. 8^ 

environs (up to 20); Apo Batu Buli Range and 
mountain plain between this range and Batu 
Lawi (22-27); P. Chai staying another weei<. 

Collections. Herb. Leyclen: c. 1600 Borneo 
(Sabah and Sarawak) numbers, including the 
collection made by P. Chai (see above). 

♦Nordin, see Forest Dept North Borneo. 

Rijksherbarium at Leyden and Herb. Bog. have 
no set, it is probably that set which was presented 
to Berlin, presumably by Reinwardt or later by 

Literature. (10) Replace by: C. G. G. J. van 
Steenis & M. J. van Steenis-Kruseman: 'The 
Plates of Javanese Plants of Francisco Norona 
with a revised evaluation of his generic names' 
(Regn. Veg. 71, 1970, p. 353-380). 

Biographical data. Add: Bull. Acad. 
Malg. n.s. 35, 1958, p. 13-16; Diet, of Maurit. 
Biography p. 757. 

Norris, Sir William 
Collections. Add: and in Herb. Oxford. 

Nouhuys, Jan Willem van 
(d. 1962, Netherlands). 
Biographical data. 

Aardr. Gen. 2e reeks 81, 


Add: Tijd. Kon. Ned. 
1964, p. \-^, portr., 



stand for Numbered Trees, distributed in the 
SAN series. 

*Nur bin Juma(a)t, Mohamad 

See sub H. M. Burkill 1959. Identical with M. 
NooR bin Jumaat (see there). 

Nur bin Mohamed Ghose, Mohamed 

(d. Nov. 1958, Singapore). 

Collections. Add: Dupl. Malay Peninsula 
(1937-59), with Sinclair, Shah & Kadim bin 
Abdul in Herb. Edinb. Sometimes as Noor on the 

Biographical data. Gard. Bull. Sing. 17, 
1959, p. 331-336, portr. 

*Nusi, A., see Boswezen Nieuw Guinea. 

Ochse, Jacob Jonas 

(d. 1970). 

Biographical data. Add: Bull. Fairchild 
Trop. Gard., July 1970, p. (6)-(12). 

*Ogata, Dr Ken 

of the Forest Experiment Station, Meguro, 
Tokyo, collected a large number of wood samples 
with reference herbarium specimens in Sabah (2nd 
half 1968); the first half of the year he was in 
Malaya, collecting at least on G. Gajah and G. 
Tempurong in Perak (Feb. 14-15). 

Ohna, Miss H. S. 
Add: took a Ph.D. in pharmacy in the U.S.A. 

Olegario Jr, D., see Philippine National Her- 

Norona, Francisco 

(r. 1748, Sevilla, Spain; Jan. 12, 1788, Mau- 

A letter (to M. Van Marum, dated March 29, 
1819; in Arch. Dutch Society of Sciences at 
Haarlem) from C. G. C. Reinwardt (see there) in 
Java, it is evident that the^latter had the dis- 
position of a set of Norona's plates. As the 


*01sen, Sven-Erik Sandermann 

(1920, Helsingor, Denmark; x) was educated at 
the Pharmaceutical Highschool, Copenhagen 
(cand. pharm. 1944). Pharmaceutical chemist, 
since 1966 at Vesterbro Apotek, Copenhagen. 

He participated in the Danish 'Noona Dan 
Expedition' (see there for itinerary etc.) from July 
31, 7967-Feb. 20, 1962. 



Cyclopaedia of Collectors — Supplement II 


Biographical data. Dansk Bot. Ark. 21, 
1963, p. 45, 244 (bibliogr.). 

Oosterzee, Louis Albert van 

Literature. (1) Add: For annotations on the 
itinerary cf. W. Vink in Nova Guinea, Bot. 22, 
1965, p. 482^83. fig. 2 and 5. 

Ooststroom, Simon Jan van 

Add: Retired Feb. 1971. 

Biographical data. Blumea 19, 1971, p. 
1-2, portr. 


accompanied E. F. de Vogel (see there) to S. 
Borneo (Kalimantan Selatan) in 1973, and evidently 
collected in a series of his own. 

*Orden, J., see Philippine National Herbarium. 

Osbeck, Pehr 

Literature. Add: (5) C. Hansen & A. Fox 
Maule: 'Pehr Osbeck's collections and Linnaeus's 
Species Piantarum (1753)' (Bot. J. Linn. Soc. 67, 
1973, p. 189-212, 6 pi., 1 fig.). 

Biographical data. Add: G. & Bo 
Petterson: 'En Reseberattelse av Prosten Pehr 
Osbeck ar 1776' (in Arsb. Varbergs Museum 1967, 
22 pp., portr. and summary). 

*Othman bin Haron, A. F. O. 

ecologist, Assistant Forest Botanist, Sandakan. 

Collecting localities. 1964. Sarawak 
(NW. Borneo): ecological expedition on Tau 
Range (July-mid Oct., 600 nos coll.); area of Bt 
Raya, Pelagus, and Bah Sama P. F., Ulu Rejang 
(Nov.); area of the Ulu Segan and Nyabau block, 
Similajau F. R., and Bintulu (Dec.).— 7969. 
Sarawak. A month's collecting expedition to the 
middle reaches of the Balieh R. (Nanga Menyong) 
and overland to Bukit Mersing (in the 4th Div.) 
for another month. 

Collections. Herb. Sandakan. 

Otto-Surbeck, Rosa Lydia 

Collections. Add: At least up to 1964; 
dupl. prcs. to Herb. Leyden. 

*Overstreet, Miss Elsie 

later married a Mr Burr, living in California. 

Collecting localities. /9i7./'ena«^(Sept. 
30); Thailand: Bangkok (Oct. 13, 16); Sumatra: 
Palcmbang (22). Lahat (23), Ranau (24, 27), base 
of G. Dcmpo (27), Pageralam to Tjoeroep (28-29); 
Java: Poentjak (Nov. 5), Borobudur (8), G. Bromo 

Collections. Small duplicate set in B. P. 
Bish. Mus. Herb. Hawaii (prcs. 1938), identified 
at Ixydcn in 1968; some specimens in ffcrb. 
leyden. All together about 1000 nos: location of 
1st set unknown. 

Oye, Paul Herman Gustaaf van 

(d. 1969). 

Biocrapmical data. Add: Hydrobiologia 
35, 1970, p. iii-xxvii, portr.; Biol. Jahrb. Dodonaca 
38, 1970, p. 7-38, portr., bibliogr.; Vakbl. Biol. 50. 
1970, p. 66-67. 


P nos 

see J. W. Purseglove (personal collections). To 
be cited under his full name. 

*Pachiappan, G. 

Laboratory Assistant, University of Malaya, 
Kuala Lumpur, will continue collecting for the 
Phytochemical Survey of Malaya after the 
departure of J. Carrick (end 1967). He is a 
vigorous field worker, who has learned many 
skills in both Botany and Chemistry Departments. 

Collections. From 1968 he will make field 
collections with Teo Leong Eng (see there); they 
will begin their numbers as T. «& P. 1 (PCSM no 

*Paie, Ilias, see Forest Dept Sarawak. 

Palla, Eduard 

Add: evidently visited S. Sumatra in Feb. 1901, 
as a Scleria was collected at Pladjoe (Palembang) 
on the 26th. 

Collections. Add: Dupl. in Herb. Vienna. 

*Palo, S. L., see Philippine National Herbarium. 

*Palser, Barbara Frances 

(1916, Worcester, Mass., U.SA.; x) was edu- 
cated at Mt Holyoke College, B.A. 1938, M.A. 
1940; Ph.D. University of Chicago 1942; In- 
structor Botany Univ. Chicago 1942-45, Assistant 
Professor 1945-51, Associate Professor 1951-60, 
Professor 1960-65; Associate Professor of the 
Dept of Botany, Rutgers University, New Bruns- 
wick, N.J., U.S.A., 1965, and Professor since 1966. 

Author of several papers on the morphology of 

Itinerary. 7969. Z,Mzo«.Touringthe Mountain 
Province (May): from Baguio (Mt Santo Thomas) 
to Bontoc (Mt Data, Mt Polls) and Banaue. 

Collections. Ericaceae, nos 6901-6924, a set 
of which in Leyden Herb.; pickled material for 
morphological investigation at home. First col- 
lection since Loher of Rhododendron zoUingeri 
J.J.S. in Luzon; also recollection of Diplycosia 
parvifolia on Mt Polls (1st collection was lost at 
Manila). Dupl. Philippine Ericaceae and Epacri- 
daceae in Herb. D.S./.R. Christch. 

Biographical data. Amer. Men of Science; 
Who's Who of American Women ed. 5, 1968-69. 

Pancho, Juan V. 

Later Professor in Botany. 

Collecting localities. Add: Collecting 
tour (Oct. 17. l964-h\n. 23. 1965) in Luzon: Mt 
Sto Thomas and vicinity. Mountain Prov.; Mt 
Makiling and Mt Banahao, Laguna; Atimonan, 
Quezon; Taal Volcano, Batangas; Mt Isarog, 
Camarincs Sur; Mayon Volcano. Albay; Bulusan 
Lake, Sorsogon; in Negros, NW. part; in Min- 
danao: Davao, Marawi. Lanao del Sur; in Basilan 
and Palawan (Puerto Princcsa). 

Collections. Add: He accompanied the 
Dutch wasp specialist J. T. Wiihks. for whom he 
collected Ficus plants (43 spp.) and a few others, 
numbered in J'ancmo's scries, 4161-4271. The 
plants were sent to CoRNtR. Herb. Cambridge. 



Flora Malesiana 

[ser. I, vol. 8^ 

♦Panoff, Mr and Mrs (Michell and Fran^oise?) 

Anthropologists working in the Jacquinot Bay 
area. New Britain, collected plants. Partly in Herb. 
Lae (no dupl.): 187 nos in total. Authors of 
'I'Ethnologue et son ombre' (Paris 1968). 

Park, Mungo 

Correct: His voyage to Bencoolen (5. Sumatra) 
took place in 1793-94, the 'Worcester' sailing from 
Portsmouth Apr. 5, 1793, returning at Moorings 
May 2, 1794.* 

Literature. (2) C/. Ch. Hardy: 'Register of 
ships employed in the Service of the Hon. United 
E.LC. etc: (London 1799) p. 61. 

Paijmans, Kees 

Add: (1921, Dordrecht, Netherlands; x) from 
1956-62 Forest Officer, Forestry Department, 
Ghana (W. Africa); from 1963 Plant Ecologist, 
C.S.LR.O., Division of Land Research, Canberra, 

His papers include a publication on Mt Albert 

Collecting localities. 1950-51. Nunukan 
I. (off NE. Borneo).^— 1963. Papua: c. 100 miles 
east of Port Moresby and 50 miles southeast of 
Popondetta in lowland rain-forest and savannah, 
sea-level to about 4000 ft.^— 1967. W. Papua: Fly 
River region to c. 400 ft, also in monsoonal 

Collections. Add: Herb. Canberra: nos Pj 
1-174 (coll. 1963), and Pj 175-453 (coll. 1967). 

Literature. (1) See internal report of the 
Indonesian Forestry Department. 

(2) K. Paijmans: 'Vegetation of the Safia- 
Pongani area, (in Lands of the Safia-Pongani area. 
Territory of Papua and New Guinea. Land 
Research Series no 17, C.S.LR.O., Australia 1967). 

(3) K. Paijmans: 'Vegetation, forest resources 
and ecology of the Morehead-Kiunga area' (in 
Land resources of the Morehead-Kiunga area. 
Territory of Papua and New Guinea. Land 
Research series no 29, C.S.R.LO., Australia 1971). 

(4) K. Paijmans & E. Loffler: 'High-altitude 
Forests and Grasslands of Mt Albert Edward, 
New Guinea' (J. Trop. Geogr. 34, 1972, p. 58-64, 
2 fig., 2 pi.). 

PB tios, see O. Beccari. 

PCSM nos 

are used by contributors to the Phytochemical 
Survey of Malaya of the Federation, Kuala 
Lumpur. If KL is on the same label, only the latter 
should be cited. 

Peekel, Gerhard 

Biographical data. Add: In Streit & 
DiNDiNGER, Bibl. Missionum 21, 1955, p. 458^60, 

*Peinberton, Cyril Eugene 

(1886, Los Angeles, Cal., U.S.A.; x) well- 
known entomologist (Hon. Sc.D. of Hawaii) of 
the Exp. Station of the Hawaiian Sugar Planters' 
Association (1919-53), collected sugar-cane var- 
ieties on behalf of that association, together with 
Jeswiet and Brandes (see those) in E. New 


Guinea in 1928; in 1929 he collected cane in the 
vicinity of Rabaul, New Britain. 

In 1937 he made a 3-months visit to New 
Guinea and the Bismarck Archipelago together 
with C. G. Lennox (see there) for the collecting 
of true seed of wild relatives of the sugar-cane. 
They visited New Hanover, New Ireland, N. New 
Britain, and Austr. New Guinea (Kainantu, Wau- 
Bulolo, Salamaua).* 

It is not known to me whether he made her- 
barium specimens. 

Literature. (1) Cf. Hawaiian Plant. Rec. 55, 
1958, p. 213; Cane Growers' Quart. Bull. 15, 1951, 
p. 41-42. 

Biographical data. In Amer. Men of 
Science (Phys. & Biol.) ed. 10, 1961. 

*Pennek, Sutan 

collected in Nunukan I. near E. Borneo (Kali- 
mantan) in March and July 1951. Evidently with 
K, Paijmans (see there) and probably numbered 
in the same series. 

Collections. In Herb. Bog.; dupl. in 
Leyden, Kew, Dehra Dun. 

*Pennington, T. D. 

(1938, Ewell, Surrey, England; x), B.A. (Botany) 
1960, and Ph.D. (1965), both at Oxford University. 
Research Officer Dept of Forestry Oxford, 1965- 

The Leverhulme Trust Fund sponsored his tour 
to the Far East, 1963-64. 

His work on generic delimitation of the Melia- 
ceae^ came to an abrupt stop in 1970, when 
financial support for tropical taxonomy was cut 
by 50%. He started his own horticultural business 
and now devotes part of his time on taxonomy, 
usually at Kew. 

Itinerary. 1963. Malaya, West Coast (Sept.); 
Sarawak (Oct.): Lundu, Rejang River, Kapit; 
Sabah (Nov.): Sepilok, lower slope Mt Kinabalu; 
Singapore and East coast Malaya to Mersing 
(Dec). — 1964. Terr, of New Guinea (Jan.): Lae- 
Wau, Bulolo; New Britain, Honiara, New Cale- 
donia, Fiji (Suva). 

Collections. For. Herb. Oxford: c. 350 
numbers, mainly Meliaceae, with a few Ebenaceae 
and Chrysobalanaceae dupl. in Sing., Kep., 
Kuching, Sandakan, Lae, Honiara, Suva. 

Literature. (1) Author of 'Materials for a 
monograph of the Meliaceae I. A revision of the 
genus Vavaea' (Blumea 17, 1969, p. 351-366, 2 fig.). 

Biographical data. Photogr. in Principes 
9, 1965, p. 110, fig. 98. 

Penzig, Otto 

Literature. (2) Add: 'Come nasce una 
foresta virgine' (La Lettura 24, 1924, p. 833-838, 
13 fig.). 

Peters, Christian 

Add: His voyage with Capt. Hagemeister was 
made in the 'Krotky'. Collections in Herb. 

*Petersen, H. J. van, see Boswezen Nieuw Guinea. 

*PetIo, G. J. 
collected no 20, Digitaria argyrostachya (Steud.) 



Cyclopaedia of Collectors — Supplement II 


Fern, at Banjoebiroe, Semarang (Centr. Java), at 
470 m, in 19 J6. In Herb. Bog. with Backer label 
and writing. 

*Phang, C. I. 

collected ferns from Bt Anak Takun, Selangor, 
Malaya (Sept. 15, 1960). Phanerogams too? 

Philippine National Herbarium 

To Suppl. Cycl. Fl. Mai. I, 5, 1958, p. cccx 
under above-mentioned entry, add the following 

1. H. G. Gutierrez*, Ern. J. Reynoso*, R. M. 


2. Melecio Agra {1955), B. Aldos {1955), G. 
Banlugan, Carolyn, P. Cordero, Mrs Elisa D. 
Dawa {1951), ViCTORiNO Gabot {1955), Gerardo 
Landingin {1955), P. P. Lazo {1955), Ricarte 
Martin {1955), V. Menor {1955), P. Narciso Jr 
(7955), D. Olegario Jr (7955), Jose Orden (7955), 
S. L. Palo (7956), R. Quitoles (7955), P. P. 
SuMAGUE (7955), J. B. Susara (7955), M. Tolen- 
tino, C. Valdez. 

Several of the 7955 collections were made in 
Jan.-Feb. on Mt Maquiling, Laguna Prov., Luzon, 
possibly by participants of a forestry course. 

Names with asterisk see separate entry. 

Collections. Herb. Manila: dupl. to Leyden, 
Kew, Arn. Arbor., Sing., U.S. Nat. Mus., Bogor, 

*Philipson, Melva Noeline 

(1925, Palmerston North, New Zealand; x) 
took her B.Sc. (1948), and her M.Sc. (1953) at the 
University of New Zealand; on the staff of the 
Dairy Division Dept of Agriculture, 1949-51; in 
the Microbiology Dept, Canterbury Agricultural 
College, 1951-55; in the Botany Division D.S.I.R., 
from 1963 onwards. 

Itinerary and Collections.* 7965. Terr, 
of New Guinea. With her husband, W. R. Philip- 
son (see there). 

Literature. (1) Melva N. Philipson: 'Plant 
Hunting in New Guinea' (New Zeal. Rhodod. 
Assoc. Bull, no 54, 1969, p. 10-14). 

•Philipson, William Raymond 

(1911, Newcastle-upon-Tyne, England; x), B.A. 
Cambridge University (1933). Research Student, 
Kcw Gardens (1933-35), Ph.D. London University 
(1947); on the staff of the Dept of Botany. British 
Museum. 1935-51 ; since 1957 Professor of Botany 
in the University of Canterbury, Christchurch, 
New Zealand. Besides in New Guinea, he col- 
lected in Jamaica (1939). Columbia (1953), New 
2:caland (1953). and the Cook Islands (1969). 

Specialist and author on Araliaceae. 

Collecting localitijs. I96H. Terr, of 
New Guinea: environs of Lae (Aug. 4). Kassam 
Pass and Kainantu (5), Okapa Road and back to 
Lac (6); environs of Wantowat. Finistcrrc Range 
(7-10); Hdic Creek (13-16); to Goroka by road 
(20); to Mt Wilhclm by air (21); to Kundiawa by 
road. Chimbu Valley (26); Kcrawagi and KorongI 
River (27), over Daulo Pass to Goroka (28); 
return to Lac by road (29). 

( or.LrcTiON.s. New Guinea collections made 
jointly with his wife, M. N. Pmnp.vjN (see there); 


mainly Rhododendron and Araliaceae, nos 3250- 
3535; first set Bot. Div. DSIR, Lincoln, New 
Zealand; not complete dupl. sets at Leyden, 
Edinb., Kew, Arn. Arbor. 

*Philpotts, Miss 
In Herb. Kew a Cleome from Singapore (culta). 

*Phytochemical Survey 

of the Federation of Malaya, Kuala Lumpur. 
Collections. Numbered in the KL series, by 
various collectors. See also PCSM. 

Pierot, Jacques 

Add : (died on the outward journey, and not on 
the home-voyage). There are contradictory data on 
his role in the foundation of the Dutch horticul- 
tural society. 

Collections. It is evident from letters in the 
archives of the Leyden Herbarium that the 
Japanese plants ascribed to J. Pierot were bought 
in Java for joint account of him and G. Bisschop 
(The Hague). They were sold to the Rijksher- 
barium in January 1844. This collection has labels 
written by Miquel on which Pierot is mentioned 
as collector. All these specimens have a 2nd label 
with a number, and generally some dates on 
locality, and vernacular names. By comparison 
Prof. Dr H. J. Venema found that the latter labels 
were written by H. BOrger and that the collection 
must be a duplicate set of the same.* In 1841 
Burger was in Holland, and it is still a mystery 
who sold this set of Burger plants. 

Literature. (2) Cf. Meded. Bot. Tuinen 
Belm. Arb. 3, 1959, p. 58. 

*Pierre, Louis 

(1833, Champ-Borne, Reunion; 1905, Paris), 
son of a sugar planter, studied medicine in Paris 
and Strasburg, but did not finish his studies due to 
family circumstances. In 1861 he went to India 
as the collaborator of Thomas Anderson in the 
Calcutta Botanic Garden. In four years time he 
developed himself in a way that the English were 
eager to keep him in India, but he preferred to be 
in a French colony, and was from 1865-1877 in 
charge of the Jardin botanique at Saigon. In those 
years he organized numerous expeditions in the 
interior of Cochinchina, Cambodia, and Siam, 
bringing together the materials for the 'Flore 
forestiere de la Cochinchine' and the later 'Flore 
dc rindo-Chine'. 

After a study tour in Java and India in con- 
nection with Cinchona culture, he returned to 
Paris to work on his tree flora and on several 
plant families. 

No pea pierrei and several other plants have been 
named after him. 

Coi I. acting localities. 7577. W. Java: 
probably at Batavia. Bogor. Ijibodas. G. Gedeh- 
Pangrango. and eventually in the Prcangcr 
Regencies (Mts Malabar, Papandajan. etc.). 

CoLLEC tions. 1/erh. Paris, small collection. 

•PinK, I'atrick, see Forest Dept North Borneo. 
V] nos, see K. Paijmans. 



Flora Malesiana 

[ser. I, vol. 8^ 

Pleyte, Didit Rudolf 

Literature. (1) Add: For sketch map of 
Misool L with principal collecting localities, see 
Nova Guinea n.s. 9, pt 2, 1958, p. 298. 

PNH nos, see Philippine National Herbarium. 

Poivre, Pierre 

Collections. At the end add: also MSS in 
the library of Angers (France).* 

Literature. (3) Add: According to Madeleine 
Ly-Tio-Fane, in 'Mauritius and the Spice Islands' 
(see sub Biogr. Data) not all the statements are 
reliable and less satisfactory details have been 
omitted or glossed over. 

(6) A summary of Poivre's observations on the 
cultivation of the spice plants at Isle de France 
(B. Angers, MS 612, p. 569, fig. 28). 

Biographical data. Add: In M. Ly-Tio- 
Fane (ed.): 'Mauritius and the Spice Trade. The 
Odyssey of Pierre Poivre' (Publ. no 4, of the 
Mauritius Arch. Publ. Fund, Port Louis 1958, \v. 
portr. and photogr. of seal); M. Ly-Tio-Fane: 
'Pierre Poivre et I'expansion frangais dans I'lndo- 
Pacifique' (Bull. Ecole Fran?. d'Extr.-Orient 53, 

1967, p. 453-511, pi. I-XIV); 'Commemoration du 
bi-centenaire de I'arrivee de Poivre a I'lle Maurice 
1767-1967' (Proc. R. Soc. Arts & Sci. Mauritius 3, 

1968, p. 101-130); 'Poivre's contribution' in 
'Mauritius and the Spice Trade' II (The Triumph 
of Jean Nicolas Cere, Paris/The Hague 1970, p. 

*PoIunin, J. 

collected in Br. N. Borneo (Aug. 1953); in 
Herb. Sing. 

*Poore, Martin Edward Duncan 

(1925, Dunkeld, Scotland; x) was educated at 
Edinburgh University (1946-47), and at Cambridge 
University (1947-52; M.A., Ph.D.). Professor of 
Botany at the University of Malaya, Kuala 
Lumpur (1959-64), Dean of Science there (1964- 
65); appointed Lecturer in Commonwealth 
Forestry Institute, Oxford, in 1965. Since 1966 
Director of the Nature Conservancy, London. As 
ecologist he joined the 2nd Royal Society Ex- 
pedition to Mt Kinabalu, N. Borneo, in 1964. 

Collecting localities. From 1959-65 in 
Malaya, especially in Selangor, the Cameron 
Highlands and Eraser's Hill. In 1964: N. Borneo 
{cf. Corner and Chew Wee Lek).^ 

Collections. Three large crates presented to 
Herb. Univ. of Malaya (KLU) (1966). 

Literature. (1) C.-Ch. Ho & M. E. D. 
Poore: 'The value of Mount Kinabalu National 
Park, Malaysia, to plant ecology' (Mai. Nat. J. 19, 
1965, p. 195-202, map, 5 photogr.). 

Popta, Sicco Martinus 

(d. Feb. 26, 1968, Bandung, Indonesia). 

Collections. Add: Many specimens for the 
educational herbarium of the 'Institut Teknologi 
Bandung', of which he was keeper since 1950. 

Porsch, Otto 
(d. 1959, Vienna, Austria). 


Posewitz, Theodor 

Literature. (1) Add: and in Peterm. Geogr. 
Mitt. 32-33, 1886-87. 

*Postels, Alexander F. 

geologist who accompanied Lutke's voyage in 
the 'Senjavin' (see sub Mertens). He collected a 
few plants in Alaska (cf. Bot. Not. 1940, p. 300) 
and might have collected elsewhere during the 
voyage. In Herb. Leningrad. 

Potts, John 

Collections. Add: Herb. Deless. (Geneva) 
(cf. Taxon 19, 1970, p. 536). It seems very im- 
probable that he collected in Sumatra; specimens 
from the latter island will have been collected 
either by Raffles or W. Jack. 

♦Pound, D. 

He might be identical with James Pound (1669, 
Bishop's Canning, Wiltshire, England; 1724) who 
matriculated at Oxford in 1687, graduated B.A. 
and M.A. in 1694; medical diploma 1697 with a 
degree of M.B. Having taken orders, he entered 
the service of the E.I.C., went out to Madras in 
1699 as a chaplain to the merchants of Fort St. 
George, whence he proceeded to the British 
settlement of Pulo Condore off Thailand. Upon 
the mutiny of the native troops there on March 3, 
1705, he escaped to Malacca and ultimately 
reached Batavia. He returned to England in 1706, 
where he was presented a rectory. He did important 
work in astronomy. His collections and papers 
were destroyed in Pulo Condore, but it seems quite 
possible that he collected some plants in Java. 

Collections. In Herb. Oxford (with Herb. 
DuBois) one specimen 'collectum a D. Pound in 
insula Java'; probably early in the 18th century. 

Biographical data. In Diet, of Nat. 
Biogr. vol. xlvi, 1896. 

PP nos, see O. Beccari. 

Pratt, Antwerp E. 

Literature. (3) Add: Some additional 
information on his stay in the Anggi Lakes region 
by W. ViNK in Nova Guinea, Bot. 22, 1965, p. 484. 

*Price, Michael 

from Columbia University, arrived at Los Bancs 
Agricultural College in 1968. He finished his 
degree in 1971. Now Research Instructor in the 
College of Agriculture Herbarium, Los Banos 
Units College, Laguna, Philippines. He is specially 
interested in ferns. Collections? 

*Prijanto, Botjah 

(1942, Djombang, E. Java; 1969, Makassar, S. 
Celebes), was educated at the (Bogor) Tjiawi 
College of Agricultural Sciences (B.Sc. 1962), and 
joined the Botany Division of the Forestry Re- 
search Institute; in 1963 he continued his education 
at the University of Edinburgh (Ph.D. in 1966, on 
taxonomic studies in the Scrophulariaceae), 
subsequently studying for a year at Stockholm 
and two months at Utrecht, in palynology. 

Collecting localities. Joined collecting 
trips by Kostermans (see there) to Udjon Kulon 
Nature Reserve in W. Java and Sumbawa I., in 



Cyclopaedia of Collectors — Supplement II 


1968 joined M. Jacobs (see there) in an expedition 
to South Sumatra. In J969 he took part in a trip 
to SiV. Celebes with E. Larsen (see there) where 
they met a tragic death in a car accident. 

Biographical data. Reinwardtia 8, 1970, 
p. 1-2, portr., bibiiogr. 

Prince, John 

Add: (1770, baptized at the Parish Church 
Greenwich, Kent, England; ?) was appointed on 
the Fort Marlborough establishment (Benkulen) 
of the E.I.C. in 1786, arriving there on 29 Aug. 
1787. Appointed Assistant to the Resident of 
Natal (Maidman) on 31 March 1790, and follow- 
ing the French capture of the northern settlements 
in 1794, he took Maidman's place, holding the 
post until April 1798. In that year he went to 
Tapanuli, where he was appointed Resident on 
20 June 1801. In December of that year he and 
his wife went to Calcutta on leave to recover their 
health, but returned to Tapanuli in 1802. In 1816 
provisionally appointed to take charge of Fort 
Marlborough, but that was revoked in favour of 
somebody else. Prince was employed by Raffles 
along with William Jack, on a mission to Pulu 
Nias in September 1820, and P. succeeded R. as 
superintending officer in charge of Benkulen 
during the transfer of the British west coast 
establishments to the Dutch in 1825. Later in the 
Straits as Resident Councillor, taking an active 
interest in the attempts to keep going the Botanical 
Garden started by Raffles. 

Dr J. Bastin was so kind to give above infor- 
mation; he found evidence quite opposed to 
Jack's description of Prince (see Cycl. Fl. Mai. I, 
1). Raffles in a letter to his sister wrote (Sing. 18 
Nov. 1825; India Office Library: Drake collection 
MSS Eur. D 742) that "it has been said that he is 
not only a Prince by name but a Prince by nature 
too". This judgement was shared by Baptists 
missionaries and others. 

Collections. Add: His despatch of seeds 
and flowers of the Sumatran camphor tree to 
Roxburgh enabled Colebrooke (who received 
them due to Roxburgh's departure for England) 
to give his excellent account of the plants in 
Asiatick Researches xii (1816). Prince's own 
description of the plant is given too (Dr J. Bastin 
inlitt. July 1972). 

Proppc, Dr Gregor Joseph 
collected at Madiun too. 

Provost, Jcan-Mathieu-Simon 

Add: (1728, Auray, France; before 1778, ? Isle 
de France) served as supercargo of the 'Compagnic 
dcs Indcs', spoke Malay fluently, and later settled 
at Isle dc France. He was picked by Poivki; as 
technical leader to obtain spice plants in the 
Moluccas, hiirly 1768 he sailed to Kcdah in the 
'Vigilant' to contact the Macassarcsc traders. I his 
resulted only in a promise from the Macassarcsc, 
and in May 1769 he set out once more and pro- 
ceeded to the Philippines and the Moluccas, this 
time with success. 

Subsequently he was made Honorary Inspector 
of the spite plants being grown at the 'J.irdin du 
Roi' in Mauritius. The next year he handed the 
care of the plants to Ci.Ri (see there) and sailed 


again in June 1771 with the 2nd Moluccan 
Expedition (see sub Sonnerat).^ His visit to 
Gebe(h) was a great success. The plants were 
brought to Mauritius and distributed to other 
French colonies, including Cayenne. 

Literature. (1) Add: Mad. Ly-Tio-Fane in 
'Pierre Poivre et I'expansion fran^aise dans I'lndo- 
Pacifique' (Bull. Ecole Fran?. Extr.-Or. 53, 1967, 
p. 472^73, etc.). 

pullen, Finisterre Range, 1904 

(2) Some pages of his diary are reproduced in 
Mad. Ly-Tio-Fane I.e. sub Liter. (1) pi. iv-v. A 
plan of the harbour of Gebe is reproduced in I.e. 
pi. vi. 

Biographical data. M. Ly-Tio-Fane, 
Mauritius and the Spice Trade, 1958, p. 10-11. 

PS nos, see O. Biccarl 

PUH nos, see Philippine University Herbarium, 
Quezon, in Suppl. Cycl. IL Mai. I, 5, 1958, p. 

•Pukol, Hasan 

Collector of the F-orest Dept at Brunei, Borneo; 
numbered in tiie BRUN series. 

Pullen, Royal 

(Oli.k riNG localitiis. Continued col- 
lecting in Auslr. New Guinea. 1957. Central 
Highland. 1958. Lower K;umi River.— 1959. 



Flora Malesiana 

[ser. I, vol. 8^ 

Lower Sepik River and Prince Albert Range. — 
1961. Papua, Southern Highlands: Mcndi, lower 
slopes of Mt Giluwe, lalibu (June 26-July 8); 
lalibu Basin (-July 17); lalibu, Kagua (-July 25); 
Kagua, Erave River valley, Mubi Valley, Lake 
Kutubu (-Aug. 12); Lake Kutubu, Benaria, Tari 
(-23); Tari, Koroba (-26); Koroba, Korenda, 
Kwongebi, Tari (-Sept. 3); Margarima, E. slopes 
of Doma Peaks, Nipa (-9); Nipa, Lai (Kawuku) 
valley, Mendi (-16); Mendi Basin, ascent of Mt 
Giluwe to 12,500 ft from W. side (20-23).— 1962. 
Papua: Port Moresby-Kairuku area (July 1-Oct. 
6), including Laloki, Goldie and Brown Rivers, 
to Kapa-Kapa and Rigo, S. foothills of Astrolabe 
Range, to Redscar Bay, Vanapa R., Sogeri Plateau 
and N. side Astrolabe Range, vicinity of Galley 
Reach, Kanozia, to Aroa R., to Hall Sound, 
Angabunga R. and Mekeo area, Biaru R., W. to 
Palipala Hills, Yule /., Hall Sound, lower reaches 
of Brown and Vanapa Rivers, mouth of Gallay 
Reach. — 1963. Terr, of New Guinea: Western 
Highlands Distr., leader of the C.S.LR.O.- 
Rijksherbarium Expedition to the Kubor Range 
(June 25-Sept. 11): northern (Wahgi valley) side 
of the range to S of Minj, camps I-III (resp. at c. 
10,600, 11,700, and 13,300 ft), with a climb to 
Mts Kinkain. — 1964. Papua: Northern Distr. 
(July 24-Sept. 14), S. side Hydrographers Range 
near Siurane, near Numba, Pongani River valley 
and Falls, western flanks of Sibium Range S of 
Toma, Barihi R. valley, upper southern slopes and 
summit ridge of Hydrographers Range to c. 6000 
ft. Terr, of New Guinea: Eastern Madang Distr., 
Finisterre Range with the Brit. Mus.-Newcastle 
uponTyne Expedition (Oct. 6-Nov. 24; itinerary see 
sub C. D. Savers).^— 1965. Not in New Guinea.— 
1967. Papua: Port Moresby area (Apr.-May), 
savannah area along Morehead R., W. Distr. (end 
of July-?).— Mid-1969. Owen Stanley Range, 
including Mt Wadimana in the Simpson Massif. — 
1972. Expedition to Mt Suckling in Papua, 
collecting June-July at low altitude with G. 
Leach (see also P. F. Stevens and J. F. Veldkamp). 

Collections.* For the distribution of CSIRO 
plants see under Hoogland. The collection 1959 
amounted to 590 nos; in 1961 (June-Sept.) col- 
lection series 2637-2890; in 1962 nos 3084-3388; 
in 1963 nos 5000-5421; in 1964 nos 5422-5950, 
5951-6192; in 7967 > 400 nos; dupl. 1964 in 
Herb. Edinb. About 300 nos (8242-8515) were 
collected by Pullen on Mt Suckling in 1972. 
Ecological samples (extra material) were allotted 
numbers in the series 3810-3823, 6193-6423. 

Literature. (1) Including a new Pittosporum, 
cf. K. Barker in Nova Guinea n.s. 9, 1958, p. 
339-340, 1 fig. 

Mosses by E. B. Bartram in Brittonia 11, 1959, 
p. 86-98. 

(2) A. C. Jermy & C. D. Savers: 'An Expedition 
to the Finisterre Mountains of Eastern New 
Guinea' (J. R. Hort. Soc. 92, 1967, p. 117-128, 
fig. 54-55, 57-63, 1 map). 

*Pulsford, Mary 

wife of R. Pulsford, an Agricultural Officer at 
Dagua, Sepik District, New Guinea (N. coast), 
collected some tens of plants between 1953-54; 
in 1968-69 Pulsford served in the Public Health 
Department, Port Moresby, Papua. 


Collections. From the Sepik Distr., Terr, 
of New Guinea, low numbers on Dagua airstrip; 
also with A. G. Floyd (see there) in the NGF 
series; in 1969 >136 nos in the UPNG series of 
the University at Port Moresby. 

Purseglove, John William 

Add: About 1967 appointed Advisor on 
Tropical Crops for the Ministry of Overseas 
Development, working at East Mailing Research 
Station, Maidstone, Kent, England. 

Collecting localities. Add: 1957. Sara- 
wak: Bako National Park (Feb. 2-18) with MOH. 
Shah bin Haji Moh. Nur. 

Quaife, Dr 

Add: He collected in the Cameron Highlands 
in Oct. 1940. 

Quisumbing, Eduardo 

Add: He retired as Director in November 1961, 
and was for some following years attached to the 
Araneta University. 

Saccolabium quisumbingii L. O. Williams has 
been named in his honour. 

Collecting localities. Add: 1960. Papua 
(Sept. \5).—1961. Batanes & Babuyanes Ex- 
pedition (Mar. 25-Apr. 4; with H. Gutierrez 
and R. del Rosario):* Bat an I., Sabtang /., 
Itbayat I., Calayan I. (May 2-10); interior of 
SanVicente in Cagayan Prov. (May); Batan I. 
(June 3-8); Mt Iraya (on the summit and on the 
Pacific side) and Mt Makatarem. — 1963. Borneo 
{Sarawak, Brunei, N. Borneo; first half of July):* 
Baku National Park and Santubong; Oran 
Reserve; Semengoh F. R. (10) and Bau Limestone 
Hills; Tutong White Sands (11), Badas peat 
swamps (12). 

Collections. Mostly orchids. 

Literature. (1) E. Quisumbing: 'The 
Vegetation of Batan and Babuyan Islands' 
(Symp. Ecol. Res. Humid Trop. Veget. Kuching 
1963: UNESCO Sc. Coop. Off. SE. Asia 1965, 
p. 31-38). 

(2) E. Quisumbing : 'Orchid Hunting in Borneo' 
(Araneta J. Agr. 11, 1964, p. 76-78). 

Biographical data. Portr. in Philip. Orch. 
Review 7, 1959, p. 33. 

*QuitoIes, R., see Philippine National Herbarium. 

Quoi, Jean Rene Constant 

Biographical data. Add: Act. Mus. Hist. 
Nat. Rouen 27, 28, 1922, p. 486; in Bibliogr. 
Austr. Entom. 1775-1930, 1932. 

Raap, Hugo 

Collections. Add: is cited to have collected 
lichens and mosses in Columbia in 1904 {cf. 
Urban, Gesch. Kon. Bot. Gart. Mus. Berl.-Dahl. 
1916, p. 289,311). 


(d. Dec. 19, 1957, Bogor, Indonesia). 

Raciborski, Marian 

Biographical data. Add: Ochr. Przyr. 29, 
1963, p. 9-16, w. portr. (w. English summary). 



Cyclopaedia of Collectors — Supplement II 


Raffles, Sir Thomas Stamford Bingley 
The Drake collection (c/. Suppl. Cycl. Fl. Mai. 

I, 5, 1958, p. cccxiii) is now in the E.I. Office, 

London (inf. by Dr J. Bastin, July 1970). 
Biographical data. J. Bastin: 'Raffles 

the Naturalist' (Str. Times Annual for 1971, p. 

58-63, col. ill.); 'Raffles in Marble and Bronze' 

{ibid, for 1972, 6 pp.). 

*Rahim bin Ismail, A. 

of the Forest Research Institute at Kepong, 
Malaya, collected 365 FRI nos (of Kepong) in 
Pahang, Trengganu, Perak, and Kelantan in 

Identical with Abd. Rahim of the Forest Dept 
North Borneo ? (see there). 

Rahman bin Mohd Ali, (Enche) Abdul 

Add: (1925, near Kuantan, Pahang, Malaya; 
24 Sept. 1969, Kuantan, Malaya) entered the 
Forest Department in 1947, and left in February 
1952 to follow a four-year course in forestry in 
Australia. After having attained his B.Sc. he 
returned to Malaya as Deputy Conservator of 
Forests. In 1957 District Forest Officer at Klang, 
in 1959 at Kuala Pilah; subsequently posted in 
Trengganu, Johore, in 1963 at the Forest Research 
Institute, Negri Sembilan, and Kuala Lumpur, 
acting on two occasions as Deputy Director. 

Biographical data. Mai. For. 33, 1970, 
p. 3-6, w. portr. 

*Ramli bin Zainuddin 

of the Forest Research Institute at Kepong, 
Malaya, collected 34 FRI nos (Kepong) in the 
Kinta Hills, Perak. 

Ramos, Maximo 

Collections. Add: Also dupl. Philippines 
in Herd. Edinb. (1907-12). 

•Rappaport, R. 

and Mrs Rappaport (associated with Professor 
Vayda, see there) operated at Simbai, Madang 
District, E. New Guinea, in 1963. Herb. Lae: c. 
420 nos: Pandaniis to Dr Stone, bamboos and 
ferns to Dr Holttum. 

Rappard, Frederik Willem 

His position in New Guinea was rightly: Head 
of the Forestry Division of the Dept of Agriculture. 

Author of a paper on New Guinea.* 

Collections. Besides plants he made large 
collections of Aphidae in Java and New Guinea. 
In Herb. Waneningen: 7 plants from G. Idjen and 
some from SE. Baluran etc., E. Java. 

Literature. Add: (2) F. W. Rappard & P. 
VAN Royen: 'Some Notes on the Vegetation in the 
VVissel l^kcs Area' (Nova Ciuinea n.s. 10, 1959, 
p. 159-176, fig. 1-3, pi. 3-8; in Dutch, English 

*Rashid bin Taggoi, sec Forest Dept Sarawak. 

•Raufcr, F". 

was probably in the 1920s or 1930s in the employ 
of the Forest Department I". M.S., as he collected 
together with R. V:. Coiomh (sec in (yd. 11. Mai. 
I, I, 1950). 


*Read, Mrs, see under Miss L. Hamilton. 

♦Reed, Dr J. G. 

collected no 14, Dodonaea viscosa, Bernam R., 
Perak, Malay Peninsula (Nov. 25, 1941); in Herb. 
Sing. = J. A. Reid? (see there). 

*Rehal, see Forest Dept Sarawak. 

Reid, John Alexander 
Add : Author of some botanical papers.^ 
Literature. (1) J. A. Reid: 'Klang Gates and 
Bukit Takun: Reflections of an amateur botanist' 
(Mai. Nat. J. 5, 1951, p. 109-123, 6 fig.); 'Our 
changing Flora' {ibid. 9, 1954, p. 84-86); 'Plants of 
the quartz ridges' (ibid. 14, 1959, p. 22-31, 2 
photogr., map). 

♦Reid, Dr L. 

linguist from the University of Hawaii. For 
ethnobotanical collection see D. A. Madulid, 
April 1971 {Luzon). 

♦Reilly, P. 

Agricultural Officer in Australian New Guinea, 
in 1968 on Kibagah Plantation, Western Highland 
District, Papua. He collected a small series of 
numbers at Garaina, Morobe District, about 1948: 
in Herb. Lae. 

Reinwardt, Caspar Georg Carl 

Literature. (9) Add: For particulars on the 
drawings of Plant. Ind. Bat. see Cl. Nissen, 'Die 
botanische Buchillustration' 1951, vol. 2, p. 193. 

♦Reynoso, Ernesto J. 

(1934, Lucena City, Luzon, Philippines; x), 
Botany-Aide of the Philippine National Her- 
barium (Bot. Dept National Museum); graduated 
B.S. at the Far Eastern University. 

Collecting localities. Philippines. 1963. 
Lipuun Point, Quezon, Palawan. — 1964. Mt 
Koronon, Sitio Gumate, Davao, Mindanao. — 1965. 
Basilan I., Mindanao, with H. G. Gutierrez & 
Fraga; Mt Apoy, Borongan, Samar, with H. G. 
Gutierrez. — 1966. Mt Guiting-guiting, Sihuyan 
I., Romhhn, with H. G. Gutierrez. — 1967, 
Victoria Peak, Palawan, with R. Espiritu.— 1968. 
Mt Pulog and Mt Tabayoc, Benguet, Luzon, with 
M. Jacohs, D. R. Mendoza, and R. Espiritu. — 
1969. .S'. Samar (sec .mb Gutierrez). Teamed up 
with the Osaka Museum of Natural History. The 
following areas were to be visited: Mt Makiling 
(Luzon), Mt Apo (Mindanao), Quc/.on (Palawan), 
Ilo-Ilo basin (Panay); sec also K. Sito. 

Collections. Approx. 545 Held numbers 
with PNH (series) numbers in Herb. Manila and 
distributed elsewhere. 

Richc, Claude Antoine Gaspard 

C()i i.i,( rif)NS. Part of his Mauritius and 
Reunion plants have been mislabelled 'Java'. 

Richlhofcn, I'crdinand von 

lorthe 'I'icussisclic l-xpcdition' sccalso Pcterm. 
Cicogr. Mitt. K. 1962, 420 424, where letters have 
been published. 



Flora Malesiana 

[ser. I, vol. 8^ 

*Ridgwell, Ray 

a Technical Officer with the Forestry Depart- 
ment New Guinea; collections in the Bulolo-Wau 
area, Morobe District, in 1955, and near Brown 
River and Trans-Vanapa (Oct. 1959), in all some 
tens of numbers in the NGF series. In Herb. Lae. 

Ridley, Henry Nicholas 

Itinerary. 7975. Add: 
(Feb. 4). 

Centr. Java: Borobudur 


Biographical data. Add: Biogr. Mem. R. 
Soc. 3, 1957, p. 141-159, portr.; Proc. Linn. Soc. 
Lond. 169th session, 1958, p. 35-38; J. Mai. Br. 
R. As. Soc. 33, 1960, p. 104-109. 

*Ridsdale, Colin Ernest 

(1944, Bristol, England; x) graduated B.Sc. 
(Hons) at Bristol University in 1966. He arrived 
at Lae, T.N.G., Oct. 1966, on a contract. After his 
return, end 1968, performing post graduate 
research on Uncaria {Rubiaceae)} Ph. D, Bristol 

Collecting localities. Terr, of New 
Guinea. Around Lae (Oct. 7965-Sept. 7965). 1966. 
Edie Creek, Wau (Nov.); near Cape Gloucester, 
ethnobotanic collections assisting Prof. Darke; 
W. New Britain: Salasea (Dec). — 1967. Bougain- 
ville {Solomon Is) (Jan.-Feb.):^ Parairi village 
Kieta and Lake Lotura area; Mt Wilhelm (Feb.); 
Morobe Distr., Kui (Mar.); loma near Popendetta 
(May). Attached to New GuinealWest Irian Border 
Survey Party with Assistant Galore (June-Sept.)' 
(see also sub Soegeng), collecting in the foothills 


of the Star Mts (c. 4000 ft, ± a fortnight), Ingem- 
bit (c. 300 ft, 3 weeks), brief stay at Kiunga, and 
Angamaruk on the Fly River (Papua), savannah 
at Weam on the Bensbach R. (Irian) about 3 
weeks. — 1968. Madang area, Kar Kar I. (Jan.). 
Open Bay New Britain Forest Survey with P. 
Katik,* and a small collection from Lacket, c. 
2300 ft, on limestone. Murray Pass, Mt Albert 
Edward with Paddy Wood (see there) (Aug.). 

Collections. Herb. Lae in NGF series, 
about 2700 nos; Murray Pass collections (1968) in 
Herb. Edinb. Number series and localities at Lae. 

Literature. (1) Author on arborescent 
Nauclea of New Guinea in Gard. Bull. Sing. 25, 
1970, p. 247-281. 

(2) Internal report at Lae. 

(3) C. E. Ridsdale: 'Botanical Results of the 
New Guinea Border demarcation expedition, 1967' 
(Papua & New Guinea Sc. Soc. Trans. 9, 1968, 
22 pp., 6 fig.). 

(4) Internal report at Lae. 

Riedel, Johann Gerard Friedrich 

Collections. Add: Herb. Leningrad: Celebes 
plants (1869). 

*Riekerk, Ch., see Boswezen Nieuw Guinea. 

*Ringers, Miss N. 

collected some Cyperaceae on Mt Kelud (Kloet) 
in E. Java (1915). 

*Robbins, Ross Gordon 

(1919, Wanganui, New Zealand; x) was edu- 
cated in New Zealand (M.Sc. Botany 1950); Ph.D. 
of the University of New Zealand in 1957, on a 
thesis on the New Zealand Forest History. From 
1942-45 with the New Zeal. Army Forces in the 
Middle East; 1951-56 University Lecturer in 
Botany, mostly in New Zealand, but two years 
in Jamaica (1951-52); 1957-60 Plant Ecologist 
C.S.I.R.O. New Guinea Land Surveys; 1961-66 
Research Fellow in Biogeography, Australian 
National University; for seven months acting as 
the first curator of the Canberra Botanic Gardens; 
in 1966 appointed Professor in Botany in the new 
university at Port Moresby, Papua, taking up the 
post in July 1967; in Aug. 1970 he accepted an 
appointment teaching biogeography at Canberra 

His major interests are phytogeography and 
rain-forest ecology in the Pacific. About 1970 he 
resigned and still lives in Australia. 

Some plants, including Dimorphantera rob- 
binsii Sleum. and Barbula robbinsii Bartr. have 
been named in his honour. 

Collecting localities. 1957. E. New 
Guinea:^ Eastern and Western Highlands (Goroka- 
Mt Hagen area) incl. Mt Wilhelm, Mt Hagen, and 
Mt Giluwe (June-Oct.). — 1958. Lower Ramu 
Valley, incl. Adelbert Ranges (July-Oct.); W. 
Java (Dec). — 1959. E. New Guinea: Lower Sepik 
Valley, incl. Prince Alexander Range (June- 
Oct.). — 1960. Western Highlands, Wabag area 
(June-Sept.).2— 1963. In Oct.-Nov. collecting trip 
in N. Thailand, and proceeding to Malaya.^ — 1965. 
Philippines. Mindanao: slopes Mt Talomo (Oct. 2), 
lower slopes Mt Apo (6), side ridge Mt Apo (c. 
2000 m) (7), Mt Talomo (c. 1400 m) (11); Luzon: 




Cyclopaedia of Collectors — Supplement II 


Mt Pulog (c. 2700 m) (24-25), reaching all the 

Collections. End 1961 totalling from 1- 
3430; duplicates infrequent. In Herb. Lae, Leyden, 
etc. This collection includes some 750 mosses,* 
of which a small set from Java. The CSIRO 
numbers include a few dozen Queensland rain- 
forest plants. Thailand collection in ? Herb. 
Canberra: dupl. {nos 3536-3689) in Bangkok; 
mosses also at Leyden (from Malaya 3690-3709). 
Some 50 nos from the Philippines sent to Leyden 
of which sterile material thrown away. His col- 
lections include numbers from New Hebrides, 
New Caledonia, Fiji, Solomons, Java, Malaya, 
New Zealand, U.S.A., Canada, and Hawaii. 

Literature. (1) R. G. Robbins: 'Montane 
formations in the Central Highlands of New 
Guinea' (Proc. S>'mp. Humid Trop. Veg. Tjiawi 
1958, 1961, p. 176-195, map); 'The montane 
vegetation of New Guinea' (Tuatara 8, 1961, p. 
121-134); 'The vegetation of New Guinea' (Austr. 
Terr. 1, 1961, p. 21-32); H. A. Haantjes, R. G. 
Robbins c.s.: 'Lands of the Goroka-Mount Hagen 
area. Territory of Papua and New Guinea' (Land 
Res. Ser. no 27, 1970, p. 1-159, 15 fig., 23 tab., 

(2) Cf. 'General report on Lands of the Wabag- 
Tari area. Territory of Papua and New Guinea 
1960-61' (C.S.I.R.O. Land Res. ser. no 15, 
Melbourne 1965; vegetation description by 
PuLLEN and Robbins, p. 100-113 with 16photogr.). 

(3) R. G. Robbins & J. Wyatt Smith: 'Dry land 
forest formations and forest types in the Malayan 
Peninsula' (Mai. For. 27, 1964, p. 188-216, 12 fig.). 

(4) Cf. H. Crum in Bryologist 62, 1959, p. 283- 
289; E. B. Bartram: 'Contributions to the Mosses 
of the Highlands of Eastern New Guinea' (Brit- 
tonia 11, no 2, 1959); 'Low Altitude Mosses from 
Northwest New Guinea' (ibid. 13, 1961, p. 368- 
380); 'Mosses of the Western Highlands of Eastern 
New Guinea' (Rev. Bryol. Lichen, n.s. 30, 1961, 
p. 185-207). 

Roberson, see Clark & Roberson. 

Roberts, Mrs D. Carson 

collected at least in Sumatra and Java (near 
Bandung) in 1930. 

•Roberts, O., see under D. W. Ives. 

Rock, Joseph Francis Charles 

Add: (d. 1962, Honolulu, Hawaii) lived for 
many years in China, and finally returned to 
Hawaii. He received numerous awards, including 
an honorary Dr.Sc.'s degree of the University of 

Itinerary. Add : 1916. Philippines, Java, and 
Singapore. — 1919. Also in Siam (Thailand) and 

BioriRAPMirAL data. Add: Newsletter Haw. 
Bot. S(x;. 2, 1963, p. 1-13, w. bibliogr.; Taxon 12, 
1963. p. 89-102, porlr., bibliogr. 

*Roe, John Septimus 

("1797 1878), a liculcnant who was with Captain 
KiN<, n'Mii III' F'akkir KiN<i, who surveyed llic 
NW. coast of New Holland. 1818-22, accom- 
panied by the botanical collector A. CUNNiN(iiiAM), 


and collected many specimens. He again visited 
the NW. coast of New Holland in company with 
Captain Bremer o/b H.M.S. 'Harcourt' and once 
more made natural history collections.^ 

Collections. Several of his plants, mainly 
from Australia and Persia, but also from Ceylon 
and Prince of Wales Island (near Cape York 
Peninsula, N. Australia), came into the possession 

R. G. ROBBINS, New Guinea, 1961 

of A. B. Lambert and were later sold. Possibly 
plants from other islands in Torres Strait, S of 
New Guinea, were collected. Now at least in 
Herb. (Geneva). 

Literature. (I) Cf. Hort. S. Miller in 
Taxon 19, 1970, p. 537-538. 

♦Rocby, H. C. J., sec Boswczcn Nieuw Guinea. 

Roemer (Rcimcr), von 

Add: Collcclcd a no 53, C'y penis iria L. in Java 
in /W/. A Goniphandra collected in the same year 
in Java in Herb. Uoiss., (ieneva. 

Coi.i.FCTioNS. Add: No news about the 
identity of the collector was fouiul. Dr I'. Makk- 
(.KAi, who formerly sorted out the material for 
Ikriin, docs remember that llic labels were copied 
by I HI vikanus. 



Flora Malesiana 

[ser. I, vol. 8^ 

Roepke, Walter Karl Johann 

(d. 1961, Wageningen, Netherlands). 

*Ro]o, Justo P. . . . . , 

of the Forestry Department, School of Agricul- 
ture, Los Banos, Luzon. From Nov. 1968 he did 
post-graduate work on plant taxonomy at the 
Rijksherbarium, Leyden, with a one-year grant. 
Now Chief of the Forests Products Collection 
Section under the Wood Technology Service, 
National Science Development Board, College, 
He collected with William L. Stern (see there). 

♦Roorda van Eysinga, N. P. P. J. 

(1894, Middelburg, Netherlands; 1940, killed m 
battle, Loosduinen/The Hague, Netherlands), 
book-keeper; from 1920-23 attached to the 
K.N.LL. (Netherlands Indian Army) at Magelang, 
Central Java. After return to the Netherlands he 
took up his old profession. In his spare time he 
studied the flora. 

Collecting localities. Centr.Java. 1920- 
23. Environs of Magelang, Kedu Residency. 
Possibly elsewhere in Java. 

Collections. Presented (1971) to Herb. 
Leyden, together with a Dutch collection partly 
made together with his son, P. N. H. G. Roorda 
VAN Eysinga (born at Magelang, 1920), Agricul- 
tural Engineer, and partly by the latter (1940-47). 
It includes also a small Swiss collection leg. J. H. 
RooRDA van Eysinga, and some cultivated plants. 


collected > 24 nos of plants in E. Borneo 
{Kalimantan), vicinity of Samarinda {1940); in 
Herb. Bog. 

*Rosario, Romecaldo M. del 

completed a 2-year study in the U.S.A., taking 
a M.S. degree in botany on a thesis on North 
American Asterella {Hepat). June 1964 he was 
back at the Manila Herbarium. In 1968 he went 
once more to the U.S.A., pursuing a Ph.D. (bryo- 
logy) at the University of Illinois, Urbana. He was 
expected to be away for three years. 

Co-author of 'Philippine aquatic flowering 
plants and ferns' (Manila 1967), of a Bibliography 
of Philippine Bryology (± 1968), and author of 
papers on liverworts. 

Collecting localities .1961. Expedi- 
tion to the Batanesand Babuyanes Is (Itinerary see 
sub Quisumbing). — 1964. Makiling National Park, 
Luzon (July). — 1965. Kalanga, Mountain Prov. 
with P. A. CORDERO Jr. — 1966. Guimaras L — 7967. 
Luzon: Dalton Pass, Nueva Vizcaja Prov. (Mar. 
3-18); Banaue, Ifugao Prov. (May 11-June 11; 
with Mendoza, see there). 

Collections. Mostly hepatics; Herb. Manila. 

*Ross, Alan E. H. 

was appointed ecologist to the Herbarium at Lae 
(Mand. Terr, of New Guinea) in April 1962. 

In 7957 he collected in the NGF series in Jimmy 
Valley, Western Highlands, near Mt Hagen, Terr. 
of New Guinea. In Herb. Lae. 

Royen, Pieter van 

Add: From 1962-65 Botanist-Ecologist at Lae, 
Territory of New Guinea. He left on completion 


of his contract to Australia in June 1965. Since 
May 1967 Curator of the B. P. Bishop Museum 
Herbarium at Honolulu, Hawaii. Author of many 
papers on the New Guinean flora; engaged in 
writing a mountain flora of New Guinea. 

The genus Vanroyena Aubrey, has been named 
after him. 

Collecting localities. Add: 1954. Singa- 
pore L (Mar. 4), Labuan I. (8).— 1955. P. Wai- 
geo.^ — 1961. Luzon (1 week in Mountain Prov.). 
W. New Guinea: Hollandia, Cyclops Mts, SE., S., 
and SW. part (May 23-Aug. 23, partly with H. 
Sleumer and J. Dukstra).' — 1962. Terr, of New 
Guinea: McAdams Park, Kauli Creek, Wau (June 
13); Lake Wanum-Wau (July 5); Oomsis logging 
area, Wau (July 9-11); Aiyura area (12); Wau, 
Bulolo area (Aug. 20-23); Keglsugl area, Lengagl 
Creek, Lake Aunde, Mt Wilhelm to the summit 
(Sept. 17-27); Edie Creek area, Wau (Oct. 9-12); 
Kauli Creek, Wau (Nov. 14-15); various localities 
around Lae. — 1963. Around Lae, Umi R. area, 
Adzare Range, Leron R., etc.; Papua: Rouna Falls, 
Sogeri Plateau (Jan. 24-25); Terr, of New Guinea: 
Rawlinson Range and Saruwaket, W of the main 
ridges (Feb. 20-Mar. 2), with Mr and Mrs Millar, 
climbing Mt Enggom too; Markham R. near 
Bridge (Mar. 17), Lae Botanic Garden (18-22); 
Lae-Mumeng Plateau-Bulolo (27); Nadzap Air- 
strip N of Lae (Apr. 28); Adzare Range (May 4); 
Lae Botanic Garden; Edie Creek, Wau, Kaindi, 
Kauli Creek (21-25); Lae Bot. Gard. (June 6); 
Adzare Range (13-16); Lae to Voco Point along 
Solomons Sea (18); Oomsis logging area (19); 
Adzare Range (20); trip to Bougainville I. (June 
20-July 8), on the way to and from making a trip 
in Gazelle Peninsula, New Britain; trips around 
Toiumonapu Plantation to Crownprince Range 
(June 28-July 4); New Britain, Vulcan I. between 
Rabaul and Kokopo (July 7), S of Kerevat (7), 
Matupi volcano (8);* Adzare Range and Lae Bot. 
Garden (15-31); Oomsis logging area to Markham 
Bridge (Aug. 11). Trip to Mt O'Dan (Sept. 2-9): 
by car from Lae to Goroka, Kundiawa, and 
Nondugl; northward to the Wahgi-Jimmy Divide 
following the Warrapuri R.; Mt Bangeta (13,500 
ft),» Sarawaket Range (Nov. 1).— 1964. Lae- Wau 
road (June 16-Aug. 4); Markham R. — Lake 
Wanum, Red Hill Swamp, Adzare Range, Lae 
Bot. Garden (Aug. 29-Sept. 9); Buzu R. (Oct. 27); 
Markham Point Ridge (Nov. 10 & 17).— 1965. 
Papua. Trip to the Upper Vanapa Valley (Jan. 7- 
26), Murray Pass (Wharton Range) and Mt Albert 
Edward, W. summit (Mt Biota) (Jan. 7-26): 
Woitape-Kosipi track (7-12), Mt Kumrne-Mt 
Amorwange track up to summit (13-17), Woitape- 
Murray Pass track (18-26, camp at Avios, 9000 ft), 
Murray Pass and Tsjidibombo-Mt Biota (22-23; 
1 1,500 ft).i» 

Collections. Add: Herb. Lae: New Guinea 
plants {1962-65), NGF series 15003-30178, usually 
in 8 or more specimens, distributed following the 
usual scheme, 1st set to Leyden, 2nd to Brisbane, 

Literature. (2) Add: 'De botanische ex- 
ploratie van Nieuw-Guinea in de jaren 1954 en 
1955' (Schakels NNG, 38, 1960, p. 28-35, 3 fig.). 
(3) Add: 'Notes on the vegetation of South 
New Guinea' (Nova Guinea, Bot. no 13, 1963, p. 
195-241, pi. 13-16, map). 



Cyclopaedia of Collectors — Supplement II 


(5) Add : 'A new Siaurogyne (Scroph.) and some 
Acanthaceae from New Guinea' (Nova Guinea n.s. 
8, 1957, p. 129-155, fig. 1-7); 'Coombea, a new 
genus of Rutaceae' (Nova Guinea, Bot. no 2, 1960, 
p. 9-12, 1 fig.). 

(6) P. VAN Royen: 'Vegetation of some parts of 
Waigeo Island" (Nova Guinea, Bot. no 5, 1960, 
p. 25-62, 5 fig., 5 pi.). 

(7) P. VAN Royen: 'An outline of the flora and 
vegetation of the Cycloop Mountains' (Nova 
Guinea, Bot. no 21, 1965, p. 451^69, fig. 1, pi. 

(8) P. VAN Royen: 'Blanche Bay Vulcanoseres' 
(Trans. Papua and N.G. Sc. Soc 1962, p. 12-15, 
1 map). 

(9) P. VAN Royen: 'Sketch of the alpine vege- 
tation of Mount Bangeta' (Trans. Papua and N.G. 
Sc. Soc. 1964, p. 14-20, 2 pi.); 'The genus Papuzilla 
(Brassicaceae)' (Nova Guinea, Bot. no 19, 1964, 
p. 427-433, 2 fig.). 

(10) P. van Royen: 'Notes on the vegetation of 
Mount Biota (Papua)' (Acta Bot. Neerl. 15, 1967, 
p. 530-534). 

RRI nos, series of the Rubber Research Institute, 

RSNB nos, of the Royal Society North Borneo 
expeditions to Mt Kinabalu (see Corner). 

RSS nos, of the Royal Society Expedition to the 
Solomons (see Corner). 

Riimke, Christian Ludwig 
(d. 1964, Utrecht, Netherlands). 

Rumpf, George E. (called Rumphius) 

Literature. (1) Add: For data on the 
drawings see also Cl. Nissen, 'Die botanische 
Buchillustration' 1951, vol. 2, p. 157-158. 

Biographical data. Add: H. C. D. de Wit 
(ed.): 'Rumphius Memorial Volume' (Amsterdam 
1959); Biologist 47, 1965, p. 42-54, 3 fig. 

*Rundi, Masirom, see Forest Dept North Borneo. 

Rut(;crs, Abraham Arnold Lodewijk 
(d. 1966, Wassenaar, Netherlands). 

Rattner, Franz 

(d. 1961. Lunz, Austria). 

Biographical data. Add: Hydrobiologia 
20. 1962, p. 193-201, photogr., bibliogr.; Arch. 
Hydrobiol. 58, 1962, p. 244-251, portr., bibliogr. 

•RiJRersma, Miss van 

collected a Cyperacea s.n. at Bogor, Java (July 
17. 1929). 

•Rijswijck, Olga van 

( 1 940, Brussels, Belgium ; x) studied anthropology 
and African Administration at the University of 
Witwalcrsrand, Johannesburg, Republic of S. 
Africa (B.A. 1962), and took her F'h.D. in An- 
thropology at the Australian National University 
(1967). She paid two visits to New Ciuinca (July- 
Dec. 1963 and March J964-Apr. 1965), sponsored 
by the Australian National University. She studied 
a resettlement of the mountain Kuni people (Ccntr. 


Distr. Papua) for her Doctoral thesis, and besides 
the resettlement of Nakanai peoples of New 
Britain. 1 In the latter island she stayed in Nakanai 
(Oct.-Nov. 1964). 

Collections. In addition to a full collection 
of Kuni artifacts, a few birds of paradise and 
botanical specimens were collected.'- In Ethno- 
graph. Mus., Dept Anthrop., Monash Univ., 
Melbourne, and part-collection at Ethnograph. 
Collect. (Inst, of Anatomy) Canberra. Botanical 
specimens (about 20) comprise dried leaves, 
medicinal herbs and odoriferous plants from the 
Central District. They are not numbered nor 

Literature. (1) Author of 'The Silanga 
Resettlement Project' (in New Guinea Res. Bull. 
10, 1966). 

(2) Cf. Pac. Sc. Ass. Inf. Bull. 19, no 4, 1967, p. 

*Saakov, Dr Semen G. 

(1903, Ibilissi, Georgia, U.S.S.R.; x) was 
educated in an Agronomical Highschool ; professor, 
Dr of Biological Sciences in the Komarov Botanical 
Institute of the Academy of Sciences of the 
U.S.S.R., Leningrad. 

He made expeditions for the investigation of 
tropical plants in Indonesia, Cuba, etc. 

Collecting localities. Indonesia. Lesser 
Sunda Islands: Komodo,^ Padar, Rindja, Kode. 

Collections. Dried plants in Herb. Bog. 
Seeds and plants for cultivation in the glasshouses 
at Leningrad. 

Literature. (1) Author of 'On the Flora and 
Vegetation of the island of Komodo in Indonesia' 
(Botan. Zjurn. 50, 1965, p. 1185-1198, 5 photogr.; 
in Russian). 

Sachse, Frans Jonathan Pieter 
(d. 1962, The Hague, Netherlands). 

Sainuddin bin Bolhassan, see Forest Dept Sarawak. 

Salverda, Zeno 

Add: Later he went to Surinam; in 1958 ap- 
pointed Inspector Staatsbosbeheer, Utrecht. 

Salvoza, F. M. 

Add: Author of 'Lexicon of Philippine Trees' 
(For. Prod. Res. Inst. Bull, no 1, 1963, 136 pp.). 

*Sam, Patrick P., see Forest Dept North Borneo. 

*Samat, A. 

collected mainly ferns (about 15) in Malaya, 
1961-69, at least in Langkawi, and on Bt Anak 
Takun and Bt Takun in Selangor. 

'Samsuri bin Ahmad 

(1936, Singapore; x). Plant Collector at the 
Singapore Botanic CJardens since 1968. 

Collecting localities. 1962. Malaya: 
Accompanying HiiRrr & Woods to lU Choping 
(Apr. 23-27), Bt Hintang (28). and from Apr. 29- 
May to P. Umgkawi, Dayang Hunting. Ci. Raya, 
Tclaga Tujoh. With UNESCO Limestone l-x- 
pcdition to Ikrtam, Gua Musang. Gua Panjang 
and Ulu Kclantan (July 22-Aug. 20). G. Jcrai 
(Nov. 9-13). G. Pulai (24); G. Lcdang (Dec. 2-9). 



Flora Malesiana 

[ser. I, vol. 8^ 

—1964. Sabah (N. Borneo): Mt Kinabalu, joining 
the Roy. Soc. Expedition (see Corner and Chew 
Wee Lek) (Jan.-May).— 1965. Malaya: G. Rapet, 
Tambun limestone, G. Pondok, Batu Tukang, Kp. 
Golok, Gua Puteri, led by Chin See Chung of the 
Univ. of Malaya (Mar. 9-14); with H. Singh in 
Maxwell's Hill areas in Taiping (May), Penang 
Hill area (15-17). Sarawak: G. Matang and Niah 
Cave with A. G. Alphonso (May 31-June 9).— 
1970. Malava: Sg. Juasseh, Labis, with Heaslett 
(June); G. Panti &. Sg. Panjang, Labis (July); Sg. 
Chabang Dua, Labis (Sept.); Jalan Mersing, 


Kluang (Oct.).— 1971. Ulu Sg. Punkin, Lesong 
F. R., SW. Pahang with Whitmore (Feb. 16-20).— 
1972. With Heaslett to Ulu Sg. Segamat, G. 
Chabang Tiga, G. Besar (Feb. 17-22); P. Tioman, 
Bt. Penak (Sept. 17-18); with Heaslett to G. Panti 
East (Oct. 18).— 1973. With Dransfield and 
Heaslett to G. Panti East (Jan.). 

Collections. Herb. Sing.: > 792, with his 
own serial numbers. Dupl. distributed to Kew, 
Leyden, Am. Arbor., Lae, etc. 

SAN nos, see Forest Dept North Borneo. 

Sanchez, Mateo 

Add: Jesuit priest and graduate student in 

Sanderman Olsen, Sv.-E., see sub Olsen. 

*Sands, Martin J. S. 

B.Sc. (1959), from Kew, accompanied Dr A. C. 
Jermy (see there) on an expedition to Central 
Celebes and S. New Ireland at the end of 1969. 
Especially charged with the collecting of Begonias 
and other phanerogams. Living material? 


Santos, Jos6 Vera 

retired in February 1973 and plans to accomplish 
his projects on botanical research. 

""Sanusi bin Sarih 

collected in Malaya at least in 1963, 1965 
(Taiping, Perak), and with Dr P. Grubb in Johore 
(Sept. 20-25, 7970) orchids from G. Belumut 
(Sept. 28). 

*Sanusi bin Tahir 

collected in Sarawak, at least in 7957. 

*Satake, Yoshisuke 

(1902, Yuzawa-machi, Akita Pref., Japan; x), 
botanist, graduating from the University of Tokyo 
(1928), taking his D.Sc. in Tokyo in 1937. Curator 
of the Nat. Sci. Mus. Tokyo 1939-67. 

He collected in the Napan District, Vogelkop, 
W. New Guinea, March-May 1943.^ His party 
included S. Ijiri and T. NiImura (see there). 

The palm genus Satakentia H. E. Moore has 
been named after him. 

Collections. In Tokyo Sci. Mus. Herb.: 
about 700 sheets of flowering plants and ferns. 

Literature. (1) Y. Satake: 'Journal of 
Researches in West Irian' (1943). In Japanese. 

Saunders, John Campbell 

Itinerary. Add: 7957. Central Highland. — 
7955. Lower Ramu River. — 7959. Lower Sepik 
R. — His last collection in 7962 in Central District, 

*SaurwaIt, F., see Boswezen Nieuw Guinea. 

Sauveur, Emile Gaston 

Add: He was in the employ of Netherlands 
Indian, later Indonesian Forest Service. He was in 
Java, Borneo (Pleihari, 1965), W. Sumatra, and 
Timor (coll. no 111 near Oesau, Nov. 2, 1960). 
On leave in the Netherlands in May 1970, after 
which he returned to Indonesia. In the years 
1971-73 he collected plants in the Jalimo Valley, 
east of Baliem, West New Guinea {Irian), partly 
together with G. Sinke; numbered from 2500- 

*Sayers, C. David 

former staff"-member of the Herbarium of the 
Division of Botany at Lae, Territory of New 
Guinea. He was seconded to the British Museum- 
Newcastle upon Tyne University Expedition,^ to 
make a dried collection of flowering plants and to 
collect plants of horticultural interest both for the 
tropical Botanic Gardens at Lae and for scientific 
collections in the U.K. of an unknown part of the 
Finisterre Mountains. 

Collecting localities. 1963, See sub 
Hartley. — 1964. Terr, of New Guinea: recon- 
naissance into the foothills of the Finisterre Range 
with E. E. Henty; in Aitapa Subdistr. (Sepik Distr.) 
(May); SE. Papua: near Abau (Apu); Saidor 
subdistr. (Madang Distr.) and Mt Wilhelm, 
Keglsugl (Aug.); Edie Creek (Sept.). Terr, of New 
Guinea. British Museum-Newcastle upon Tyne 
University Expedition, to the Finisterre Mountains 
(end Sept.-Dec.):^ from the Markham-Ramu 
River divide at c. 1000 ft up to the Naho (Gusap) 



Cyclopaedia of Collectors — Supplement II 


R. to the highest point of the ridge on Mt Abilala 
(11,320 ft), proceeding on foot via Daimandi, 
Budemu (4200 ft), Moro (5200 ft), Sevve, Lake 
Naho (8700 ft), Freybourg Pass (9600 ft), summit 
(11,300 ft).— 1965. With the 3rd part of the ex- 
pedition to Wagau and the Herzog Mts (Jan.); 
Morobe District (Feb.); New Britain: Fullerton 
Harbour (Feb.). 

Collections. Numbered in the NGF series, 
but partly in the series of T. G. Hartley. In 
Herb. Lae and elsewhere. 

Literature. (1) A. C. Jermy &l C. D. 
Sayers: 'An Expedition to the Finisterre Moun- 
tains of Eastern New Guinea' (J. R. Hort. Soc. 92, 
1967, p. 117-128, fig. 54-55, 57-63, 1 map). See 
also sub Jermy. 

Schadenberg, Alexander V. 

Literature. (1) Add: Ethnological papers in 
Zeitschr. f. Ethnol. 1880 and 1885. 

Schebesta, Rev. Father Paul 

Add: (1887, Gross-Peterwitz, Austria; x) was 
educated at the Mission Seminary St. Gabriel at 
Modling near Vienna; Dr.Phil. of the University 
of Vienna. From 1912-16 missionary in Portuguese 
East Africa. Because of World War I he was in- 
terned and stayed from 1916-20 in Portugal. In 
1924-25 he stayed for 20 months in Malaya, 
Sumatra, and Siam (Thailand); in 1929-30 and 
1934-35 for respectively 20 months and a year 
with the Ituri pygmies in Africa; in 1938 for 8 
months in the Philippines and Malaya; in 1946 
and 1954 two expeditions to the pygmies. He is a 
member of the Anthropos Institute, member of 
the board of the Africa Institute, and honorary 
member of the London Anthropological Institute, 

In 1967 he stayed in the Mission Seminary at 

Author of numerous other books, including 
'Tanah Malaya' (Wien 1960). 

Literature. (1) Add: a.o. in Peterm. Geogr. 
Mitt. 72, 1926, p. 253-257, map. 

Biographical data. By courtesy of the 
Dutch Radio Broadcasting on Mission Work. 

•Schiefenhcivel, Wulf 

(1943, Siegen/Westfalen, Germany; x) medical 
student (1963-65 Munich University) was in E. 
New Guinea as a member of a small scientific 
expedition conducted by Dr Th. Schultze- 
Wkstrum, zoologist at Munich,' in 1966. His 
interest was native medicine. 

After his return in Germany he continued his 
study at the University of Eriangen. 

Collecting localitics.' 7966 (Jan. -Sept.). 
E. New Guinea, Papua: delta Kikori River in the 
Gulf of Papua, based at Aird Hill (4 months, 
Jan. -Apr.); lower reaches of St. Joseph R., N of 
Yule I.. ( cntr. Distr., based at Maria F>a Plan- 
tation near Lpo (2 months); Terr, of New Guinea: 
15 km NNW of Ml Bosavi, Western Highlands 
Distr.. based at Bosavi Station near Didissa (I 
month). -1970. Papua and 7 err. of New Guinea 

CoLLEfTioN!?.' 1 50 mcdicina! and other useful 
plants in the course of his work on the medical 
knowledge and practice among the indigenes of 


New Guinea; material obviously mostly sterile. 
Many fragmentary and sterile suitable specimens 
in Herb. Lae. 

Literature (1) Cf. Fl. Mai. Bull, no 22 
1968, p. 1540; Th. Schultze-Westrum : 'Ergeb- 
nisse einer zoologisch-volkerkundlichen Expe- 
dition zu den Papuas' (Umschau in Wiss. u. 
Techn. 68, 1968, p. 295-300, map, 6 fig.); 'Neu- 
guinea (Papua-Urwelt im Aufbruch)' (Kummerley 
& Frey, Bern). 

W. Schiefenhovel: 'Medizinmann, Magier und 
Arzt' (Zeitschr. f. Mediz. Stud. & Assist. I, 1969, 
p. 20-24, 7 fig.; not botanical). 

Y. SATAKE, June 1967 

♦Schierbeek, Abraham 

(1887, Leeuwarden, Fr., Netherlands; x), 
biologist, educated at Groningcn University 
(1906-12), where he took his Dr's degree in 1917. 
From 1910 teacher in natural history in secondary 
schools, from 1912 onwards in The Hague; he 
lectured at Leyden University in didactics and 
later also in the history of biology, 1927-53. 

He was enabled to visit the Dutch East Indies 
as a 'Dcterdinger', sent out by the Jan Pielersz. 
Cocn Society (Amsterdam). He wrote numerous 
small popular papers on iiis travel impressions. 

Editor of 'Allc dc Hricvcn van A. v. Lliuwen- 
hoi.k' (I he Collected Letters). 

Itinerary. NM (2nd half). He visited: 
Sumatra Coast: Mcdan and north of it, G. 
Sibajak; Java, E of Batavia, G. Bromo (Oct.), 
liilliton, Bali, and Mailoera. 

('OLLICTIONS. I'or the main part Sumatran 
tobacco weeds presented to him by Dr S. J. C. 
Jo< MP.MS. Not numbered. In Herb. Leyden. 



Flora Malesiana 

[ser. I, vol. 81 

Schierbrand, Wolf C. von 

Add: (1807, Bautzen, Germany; 1888, Dresden, 
Germany), Colonel, with the Royal Engineers 
from 1853-?; honorary member of the Natural 
History Society Isis at Dresden. 

Biographical data. De Indische Navor- 
scher 3, 1937, p. 94, portr. 

Schiffner, Victor Felix 

(d. 1944, Baden near Vienna, Austria). 

Collections. Add: His main cryptogam 
collection was bought by Farlow Herb. (Cambr., 


Biographical data. Scient. Pharm. May 
1942, 2 pp.; Prof. Rechinger in litt. (Nov. 4, 
1970); Fl. Mai. Bull, no 25, 1971, p. 1878. 

*Schipp, William A. 

was on the staff of the Botanical Gardens, 
Darwin, Australia, at least from 1924-28; at that 
time probably in his forties. 

He made extensive collections in British Hon- 
duras,^ and previously (about 1928) collected 
plants in New Guinea, Java, and the Dutch East 
Indies. He came to San Francisco, Cal., from Java, 
late Jan. 1929. 

Collections. Through E. D. Merrill he 
obtained subscribers to his herbarium sets (prob- 
ably from Br. Honduras only). Many letters are 
present at the Univ. of California, Michigan, etc. 

Literature. (1) R. M. Lowden: 'William A. 
Schipp's botanical explorations in The Stann 


Creek and Toledo Districts, Brit. Honduras 
(1929-1935)' (Taxon 19, 1970, 831-861). 

Schlechter, Friedrich Richard Rudolph 

Collections. Also orchid dupl. in O. Ames 
Herb, and Herb. Mo. Bot. Gard.^'' 

LiTERATU re. (12) Add: P. J. B. Woods: 'The 
whereabouts of Schlechter's New Guinea duplicates 
of Liparis (Orch.)' (Not. R. Bot. Gard. Edinb. 
26, 1965, p. 361-364). W. H. Lewis: 'Schlechter's 
New Guinea duplicates of Liparis (Orch.) at the 
Missouri Botanical Garden' (Ann. Mo. Bot. Gard. 
53, 1966, p. 259^260). 

*Schmutz S. V. D., Father Erwin 

(1932, Ingolstadt/Donau, Germany; x), at first 
a druggist, studied theology from 1956-62, and set 
out for Indonesia as Steyler Missionary in Flores 
(Lesser Sunda Islands) in 1963. 

Collecting localities. From 1964 on- 
wards in W. Manggarai, Flores. — 1968. Trip to 
Timor (June-Aug.): Atapupu (June 14), Atambua, 
Atapupu (16) (Kolo Guerita), Mota Bulk (river 
between Atambua and Neniik Lalina), Lahurus 
(20), ascent of the Laka'an (1585 m; 21) and 
descent on the W. slope: Mota Oe (river; 25); 
Lalian Nenuk and Mota Telau (25), Kefamenanu 
(28), Tua Mese, direction Mutis to Oeolo (700 m; 
30); ascent G. Mutis (July 3-4; 2427 m); Soe (10; 
1200 m) and Nikiniki (12; 750m); from Soe to 
Kupang (14); to Rote (= Roti) (17-20); return to 
Kupang (staying 27-Aug. 2); to Maumere by 
plane (Aug. 4); Centr. Flores (Aug. 8-30), visiting 
Lomblem (= Lomblen) (13); return in W. Flores 
(Aug. 31). 

Collections. Private herbarium; originally 
ferns only {1964: nos 01-01 16; 1965: nos F1-F143), 
subsequently phanerogams and mosses also 
(1965-67: nos 1-1588). All specimens at least with 
one duplicate which is intended for Leyden Herb. 
(pres. 1970). Several of the numbers are lacking. 
Garcinia specimens were sent to Dr Kostermans 
at Bogor (Indonesia). In 1974: 3200 nos. 

Schneider, Gustav 

Add: (d. 1958, ?Basel, Switzerland). In 1888 
he visited Sumatra for the first time, accompanying 
the geologist and Director of the Zoological 
Museum Zurich, Prof. C. Mosch. He got his 
training at the Zoological Institute at Basel, but 
was later custodian of the musea at Kolmar and 
Miilhausen. He traded in natural curiosities, and 
lived at Basel. In later years he visited the U.S.A. 
and the Bermuda Islands. 

Literature. (1) Add: 'Die Orang Mamma 
auf Sumatra' (Vierteljahrschr. Naturf. Ges. 
Zurich 103, 1958, p. 213-286). 

Biographicaldata. Vierteljahrschr. Naturf. 
Ges. Zurich 103, 1958, p. 215-216, portr. 

*Schodde, Richard 

(1936, Adelaide, S. Australia; x) was educated 
at the University of Adelaide, graduated with 
B.Sc. Hons in Botany in 1961. Between 1958-59 
he worked in the State Herbarium of S. Australia, 
and since April 1960 he was appointed systematic 
botanist with the Division of Land Research and 
Regional Survey, C.S.I.R.O., Canberra, Australia. 



Cyclopaedia of Collectors — Supplement II Schultze Jena 

In 1967 he was working for a Ph.D. on Monimia- 
ceae in Adelaide; author of papers on Papuasian 
botany. Later he resumed his work at Canberra, 
and about 1970 joined the Wild Life section of 
C.S.I.R.O. as an ornithologist. 

Collecting localities. 1960. Mand. Terr, 
of New Guinea: with R. D. Hoogland in Morobe 
Distr. : Labu swamps near Lae (June 15), Zenag 
Hill (16); Western Highlands Distr.: vicinity of 
Tchak and Tale River valleys, near Wapana- 
manda (23-July 10), Mount Sugarloaf grassland 
area (11-24), vicinity of Kepilam, Lagaip valley 
(30-Aug. 13), Yobobos, headwaters of Lagaip 
River (16-Sept. 7).— 1961. Neth. New Guinea, 
Central Range: Wamena (Baliem valley) to foot of 
Wilhelmina Top. c. 13,000 ft, ± via Wamena 
River valley (June 1-8). Papua, Southern High- 
lands Distr.: vicinty of Kiburu, near Mendii, c. 
5500-6500 ft (June 26-July 9), Mendi, 5200 ft and 
Dimifa, 7200 ft (June 27), Ebenda, SW. foot of 
Mount Giluwe, 6500-7000 ft (July 16-28), SW. 
slope of Mt Giluwe, 8500 ft (Aug. 3), western 
summit grasslands of Mt Giluwe (Aug. 7-23 ; 26), 
western slopes of Mt Giluwe above Karel, 8300- 
9100 ft (25-Sept. 2), Enep-Dimifa, 7100 ft (Sept. 
7), around Lake Kutubu,i c. 2500-3000 ft (16- 
Oct. 9). — 1962. Papua: Brown River area near 
Port Moresby (July 1-31), Rigo area (Aug. 6-17), 
Sogeri plateau (Aug. 27-Oct. 1). — 1964. Solomons: 
Bougainville (July 16-Sept. 23). — 1966. Papua: 
Gulf Distr. (Jan. 9-Mar. 18); Aseki area, Kuku- 
kuku ranges, Morobe Distr. (Mar. 31-Apr. 29), 
with L. A. Craven. — 1969. Owen Stanley Range 
and Mt Wadimana (E. summit Simpson massif), 
middle of the year. — 1970. With A. Kanis in 
Papua: Astrolabe Range and near Efogi on the 
Kokoda Trail. 

Collections. With Hoogland in the NGF 
series (1960), Herb. Lae; c. 1150 nos (1306-2456) 
in Herb. Canb. (C.S.I.R.O.), dupl. distributed as 
those of Hoogland.* Wood-samples with the Div. 
of Forest Products, C.S.LR.O., Melbourne, Dept 
of Forests, Lae and Port Moresby, and C.S.I.R.O. 
Herb. Canberra. Bougainville collections ser. 
3581-4131; 1966 N. G. coll. nos 4200-5090 were 
made with the assistance of L. A. Craven. He 
collected in Australia and Tasmania too. 

Literature. (1) R. Schodde & W. B. 
Hitchcock: 'Report on the birds of the Lake 
Kutubu area, Terr, of Papua and New Guinea' 
(Div. Wild Life Res. CSIRO, Canberra, Tcchn. 
Paper 13, 1968, p. 1-73, 5 fig.). 

(2) E. B. Bartram: 'Mosses of the Western 
Highlands of Eastern New Guinea' (Rev. Bryol. 
Lichen, n.s. 30, 1961, p. 185-207). 

•Schonian, W. 

of Bremen, Germany, made a geological 
expedition in NW. New Guinea in 1929. At least 
part of his plant collcclion in Herb. Berl. where 
special labels were printed 'Cir. Gcllvink (=^ 
Gcclvink)-Bai. Bcrgc siidl. von Nabire Trop. 
Urwald; 100 400 m" (Ace. 10.1.1944). 

Schol-Schwarz, M. B. 

(d. 1970). 

BiooR aimiical data. Jaarb. Vcrsl. & Mcded. 
Kon. Ned. Bot. Vcr. over 1969, 1970. p. 6. 


*Schouten, see sub J. J. Loeters. 

Schram, Ferdinand August Willem 

Add: He helped as field assistant during part 
(Oct.-Dec. 1961) of the 2nd Rijksherbarium New 
Guinea Expedition (see Sleumer). 

Diplycosia schramii Sleum, has been named 
after him. 

Collecting localities. Add: 1958. Beriat 
and Wersar, near Teminaboean (Apr. 18-May 5); 


Ajamoera area (Ajamoera, Semoe, Jokwer, 
Kamboeaja, Djitmao, Fan, Tehagh, Aitinjo, 
Erokwero, Joksoro, Awit, Ewai, Mos, Kami, 
Itaro, Koa, Teminaboean, Skendi) (May); W, 
part of Arfak Plain (Sidei, Wariki) (July -Aug.): 
Kebar Valley (Anisandau) (Oct. -Nov.).— 1959. 
Sarmi Hinterland (Kwansem, Sidoearsi Mts, 
Ocreb R., Abcnti, Kondirdjan) (May 12-June 
18); Biak I. (Karadori. Mas!) (Nov. Dec.).- 1960. 
Sckoli Plain (Feb.-Mar.); Wissel Lakes (May 11- 
30, sec Vink); Japcn I. (Seroci, Randawaja, 
Soembcrbaba, Aisao, Socmberbaba, Seroci) (Sept. 
20-Oct. 10).-- 1961. Warsamson Valley, H of 
Sorong (July 16 Aug. 24).- 1962. Wandainnicn 
Peninsula (Wondiwoi Mts, Wasiot) and Job I. 
(I-eb. 24-Mar. .30): Mens Norm (July 31 Aug. 
11).- 1958-62. Surroundings of Manokwari. 
C'OLLh( noNS. Add: About 2(i()0 nos. 

Schult/.e Jena, Lconhard 

fd. 1955, ?. Obcrhcsscn. Germany), from 1913- 
1937 Ordinary Professor of (iconiapliy at the 
Philipps-Univcrsitat, Marburg. When retired he 
devoted himself to the study of Mexican and 



Flora Malesiana 

[ser. I, vol. 8^ 

Central-American cultures (information courtesy 
Marburg University). 

'"Schuurman, Jan Albert 

(1888, The Hague, Netherlands; x), a garden 
enthusiast, was for many years in the employ of, 
the Dutch Foreign Service; he practised gardening 
in Canada, Louisiana, Netherlands, Java and 
Brisbane. From 1949-55 he lived at Djakarta 
(Indonesia), teaching at the Indonesian College for 
Foreign Affairs. After a short visit to Holland, 


he returned to Java, giving radio talks for Radio 
Indonesia and making himself useful in the 
Herbarium and Botanic Garden at Bogor. 

He translated popular botany books in English 
and wrote papers in horticultural magazines. 

Since 1959 settled near Brisbane in Australia. 

Collections. No 137, Otophora spectabilis, 
coll. 29-7-1957 in Hort. Bogor. In Herb. Bog; 
several dupl. distributed. 

Scott, J. 

(see Suppl. Cycl. Fl. Mai. I, 5, 1958) might be 
John Scott, officer in the employ of the Hon. 
United E.I.C. ; he sailed in the 'Admiral Gardner' 
to Bengal and Bencoolen in Sept. 1797, and in the 
'Lord Duncan' by the end of March 1801.^ 

Literature. (1) Cf. Ch. Hardy: 'Register of 
Ships employed in the service of the Hon. United 
E.I.C. etc: (London 1799) p. 92, IIL 

Seemann, Bert(h)old Carl 

Literature. (1) Add: For the itinerary in 
Alaska see Bot. Not. 1940, p. 300-301. 

(2) Add: For dates of publication, see W. T. 
Stearn in J. Soc. Bibliogr. Nat. Hist. 4, 1963, p. 


(4) Add: For details on the drawings see Cl. 
NissEN, 'Die botanische Buchillustration' 1951, 
vol. 2, p. 169. 

Biographical data. Add: In Hemsley & 
Hooker, Biol. Centr.-Amer. 4, 1887, App. p. 132. 

*SeIveraj, P. 

Assistant of the Forest Research Institute, 

Collecting localities. Malaya. 1968. 
Perak, G. Bubu (Aug. 27-30).— 7969. G. Bubu 
(June 17-20); Trengganu, G. Tebu F. R. (July 

Semper, Carl Gottfried 

Literature. (1) Add: On publication dates 
of the (zoological) scientific results, cf. J. Soc. Nat. 
Hist. Bibl. 5, pt 3, 1969, p. 144-147. 

Serrurier, Dr Cornelia 

will be M. C. Serrurier (see Cycl. Fl. Mai. I, 1, 
1950), the M. probably standing for Mademoiselle. 

(d. 1962, Nijmegen, Gld., Netherlands, at the 
age of 82). She was educated at Lausanne 
University and was a private lecturer on French 
literature at Leyden University from 1913-1934. 
In the latter year she settled at Nijmegen. 

*Setchell, William Albert 

(1864, Norwich, Conn., U.S.A.; 1943, Berkeley, 
Cal., U.S.A.) was educated at Yale and Harvard 
Universities, obtaining his doctorate in 1890. 
Subsequently he served as Assistant Professor at 
Yale and in 1895 came to the University of 
California as Professor of Botany, retiring in 1935. 
His special field of interest was algology. He made 
two round the world tours, viz in 1903-04 and 

He visited Java as a member of the 4th Pan- 
Pacific Science Congress in 7929, and evidently 
attended some of the excursions. 

Collections. General collection from Java in 
Herb. Berkeley {U.C.). Others were made during 
the round the world journeys when to my knowl- 
edge the Malesian region was not visited. 

Extensive collections of Algae. 

Biographical data. T. H. Goodspeed (ed.) 
in Essays in Geobotany in Honor of W.A. Setchell 
(Un. Cal. Press 1936), with bibliogr. and portr.; 
Science 97, 1943, p. 458; Madrono 7, 1943, p. 91- 
93, portr. 

*Seto, Ko 

pteridologist and a member of a joint ex- 
pedition of the Osaka Museum of Natural History 
and the National Museum, Manila, in the Philip- 
pines. They collected on Mt Apo in Mindanao 
(Davao) from Nov. 19-Dec. 20, 7969.^ 

Collections. Shared equally between Osaka 
Mus. and Herb. Nat. Miis. Manila. 

Literature. (1) Cf. Fl. Mai. Bull, no 25, 1971, 
p. 1891-1892. 

Shah bin Haji Mohamad Nur, Mohamad 

(1935, Singapore; x). Plant collector at the 
Singapore Botanic Gardens from 1956-58, subse- 
quently Herbarium Assistant. 

Collecting localities. 1955. Sarawak, 
with Purseglove at Kuching, Batu Kitang, Bau." 



Cyclopaedia of Collectors — Supplement II 


Setapok and Sebanding, G. Gading, Lundu, 
Sematan, Kp. Pueh (Sept. 12-27).— 1956. Sarawak. 
with PuRSEGLOVE and Gilliland to Bako National 
Park, Telok Asam, Telok Tajor, Lokosan, Sg. 
Serai, Telok Wakang, Telok Delima, Telok Lakei, 
P. Lakei, Telok Limo (May 16-22); Sana, Sg. 
Tau, Sg. Mayeng (Tau Range), Bt Mayeng, Bt 
Mersing, Tatau, Bt Buan (May 28-June 20).— 
1957. Malaya: Batu Berendam (May 6-9); 
Jenderam Estate near Nilai (Sept. 18-23). — 1958. 
Sabai Estate near Bentong (Jan. 26-Feb. 3); 
Bukit Paloh Estate (Apr. 2-9).— 1959. With 
H. M. BuRKiLL day-trip to Kota Tinggi (Feb.); 
with K.ADIM BIN Tassim to Kp. Gobek (Feb. 28- 
Mar. 8); Eraser's Hill and Pine Tree Hill with 
H. M. BuRKiLL and Mohd. Nook (Aug. 24-29).— 
1961. Sabah: With the Roy. Soc. Expedition to 
Mt Kinabalu (see Corner and Chew Wee Lek) 
(May-Aug.). — 1964. With Roy. Soc. Expedition 
to Mt Kinabalu (Jan.-Apr.). — 1965. Malaya: G. 
Hijau in Maxwell's Hill, Bt Berapit, Bt Kemunting 
and Batu Hampar in Taiping (Dec. 3-9). — 1968. 
With Whitmore to Taman Negara, Kuala Tahan, 
Sg. Teku, Sg. Tahan, G. Pondok II, G. Tuan Sket, 
Jeram Panjang, Kuala Keniyam, Bt Keliyu, Gua 
Luas, Bt Terom, Sg. Redap & Sg. Belar (Feb. 16- 
Mar. 10).— 1970. Sg. Bantan, Labis F. R., with 
Heaslett (May). With Whitmore and Mohd. 
NooR in Taman Negara: Ulu Sg. Sat, Ulu Sg. 
Sepia, Kuala Aur, Jeram Perahu, NW. Tanjong 
Bungkal (July); Ulu Sg. Sedili Kechil, Bt Tangga 
Tujoh, with R. Hill (Aug. 17-19); Jalan Mersing, 
Kahang, G. Blumut with Grubb (Sept. 19-27).— 
1972. With Whitmore to G. Rabong, Ulu Sg. 
Ketil (Mar. 7-19).— 1973. Gua Musang, Sg. Ketil, 
Sg. Kundor (Jan. 29-31); Bertam, G. Brangkat, 
G. Brong, Gua Brong, Bt Bang in Kelantan (Feb. 

Collections. Herb. Sing.: > 2800, in his 
own serial numbers. Dupl. to Kew, Leyden, Arn. 
Arbor., Lae, etc. 

Shelford, Robert Walter Campbell 
Rightly: d. at Margate, England. 
Biographical data. In Bibliogr. 

Entom. 1775-1930, 1932. 


*Shimizu, Tatemi 

(b. 1932), a member of the Kyoto University 
Biological Expedition to Thailand and Malay.via, 
1967. He paid special attention to the limestone 

Itinerary. See sub K. Iwatsuki. 

Collections.' 1st set in fferb. Kyoto: 2nd 
set in For. Herb. Bangkok. 

Literature. (1) Elaborated in 'Contributions 
to the Flora of Southeast Asia* in Tonan Ajia 
Kcnkyu (The Southeast Asian Studies) vol. 8, no 
2, 1970, and more to come. 


(cf. Fl. Mai. I, 1. 1950, p. 484b) must have been 
Jo(m)ann»s SiHER(i (1740. Rotterdam, Nether- 
lands; 1817, Batavia. Java), who sailed to Java as 
gunner's mate in ' 't Jlof Duna', Zealand Chamber 
of the li.ast India Company, in 1758. At his arrival 
he resigned and had a long career with the Com- 
pany. As a Senior Merchant (1776) he was sent to 
Padang, Sumatra West Coast. In 1780 he returned 


to Java, held several high offices, finally as Gover- 
nor General (1802-05). 

He was hot-tempered, ambitious, and unreliable. 
His marriage to a daughter of Governor-General 
Alting might account for his reaching the top. After 
his resignation he stayed at Batavia and still had a 
considerable influence for many years. 

biographical data. F. W. Stapel: 
Gouverneurs-Generaal van Nederlandsch-Indie' 
(Den Haag 1941, no xxxiii, w. portr.). 





collected with Shah and Purseglove (see there) 
in the Malay Peninsula and Sarawak (1957). 
Plants in Herb. Sing.: dupl. a.o. in Herb. Edinb. 

*Siebenhaar,W.E.F., see BoswezenNieuw Guinea. 

*Sikajat, see Forest Dept North Borneo. 

*Siha, K. 

collected at least 10 plants on Lou Island (Feb. 
1962), near Manus, Terr, of New Guinea. In 
Herb. Brisb. 


biologist of Bandung University, accompanied 
Kostermans to NE. Kalimantan. 

*Simcon, M., see Forestry Bureau. Manila. 

Simmonds. N. W. 

Add: Author of some books on bananas. 


collected Nothofagus Jlaviramea Stlen., no 69, 
at Chimbu, E. New Guinea. 



Flora Malesiana 

[ser. I, vol. 81 

^Sinanggul, H. T., see Forest Dept North Borneo. 

Sinclair, James 

(d. 1968, Kirkwall, Orkney, N of Scotland 
Add: He retired by Dec. 31, 1962, by com- 
pulsion of the Malayanisation scheme, but 
continued his work on a contract with the Singa- 
pore Government in an honorary capacity, till 
he left Singapore in May 1967. 


Collection localities.* Add: 1957. 
Malaya: Malacca (Apr. 20-21); A^. Borneo: 
Sepilok, Sandakan (June- July); Malaya: Mersing 
(Dec. 4-5).— 1958. Japan (May 13-16); Philippine 
Islands, Luzon (May 21-July 5), partly with 
Edano; Malava: Selangor, Perak and Cameron 
Highlands (Oct. 7-Nov. 5).— 1959. W. Java: 
Bogor (Feb.-Mar.), visiting Udjong Kulon, P. 
Handeuleum Sisi, and P. Peutjang (4 days), G. 
Salak, Tjibodas, also collecting in Hort. Bog.; 
Malaya: Pahang, Johore, Kelantan, Perak. — 1960. 
NW. Borneo (Aug.-Sept.) : Sarawak (3 weeks from 
July 31), and Brunei (2 weeks). — 1962. Malaya: 
Selangor (Nov. 12-20). 

During the years /949-Feb. 7967 numerous 
one-day trips in Johore, and Singapore. 

Collections.* Herb. Sing.: including 416 nos 
from Borneo 1957, and 374 nos from Borneo 1960. 
His own series runs to 10922, the last number 
dated Feb. 25, 1967. In Singapore he originally 
numbered in the SFN series, but when he had 
sufficient duplicates, one was laid aside for his 
private herbarium and given a number in his own 
series too. In some of his papers certain specimens 
are cited by both numbers. The SFN series was 
discontinued in 1959. Many dupl. from Singapore 
in Herb. Edinb. 

Literature. (4) H. N. Burkill gives 'A 
summary of all Sinclair's collecting in Asia, 
Singapore excepted', in Gard. Bull. Sing. 23, 1968, 
p. xx-xxiii. 


Biographical data. Gard. Bull. Sing. 23, 
1968, p. i-xxiii, pi. 1-3, incl. bibliogr., list of 
Sinclairan taxa, summary of collecting, portr., 
facs.; Fl. Mai. Bull, no 23, 1969, p. 1671-1672. 

Singapore Botanic Garden 

The SFN series was discontinued from Jan. 1st, 
1959, and had been in declining use for some years. 

*Singh, Hardial 

botanist of the Singapore Herbarium, collected 
together with Ang Gek Choo and Samsuri bin 
Ahmad (see there) in 1965; Selangor (15-26 Jan. 
1966), Johore (Nov. 21-26, 1966). In 1970 on study 
leave at the University of Hawaii. 

*Singh, J., see Forest Dept North Borneo. 

*Sinke, Guljaam 

(1945, Driewegen, Zld, Netherlands; x) attended 
the Horticultural School at 's-Hertogenbosch 
(Br.), and went to Irian in October 1970 on behalf 
of the Board of Mission of the Netherlands 
Reformed Church (in co-operation with the 
Rhinish Mission). He is stationed at Angguruk in 
the Jalimo Valley. 

Collecting localities. 1971-73. West 
New Guinea {Irian): Jalimo Valley, east of the 

Collections. Herb. Ley den. Partly with 
Sauveur (see there) in his series 2500-2707, and 
65 nos with his own numbers. 

Skottsberg, Carl Johan Fredrik 

(d. 1963, Goteborg, Sweden). 

Biographical data. Add: Hawaiian News- 
letter Oct. 1963, p. 99-100; Adansonia 3, 1963, 
p. 310-312; Taxon 13, 1964, p. 1-7, bibliogr. 
(onwards of 1940), portr.; Proc. Linn. Soc. Lond. 
175, 1964, p. 189-190; Biogr. Mem. Fellows R. 
Soc. 10, 1964, p. 245-256, portr., bibliogr. 

*Sleumer, Hermann Otto 

(1906, Saarbriicken, Germany; x) was educated 
in pharmacy in Munich, and subsequently studied 
botany and geology (Ph.D. Freiburg 1932). In 
1933 appointed Assistant in the Berlin Museum, 
and since 1946 Professor of Taxonomical Botany 
and Plant Geography at Berlin University, giving 
courses in pharmacognosy also. In 1949 to S. 
America as Extraordinary Professor at Tucuman 
(Instituto Miguel Lillo). Since mid-1953 on the 
Foundation Flora Malesiana staff at Leyden and 
from July 1956 on the staff of the Rijksherbarium ; 
he retired March 1971. Specialist in Ericaceae. 

With Dr P. VAN Royen he made an expedition 
to New Guinea in 1961-62. 

Itinerary. 2nd Rijksherbarium New Guinea 
Expedition, 1961-62.^ Travelling by way of Luzon, 
collecting (May 6-21) with Mrs M. L. Steiner on 
Mt Santo Tomas, Mt Data National Park, along 
the road to Bontoc-Banaue, and on Mt Polis. 
Dutch New Guinea, with P. van Royen, and J. 
DiJKSTRA of the For. Service: Cyclops Range, 
base camp at Ifar (c. 380 m; May 31-Aug. 28), 
making 5 major trips: to Ormoe with camps at the 
Faita R. (c. 990 m), below the saddle (c. 1220 m) 
on the S. slope, and to the highest summits in the 
central part (Mt Rara, 1700 m; Mt Raveni, c. 



Cyclopaedia of Collectors— Supplement II 


1880 m), between June 6 and July 5; in the W. part 
as a transect from Dozai via Baimungun creek 
(and Mt Dafonsero, c. 1570 m) to Tabiasufa — 
Cape Tanahmerah, and back to Dozai (July 29- 
Aug. 8); in the E. part 2 ascents from Kotanica 
resp. the Kujabu R. on the S. slopes of Mt Makanoi 
up to 800 m (July 17-23, resp. Aug. 16-19); 
numerous minor trips from Ifar into the lowlands 
on the S. side of the Cyclops Range on to Sabron 
and Depapre, along Lake Sentani, around Ifar, 
Kemiri, Kotanica, along the Jabau and Kujabu 
rivers, and near Hollandia (Pinksterbivak, Skyline, 
Base G), up to Aug. 28. After the departure of 
Dr VAN RoYEN (see there) to Lae in the Territory 
of New Guinea: with J. S. Womersley collecting 
in the Bulolo area (Edie Creek, Merri Creek, Mt 
Kaindi (c. 22(X) m) and several miles along the old 
army road from Edie Creek to Bulldog (Aug. 30- 
Sept. 3), and around Marafunga (2400-2500 m) 
near Goroka (Sept. 5-8); alone visiting Kundiawa 
and the Upper Chimbu R. valley up to Dengalagu 
(= Toromambuno, 2280 m) Mission Station (9- 
11), climbing the E. slope of Mt Wilhelm via 
Pengagl Creek (12), staying near Lake Aunde (c. 
3350 m) up to Sept. 16, collecting up to c. 38(X)m 
(mainly Ericaceae); from Sept. 17-22 lower part of 
the ascent near Keglsugl and along Pengagl Creek 
up to c. 2800 m. By air returning to Dutch Nov 
Guinea: Manokwari (25); base camp at Andjai 
(from Oct. 6), Kebar Valley, for the 2nd part of 
the expedition, making minor trips on the Kebar 
plain and into the lower part of the mountain 
(Mt Tobi-Nertoi area, Mt Nettoti, up to c. 100 m); 
via Amboaki R. bivouac to Soeroerem on the 
Aifat R. (2nd base camp), from where Van 
RoYEN, and F. Schram (of the For. Service) along 
the northern foothills of the Tokiri Range, the 
Aifat R. and the Ischon R., exploring Mt Kuse- 
mun (1400 m) in the central part of the W. 
Tamrau Range, returning to Andjai on Nov. 16; 
Slf.l'Mer collecting on the crest of the E. part of 
the Tokiri Range (1250 resp. 1400 m), Sept. 30 
marching along the Aifat R. westwards to the 
Waumi R., ridge W of and parallel to the Waumi 
R., and from a transitory camp at 1420 m to the 
waterdivide in the E. part of the Tamrau Range 
(Wamsuf massif) at c. 2100 m (Nov. 2); back at 
Soeroerem, camping on the N. flank of the E. 
part of Tokiri Range at 1 100 m, reaching the 
summit (c. 1550 m) on Nov. 8; return to Andjai 
for a short period of rest; camp on the S. shoulder 
of Mt Nettoti at c. 1750 m (Nov. 27); climbing the 
crest of the Nettoti Range both to the W and F 
up to 2100 m, marching over the top (c. 1950 m) 
(Dec. 3) towards the Wckari R. (r. 1550 m, staying 
2 days in the rcsthousc); finally lo Andjai (Dec. 6 
and 7); various minor trips from Andjai to the 
Kebar Plain and adjacent mountains up to KXX) m 
opposite Andjai; Dec. 21 by plane to Manokwari 
(staying 21-29), and return of Van Royi n to the 
Netherlands; with W. Vink (sec there) flown from 
Ransiki to Lake Angi Cijta resthousc 'Noordpool' 
(IH40m) on the Arfak Range (Jan. 8 9 1962). 
collecting in the marshland, slopes, especially the 
N. slopn: of Mt Mcscnuk (along the so-called 
I iilMfspad) up to the top (r. 2150m; 9 11); to 
Icsicivi on the opposite side of the lake; Mt 
Moiigrcmotti (2180 m) and return to Fcstcgii; 
ascent to the range between Lake Angi Gila and 


Lake Angi Gigi, called Koebre (or Kobre-mot, 
2300 m) and Tembrok (2400m) on Jan. 14; 2nd 
ascent of Mt Koebre (16), descending to the village 
Ukdehot-mamot (2100 m) and Surerei on Lake 
Angi Gigi (1920 m); from there exploring Mt 
Sensenemes (18, up to 2600 m), and via the SE. 
slope up to the summit (20); Mt Gwamongga (21, 
to the top, 2570 m); via the Sikohirie (2200 m) and 
the villages Tridaga and Siskado to Iray (22); 
from there making several trips, e.g. via Tidjei to 
the NE. part of Koebre ridge up to 2430 m (24), 
returning via Disrebei; to Mt Saru-mot (2350 m, 
26), Mt Tembrok via Tidjei and Koebre up to 
2510m (28); by helicopter from Iray to Mano- 
kwari (29); Feb. 7 by helicopter with Zieck (see 
there) to a camp above the Warjori R., collecting 
on an almost inaccessible ridge and steep slope; 
leaving Manokwari (9) and returning via Lae, 
Australia, Hawaii and the U.S.A., arriving at 
Amsterdam on March 21. — 1963.- In June some 
days in the Cameron Highlands (nos 4661-4675), 
Malaya. N. Borneo: Mt Kinabalu (July 25-28) 
and the Mesilau R. up to 2250 m (July 19-25), 
accompanied by Mrs Collenette, Dr H. P. Fuchs 
(Shell Co. The Hague), and Dr W. Meuer; 
Sarawak: Mt Matang (July 30-Aug. 1). Thailand 
(Aug. 15-Sept. 11; numbered Sleumer & Smitin- 
AND ±1000-1400, and Sleumer 4745-4792). 

Collections. Total collections of the ex- 
pedition amount to c. 3500 numbers with numer- 
ous duplicates; 1st set in Leyden, from where the 
dupl. are distributed, also to Herb. Edinb. The 
bulk of the collection is labelled Van Royen & 
Sleumer, nos 5625-8259 (continuation of Van 
Royen's series); in E. New Guinea the nos 
Sleumer 4125^190, in the Arfak Mts (W. New 
Guinea) the nos 4191-4490 and Sleumer & Vink 
BW nos 14001-14359; during the 2nd Arfak trip 
the nos Sleumer 4491^507. 

Literature.(1)C/.F1. Mai. Bull. «o 17, 1962, 
887-890; ibid, no 18, 1963, p. 988; H. Sleumer: 
'Rhododendron in Neu-Guinea' (Jahrb. Rhodod. 
Ges. 1963, p. 7-32, 39 photogr. 1 map, incl. 

(2) H. Sleumer: 'Rhododendron in Borneo and 
Thailand' (Jahrb. Rhodod. Ges. Bremen 1965, p. 
7-27, 26 photogr.). Cf. Fl. Mai. Bull, no 19, 1964, 
p. 1113-1114. 

Biographical data. Blumea 19, 1971, p. 
199-210, portr., bibliogr. 

Smith, Eryl Glynne 

Add: (born at Glyndyl, Bangor, North Wales, 
G.B.) took a medical degree in London in 1918, 
and came to Siam ( Ihailand) as Dr Mai < olm A. 
SMirii's wife in 1921. Besides collecting in penin- 
sular Siam, she visited Malaya, Hainan (1923), 
Cambodia (1925), and Celebes and limor (1925). 
Returned to England in 1925, afterwards working 
on ferns at Kew. 

Several plants have been named in her honour. 

Collections. Add: With Kirr's private 
herbarium in Herb. Brit. Mus. Total collections 

BiociRAPtiiCAL DATA. J. Siam Soc. Nat. 
Hist. Suppl. 8. 1930, p. 128-129. 

*Smllh, Gordon 
(1941, Abcrfcldy, Scotland; x), was educated at 



Flora Malesiana 

[ser. I, vol. 81 

Aberdeen University (B.Sc. Hons 1963, Ph.D. 
1968). From 1968 Lecturer, School of Biology, 
University of Malaya, Kuala Lumpur. 

Collecting localities. 1968. West Mal- 
aysia, mainly in Selangor, Pahang, and Perak 
States. — 1969. With E. Soepadmo in Borneo. 

Collections. Nos 150-400 in Herb. Aberd. 

♦Smith, Jeremy Michael Bayliss 

(1945, Stamford, England; x), botanist, edu- 
cated at Cambridge University, England, 1964-67 
(Upper 2nd Hon.), and University of Singapore 
1968-69 (Dipl. in Education). 

He was a member of the Cambridge Kinabalu 
Expedition, 1967; other members were J. W. 
Patterson and P. R. Williams, both reading 
Zoology in the University of Cambridge. 

He chose to make an ecological study of certain 
high altitude herbs {in casu Ranunculus lowii and 3 
species of Potentilla) so as to discover something 
of the factors controlling their distribution.^ 

He remained in the East, working as a demon- 
strator in the Dept of Biological Sciences, at the 
University of Malaya. Now in Canberra, School of 
Pacific Studies, A.N.U. 

Collecting localities. Cambridge Kina- 
balu Expedition, 1967. Two months in Sabah 
during the summer vacation. Base camp at Park 
Headquarters Kinabalu. Smith spent the first 
three and a half weeks close to the summit, making 
use of Panar Laban mountain hut (11.000 ft). The 
three of them made an ascent to Low's Peak 
(13.455 ft) and subsequently returned to Park H.Q. 
The second half of the expedition the eastern track 
to King George's Peak (13.340 ft) was followed, 
partly different from the one followed by Corner. 
Some time was spent at Tuaran before leaving 
Borneo and paying a visit to Singapore and the 
University of Kuala Lumpur. — 7965. Mt Kina- 
balu, East Summit Party (Apr.),^ with J. B. 
LowRY, A. HiNDE, G. Jacobsen, Dr J. Sheridan, 
and Miss R. Barker. 

Collections. ? Cambridge, and ? Herb, 
Kuala Lump. 

Literature. (1) J. M. B. Smith: 'Herbaceous 
plant communities in the summit zone of Mount 
Kinabalu' (Mai. Nat. J. 24, 1970, p. 16-29, 2 fig., 
2 tab., 1 map, pi. 6-7). 

(2) J. M. B. Smith & J. B. Lowry: 'Further 
exploration and observations on Mount Kinabalu 
East' (Mai. Nat. J. 22, 1968, p. 29^0, 3 maps, pi. 

Smith, John Swire 

Retired as Director Forest Service, Malaya. 

Biographical data. Mai. For. 23, 1960, 
p. 84. 

♦Smith, J. W. 

collected Vitex siamica Will, from G. Baling 
Kedah, Malaya (July 19, 7956). 

Smith, Lindsay Stuart 

(d. Sept. 12, 1970, Mt Barney, W. of Brisbane, 
Queensland, Australia). 

Biographical data. Courier-Mail, Bris- 
bane, Monday, Sept. 14, 1970, p. 1; Fl. Mai. Bull. 
no 15, 1971, p. 1878. 


Smythies, Bertram Evelyn 

Add: Served in Brunei up to 1959, and was 
appointed Conservator of Forests in Sarawak in 
September 1959; retired in 1964. 

He recruited and looked after a team of Iban 
tree-climbers to whose efforts many valuable tree 
collections are due. He settled in Spain. 

He has a large interest in ornithology and is the 
author of books' and papers on birds. 

Parashorea smythiesii Wyatt-Smith ex Ashton 
has been named after him. 

Collecting localities. NW. Borneo. Prior 
to March 1957 in Brunei and in the neighbouring 
Limbang and Lawas Districts of Sarawak. — 1957. 
Temburung Distr. (Mar. 15-Apr. 10, with G. H. S. 
Wood and P. S. Ashton); Bt Sagan (535 m), 
Limbang Distr. (Apr. 19-20); Andulau F. R., 
Belait Distr. (23-27 with Wood and Ashton); 
Labi road and Bt Teraja (417 m), Belait Distr. 
(May 16-23, with Ashton); Kuala Ingei, Ulu 
Belait (July 5-12, with Ashton); Ulu Temburong 
(Oct. 31-Nov. 8, with Ashton); Andalau F. R. 
(Dec. 4-8, 24-27).— 1958. Sg. Iran, Ulu Sg. 
Pelagus, Kapit Distr. (Sept. 14-16); Sg. Medamit, 
Limbang Distr. (Dec. 9-17).— 1959. N. Borneo: Mt 
Kinabalu along normal route from Tenompok to 
summit (Feb. 25-Mar. 12, nos S 10601-10650 and 
10951-10959).— 1960. Sarawak: Mt Matang 
(910 m) near Kuching (Mar. 13, Apr. 3, May 29, 
July 4), G. Gading (884 m) Lundu Distr. (Apr. 
12-14); Tiang Bekap on Padawan Rd, Kuching 
Distr. (June 3-6, with J. A. R. Anderson); Sg. Sabal 
Tapang, Serian-Simanggang Rd (Aug. 4-9, with 
J. Sinclair and J. A. R. Anderson); G. Berumput 
and G. Kanyi (1554 m) in Pueh Range, Lundu 
Distr. (Aug. 25-30); Sg. Lajang, Sg. Bena (Ulu Sg. 
Sut), Kapit Distr. (Oct. 2-7).— 1961. Mt Matang 
and Mt Selang area, near Kuching (Sept. 29 with 

E. J. H. Corner, Oct. 24-27); Sg. Sabal Tapang 
on Serian-Simanggang Rd (Oct. 16-18); G. Pueh 

F. R. at Sungei Sebat Kechil near Sematan 
(Nov. 4-10). N. Borneo: Mt Kinabalu, Hot 
Springs area with Roy. Soc. Expedition (July 4— 
24).— 1962. With B. L. Burtt and P. J. B. Woods 
to Bako National Park (May 17-20), Niah cave 
(June 3-11), Pelagus Rapids area Kapit Distr. 
(July 19-24). 

Collections. Specialising in Dipterocar- 
paceae, Nepenthaceae, and Ericaceae, and all 
trees attaining timber size. Brunei, Limbang and 
Lawas collections in Herb. Brunei, dupl. distributed 
through Kepong. Collections 1957-59 jointly with 
P. S. Ashton and distributed under Brunei nos. 
From Sept. 1959 onwards in Herb. Sarawak. 

Literature. (1) 'The Birds of Burma' 
(Edinburgh 1953); 'The Birds of Borneo' (Edin- 
burgh 1960). 

*Soares, F. A. 

collected 71 specimens of grasses in Portug. 
Timor in 7967; in Herb. Lisbon.^ 

Literature. (1) Cf. Margarida de Castro: 
'Contribuicao para o conhecimento das Gramineas 
de Timor' (Garcia de Orta 12, 1964, p. 45-66; 
see p. 45). 

*Soegeng Reksodihardjo, Dr W. 

In 1967/68 he taught botany in Honolulu 
University, Hawaii. Then Assistant Director of the 



Cyclopaedia of Collectors — Supplement II 


Seames project BIOTROP, the Southeast Asia 
Centre of Biology; in 1972 he had resigned. 

Collecting localities. 7966. With Kos- 
TERSiANS (see there) to New Guinea. — 1967. With 
Nedi in Irian, as a member of the border ex- 
pedition with Australian New Guinea (see sub 
Ridsdale). — J970. E. Kalimantan. A joint 6-weeks 
sur\'ey to Kutai Nature Reserve (May), sponsored 
by BiOTROP. 

Collections. Presumably in Herb. Bog. 

Soejatmi, see Soenarko. 

•Soenarko, Miss Soejatmi 

agrostologist at Bogor/ about 1972 she received 
a grant for attaining a Ph.D. at Reading, England. 
She made field trips in W. Java {1970) with 
KosTERMANS and J. Dransfield. 

Literature. (1) Author of: 'Rumput* di 
Pulau Peutjang (Udjong Kulon)' (Berita Biol. I, 
pt 3, 1971, p. 8-11). 

*Soepadmo, Engkik 

(1936, Java, Indonesia; x), educated at the 
Tjiawi Academy of Biology (B.Sc. 1959), and since 
then Assistant Botanist of the Herbarium Bogo- 
riense. He followed a field training course in 
Sarawak under Dr Anderson. 

For some years, till 1966, working under 
supervision of Prof. E. J. H. Corner at Cambridge 
on a revision of the genus Quercus for a Ph.D. In 
1966-68 he was at Leyden to work on the Fagaceae, 
resulting in a revision for Flora Malesiana. 

In 1968 appointed Lecturer in tropical plant 
taxonomy at the University of Malaya, Kuala 

He published a monograph of the genus 
Xeesia, and is now working on Ulmaceae. 

Collecting localities. 1958. Bali (June 
10-July 14);' W. and Centr. Java (Oct. 29-Nov. 1): 
Pangandaran, Nusa Kambangan, Tjikatok, Pend- 
jalu; W. Java (Nov. 8-15); Peutjang /., Udjong 
Kulon.— /959. W. Ceram (June 1-26); Ambon 
(July 5-Oct. 8); W. Java: Bogor Bot. Card.; 
Tjibodas (Dec. 23). — 1960. Sumatra East Coast: 
Pakanbaru, Tenajan R. (Aug. 10-27).— 796/. Sum. 
E. C: Sibolangit (Apr. 11-19); Centr. Sumatra: 
Pakanbaru (Aug.-Sept.); Singapore /.: Bt Timah 
(Nov. 5-6); Peutjang I. (Dec. 16-28).— 7969. In 
Borneo with G. Smith. — 1970. Malaya. G. Tahan 
expedition (Aug. 26-.Sept. 8)^ from Kuala Tahan; 
summit .Sept. 5. — He will occasionally continue 
collecting in Malaya. 

Collections. Herb. Bog.: plants from Bali 
(together with Soegeng, Kuswata and Koster- 
mans, nos 1-346), Java (with ditto, nos 347-423), 
Ceram and Ambon (sec Kuswata), Java 1959 (nos 
l-ll), Sumatra (nos 12-252, 254-261), Singapore 
(nos 262-267), Peutjang I. J 961 (nos 268-331). 
Malayan plants presumably in Herb. Univ. Malaya, 
Kuala Lumpur. 

All dupl. from Sumatra I960 to Copenhagen 
where they forwarded the funds for this trip. One 
set will go to Leyden: also dupl. in Herb. I.dinb. 

Literature. (I) Cf. Pcnggcmar Alani 40, 
1961. p. 25-43. 

(2) F. SrjEPADMo: 'Plants and Vegetation along 
the path, from Kuala lahan to CJunong lahan' 
(Mai. Nat. J. 24, l'>»71. p. 118-124, pi. 33a 40b). 


Solander, Daniel Carl 

(rightly: b. 1733). For additional literature on 
Cook's first Voyage see sub Banks in this Supple- 

Biographical data. Add:R. A. Rauschen- 
berg: 'Daniel Carl Solander naturalist on the 
Endeavour" (Trans. Amer. Philos. Soc. Philad. n.s. 
58, pt 8, 1968,66 pp.; r/. also book notice by A. M. 
Lysaght in J. Soc. Bibliogr. Nat. Hist. 4, 1970, 
p. 329-331). 

SUl. I'AUMd 


Sonncraf, Pierre 

Lrn rature. (1) Add: Cf. also M. Ly-Tio- 
Fane in 'Pierre Poivre et I'cxpansion fran(; 
dans rindo-Paciliquc' (Hull. Lcolc Frani;. Lxtr.- 
Or. 53, 1967, p. 473-478). 

Soriann, J(wentino D. 

a M.S. Botany graduate. U.P.; Ph.D. in genetics 
in U.S.A. Conncclcd with liie Philippine Dcpt of 
Botany, College of liberal Arts, U.P. 

(oM.KMNii locAiims. Add: Mindoro: 
Calapan; Luzon: Ki/al Prov. 

Sow l)in I'andang 

(d. Jan. 25. 1971) was also in Sabah. Sarawak, 
and Brunei; retired in March I9()9 after 42 years 
service as plant collector at Kcpong (Malaya). 

Hio(iRAPni< AL DArA. Mill, l-orcstcr 33, 
1970. p. 282. 



Flora Malesiana 

[ser. I, vol. 8^ 

Spare, Gordon H. 

Collections. Add: Also dupl. in Herb. 
Edinb., Malay Peninsula 1938. 

Sparrman, Andreas (or Anders) 

He returned to the Cape Colony when Cook's 
2nd Voyage touched again at Capetown.* 

Literature. (2) Add: A. Sparrman: 'Un 
compagnon Suedois du Capitaine James Cook. 
La coquille qui chante' (Oslo 1939; transl. from 
Resa till Goda Hopps-Udden ... 1783, 1802). 


the late chief tree climber at Kepong, 1967. 

(4) In Suppl. Cycl. Fl. Mall I, 5, 1958 add: 
H. St. John's statement (Natur. Canad. 98, 1971, 
p. 563) that Sparrman was asked to join the 
expedition at the Cape at the instigation of the 
FoRSTERs seems doubtful with a view to the 
earlier recommendation by Ekeberg. 

Sporleder, Friedrich Wilhelm 

Add: (1787, Wemigerode, Germany; 1875, 
ditto) studied law, but had a large interest in 
natural history and geology. He had an extensive 
correspondence with Kunze, Sprengel, Slechten- 
DAHL, and others, and owned private collections, 
including 200 portfolios of dried plants, which 
were bequeathed to the royal collections at 


Wernigerode. The Java plants (cf. Fl. Mai. I, 1, 
1950, and ibid. II, 1, 1963, p. 159) were certainly 
not collected by himself. 

Biographical data. Allg. Deutsch. Biogr. 
35, 1893, p. 275-277. 

*Stainton, John David Adam 

(1921, London, England; x) was educated at 
Winchester College and Christchurch Oxford. 
Hons degrees in History and Law; barrister-at- 
Law. Assistant in the organization of the Royal 
Society North Borneo Expedition 1961. Plants 
were collected with Chew and Corner (see there) 
in the RSNB series. His real interest is Himalayan, 
but he collected in the Balkan and Greenland too. 

Collections. For Borneo see under Corner. 

♦Start, A. 

collected Mucuna biplicata T. & B. from Bukit 
Anak Takun, Selangor, Malaya (Jan. 23, 1972). 

*Stauffer, Hans Ulrich 

(1929, Aarau, Switzerland; 1965, Aarau, 
Switzerland), botanist educated at the University 
of Zurich (Ph.D. 1959); later University Lecturer 

He visited New Guinea when on his world tour 
(1963-64) for herbarium and field study of the 

Collecting localities. 1963. E. New 
Guinea: Lae, Bulolo, Kainantu, from Keglsugl 
to the lakes E of Mt Wilhelm, Port Moresby, 
Sogeri. He then proceeded to New Caledonia, Fiji, 
and Hawaii, returning to Switzerland in Aug. 1964. 

Biographical data. Fl. Mai. Bull, no 20, 
1965, p. 1234-1235; Adansonia 6, 1966, p. 23-24; 
Mitt. Aargau. Naturf. Ges. 27, 1966, p. 279-286, 
portr., illustr. ; Vierteljahrschr. Naturf. Ges. 
Zurich 111, 1966, p. 457-458. 

Steenis, C. G. G. J. van 

Add: In 1959 he was awarded a honorary Dr's 
degree at the Universite de Montreal. From 1962- 
72 Director of the Rijksherbarium, Leyden. 

Literature. (40) Add: R. Geesink: 'Two 
new species of Swertia (Gentianaceae) from N. 
Sumatra' (Blumea 20, 1972, p. 132). 

Biographical data. Blumea 20, 1972, p. 
1-6, portr.; ibid. p. 7-24, 6 fig.; Trop. Ecol. 13, 
1972, p. 1-4, portr. 

Stefels, Casper Jacobus 

Add: Later employed in the Dutch Forestry 
Service, stationed at Zwolle (O.). 

Collecting localities. 7955. Anggi Lakes 

Literature. (1) C. J. Stefels: 'Orientatie- 
tournee Bosverkenning Ransiki-Anggi-meren' 
(Manokwari, 1955; typewritten report). 

Itinerary with annotations by W. Vink in Nova 
Guinea, Bot. 22, 1965, p. 490-491, fig. 4. 

Steiner (nee Lindenberg), Mona Lisa 

Add: made an exploration in Mt Pulog area 
(Mar. 1961), revisiting it in May of the same year 
with H. Sleumer and P. van Royen (see those). 

The family later returned to Europe. 

Collections. Add: Dupl. Philippine Is. 
{1961) in Herb. Edinb. 



Cyclopaedia of Collectors — Supplement II 


*Steinkraus, W. 

plants (7965). 

Herb. Edinb.: New Guinea 

*Sterly, Joachim 

(1926, Bremen, Germany; x) took his Ph.D. at 
the University of Cologne, Germany (1963), and 
subsequently got a grant from the 'Deutsche 
Forschungsgemeinschaft' (1963-66).' Founder and 
chairman of the 'Arbeitsgemeinschaft Ethno- 
medizin' (1970), and Editor of the periodical 
'Ethnomedizin' from 1971 onwards. From 1970-73 
he was once more awarded a grant and made a 
study of ethnobotany, medicinal plants and 
medicinal sociology in the New Guinean high- 
lands.* End 1973 appointed lecturer of ethnology 
and theoretical anthropology at the University of 

Itinerary. 1971. Territory of Papua and New 
Guinea (June 1-Dec. 30): Port Moresby, Lae, 
Madang, Rabaul (^New Britian), Goroka, Mt 
Hagen, and Kundiawa. Collecting was done in 
Chimbu Valley, Chimbu Distr., Central Highland, 
viz: in Lower Chimbu, territory of the Kamanuku 
tribe (environs of Kundiawa and Porol Range), 
and in Upper Chimbu, territory of the Kuglkane 
tribe (environs of Womatne south of Gembogl). 

Collections. About 300 useful plants of the 
Kamanuku and Kuglkane, Herb. Hamb. 

Literature. (1) His bibliographical study 
resulting in 'Heilpflanzen der Einwohner Melan- 
esiens' (Hamburg 1970). 

(2) J. Sterly: 'Krankheiten and Kranken- 
behandelung bei den Chimbu im Zentralen 
Hochland von Neu-Guinea' (Buske Verlag 
Hamburg 1973). 

•Stern, William Louis 

(1926, Paterson, N.J., U.S.A.; x), botanist, 
educated at Rutgers University (1950, B.S.), Univ. 
of Illinois (M.S. 1951), Ph.D. 1954. Instructor, then 
Assistant Professor Yale School of Forestry, 1953- 
60, Curator Div. Plant Anatomy Smithsonian 
Institution 1960-64. 

Acting Chairman of the Smithsonian Institution, 
Washington, D.C., Botany Department, onwards 
of 1964. 

F.A.O. Forestry Officer and Wood Technologist 
at FPRI College, Laguna, Luzon, 1963-64.* 

Collecting localities. Philippines. 1963. 
Luzon: Laguna and Batangas Provinces. Mindanao: 
Prov. of Agusan, Tungao, drainage of Agusan R., 
26 km S of Butuan, concession of Nasipit Lumber 
Co., Inc. (Sept. 28-Oct. I). Luzon: Prov. of 
Quezon, Maliliit na Bato, drainage of Dakil R., 
17 km E of Pangil (Alt. 1400 ft), concession of 
Intern. Hardwood and Veneer Co. (Oct. 24); 
Mountain Prov. (Nov. 25-26, coll. by and with 
Beniono a. Lomiuao). — 1964. Luzon: Mountain 
Prov., Subprov. of Bontoc, at Bauko, forests on 
slopes of Mt Data between 7000 and 8000 ft alt. 
(Feb. 10-13. with Ji;sto P. Rojo). 

Coli.p.ctions. Probably in Washington and in 
the Philippines. 

Ln hRATURE.(l) W, L. Stern ALT. Zamuco: 
'Identity of 'Tiaong' (Dipt.)* (Brittonia 17, 1965, 
p. 35 46, 5 fig., I tab.). 

BiociR APMicAL DATA. Who's who in America 
vol. 34, 1966/67. 

Steup, Ferdinand Karel Marie 
(d. 1971, Perigord, France). 

♦Stevens, Peter F. 

from Edinburgh, went to the Botany Division, 
Lae, May 1970. In 1971 he collected with M. J. E. 
Coode.* Leader of an expedition to Mt Suckling 
{Papua) in mid-7972 (see also J. F. Veldkamp and 
R. Pullen). 

Joint author on a list of plants of Mt Wilhelm.^ 

Probably he collected in the series of the 
institute; Herb. Lae etc. 

Literature. (1) R. J. Johns & P. F. Stevens: 
'Mount Wilhelm Flora. A Check List of the 
Species' (Bot. Bull, no 6 Div. Bot. Lae, 1971, 
60 + 4 pp., 7 fig., 1 map). 

(2) Authors of: 'Notes on the Flora of Two 
Papuan Mountains' (Pap. New Guin. Sc. Soc. 
Proc. 1971, 23, 1972, p. 18-25). 

Stok, Johan Ewald van der 
(d. 1958, Naarden, N.H., Netherlands). 

Stokey, Alma Gracey 

(d. 1968). 

Biographical data. Amer. Fern J. 58, 
1968, p. 145-154, 1 photogr.). 

Stomps, Theodoor Jan 

(d. 1973, Amsterdam, Netherlands). 

Biographical data. Vakbl. Biol. 53, 1973, 
p. 223-224, portr. 

*Stone, Benjamin Clemens 

(1933, Shanghai, China; x) was educated in 
California (B.A. Pomona College, Claremont, 
1954), and did one year of graduate work at 
Washington University under R. E. Woodson and 
others; Ph.D. in 1961 at Hawaii University. From 
1961-65 in the Department of Biology, College 
of Guam, finally as Professor. In 1965 moved to 
the University of Malaya, Kuala Lumpur as 
Lecturer in Botany and Curator of the University 
Herbarium. Founder of the journal 'Micronesia'. 

Author of numerous papers on Pandanaceae 
and other subjects. 

Heliconia stonei Lane is or will be named after 

Collecting localities. 1957. New Britain: 
Rabaul and vicinity (Dec). — I95S. New Ireland: 
Kavieng and vicinity (Jan.). — Malaya (from Apr. 
1965-67): Sclangor, Pcrak, Pahang, Penang (Feb. 
7966), Johore, Singapore (Apr.), Frascr's Hill 
(Apr.-May), Pcnang (May), Ipoh, Langkawi Is. 
(May /967).— In Borneo: Sabah (Mar. 7967). 
Sarawak (Apr.). — 7977. E. New Guinea: Mt 
Wilhelm (Apr.). 

CoLLEcnoNS. Up to June 1967, 7131 nos; 
1-725 from the U.S.A. and Switzerland in San 
Diego Nat. Hist. Mus.; nos 726^ 2588, 2M2 5512 
mostly from the Pacific region, of which IKi-'UlA 
in Herb, liish. Mus., 3775 5512 in Coll. Guam 
Herb., with dupl. at Leydrn, U.S., Bish. Mus. and 
a few elsewhere. The nos 5513 onwards from 
Malcsia will be distributed to Leydrn, Sing., Bish. 
Mus., Kew, etc. A few holotypcs will be deposited 
at the Univ. of Malaya llcrh. In New Guinea LAE 
530*; 1-53254 from Mt Wilhelm; Herb. Lae. 




Flora Malesiana 

[ser. I, vol. 8^ 

♦Stone, Oct. C. 

A naturalist who, about 1875^ and possibly also 
later,- visited the area near Port Moresby in E. 
New Guinea (Papua). He made zoological col- 
lections near Baxter and Laloki Rivers etc., and 
shipped or carried 8 varieties of sugar-canes to 

STONE and son 

Literature. (1) O. C. Stone: 'Letter from 
Papua' (The Queenslander, Brisb. Feb. 12, 1876). 

(2) O. C. Stone: 'A few months in New Guinea' 
(London 1880; review in Ind. Gids22, 1880, p. 145). 

(3) Cf. J. H. Warner & Carl O. Grassl in 
Hawaiian Plant. Rec. 55, 1958, p. 211, 213. 

Stonor, C. R. 

Add : In 1949 he again collected on Mt Wilhelm 
(Bismarck Mts). 

Straelen, Victor Emile van 

(d. 1964). 

Literature. (1) Add: See some annotations 
on the Arfak area route by W. Vink in Nova 
Guinea, Bot. 22, 1965, p. 486. 

Biographical data. Add: Le Flambeau: 
'Hommage ^ V.-E. van Straelen, 1889-1964' (162 
pp., 2 pi., portr.); Bull. Soc. Linn. Norm. X, 5, 
1964, p. 12-19; Natur u. Mus. 94, 1964, p. 217. 

♦Street, John Malcolm 

(1924, Mcintosh, South Dakota, U.S.A.; x), 
B.A. of the University of California, Berkeley, in 

1948; Ph.D. at the same institution in 1960. Since 
1960 a member of the geography faculty at the 
University of Hawaii, Honolulu. He made 
expeditions to Haiti (1952 and 1953), and to New 
Guinea (see below). ^ 

Itinerary. Terr, of New Guinea. 1964. 
Bismarck Mountains via Simbai Patrol Post 
(July), collecting at the villages of Tsembaga, 
Gunts, Kwima, and Kompiai until Jan. 1965. — In 
7967 Bismarck Mts with his assistant H. Manner 
(see there) via the Koinambe Anglican Mission in 
April, collecting in the vicinities of Koinambe, 
Kompiai, Kupeng, and Tsuwenkai until Sept. 
Later collecting was done by Manner. 

Collections. Herb. Lae: 194 specimens New 
Guinea 1964 for identification and storage; 599 
specimens coll. Street & Manner New Guinea 
1967-68. Dupl. sets 1967-68 in Leyden, and in 
Geogr. Dept Univ. Hawaii. 

Literature. (1) Author of 'Grasslands on the 
Highland Fringe in New Guinea' (J. Queens!. 
Geogr. Soc. 3, 1966, p. 9-12); (with W. Clarke): 
'Soil Fertility and Cultivation Practices in New 
Guinea' (J. Trop. Geogr. 24, 1967, p. 7-11). 

Biographical data. American Men of 
Science; Who's who in Amer. Education. 

*Streimann, Heinar 

(1938, Tartu, Estonia; x), from 1959-61 at the 
Bureau of Meteorology, Melbourne, assisting in 
research; 1961-63 Dept of Forests, Bulolo, New 
Guinea — ^road locations, land surveys and map- 
ping; 1964-71 lecturing in botany and timber 
identification at the Bulolo Forestry College, 
establishing and maintaining a teaching and 
reference herbarium at the College; 1971-72 
Senior Technical Officer at the Division of Botany, 
Lae, in charge of botanical collections;* Nov. 
1972- in charge of the Herbarium at the 
Canberra Botanic Gardens. 

Collecting localities. 1964-72. Papua & 
Austr. New Guinea, except for the following 
districts: Southern Highlands, Madang, New 
Ireland and Bougainville. Much collecting in the 
Morobe District. 

Collections. In NGF series, often with A. 
Kairo, between 8500-9000 numbers, sets in 
Bulolo, Lae. Dupl. distributed by Lae. 

Literature. (1) Author of 'A new Balano- 
phoracea, subfamily Langsdorfieae, from New 
Guinea' (co-author J. S. Womersley, in Papua & 
New Guinea Scient. Soc. 1971). Timber Species 
Leaflets 1-10 published, 225 to be published. 

Stresemann, Erwin 

Biographical data. Portr. in Ornith. biol. 
Wissensch. Heidelberg 1949; list of ornithological 
publications in H. M. Whittell, 'The Literature 
of Austral. Birds' 1954, pt 2, p. 690-692. 

♦Sudjana Kasan 

Curator of the Bogor Botanic Gardens, 
collected a new Begonia near Padang, Sumatra 
West Coast. ^ 

Literature. (1) C. A. Jannsson: 'Ein neue 
Begonia aus Indonesien' (Act. Hort. Gotob. 26, 
1963, p. 1-4, 2 fig.). 




Cyclopaedia of Collectors — Supplement II 

T. G. H. 

♦Sumague, P. P., see Philippine National Her- 

Sun Hong Fan 

Collecting localities. Add: 1963. S. 
Sumatra (June-July): Baturadja, Palembang, 
Telok Belong. — 1964. Lesser Sunda Is: Lombok 

Collections. Add: 550 nos from Lombok 
(1964) in Herb. Bog.; dupl. Herb. Leyden. 

His total collections number to about 10.000 
(Flora of the Southsea Islands). 

*Suppiah, T. 

Assistant of the Forest Research Institute, 
Kepong, Malaya. 

Collecting localities. Malaya. 1966-67. 
JohOTC— 1968. Kelantan Ulu Sat F. R. (June 6- 
20); Selangor, Ulu Langat (May 2-7); Negri 
Sembilan, Jelebu (July 1-10). 

Collections. In Herb. F.R.L Kepong. 

Surbeck, Heinrich 

Collecting localities. Add: 1941. N. 
Sumatra (Sept.). 

Collections. Add: 1941 collection also in 
E.T.H. (Techn. Coll.) Zurich. 

''Susard, J. B., see Philippine National Herbarium. 


collected >94 nos at Tjibodas (Oct. 12, 1957), 
W. Java; in Herb. Bog.; partly in Herb. Leyden. 

*Suwanda Amongprawira 

of the Forest Research Institute at Bogor, 
accompanied the Tjendrawasih Expedition to the 
Central Highlands of New Guinea {Irian) from 
Inarutali to Byoga (Dec. 796i-Jan. 1964), col- 
lecting c. 100 nos. 

Sijde, Hendrik Adolf van der 
Add: Later he went to S. Africa. 

Symington, Colin Fraser 

Biographical data. Add: Cf. Mai. For. 
26. 1962, p. 107-108 (on the fate of his book on 

Sijpkens, J. P. 

Collections. Add: Herb. Wageningen: a 
dozen plants from Tjikopo Zuid (W. Java). 

•Srent-Ivany, J. J. H. 

collected a few plants on Mt Strong {Papua) at 
the end of August 1963; in Herb. Lae. 

•Tamavaschi, Dr 

from Rumania, accompanied A. (sec 
there) (jn a tour to Bali to study pioneer plants on 
Mt Agung in 1963. 

•Taylor, Edward 

collected a plant with D. H. NicOLSON (sec 
there) in Sarawak, Aug. I960. 

TCW nos, sec T. C. Whitmore. 

•Tclussa, A. W.. 
sec Boswc/cn Nieuw Guinea. 

*Teo Leong Eng 

(1938, Malacca, Malaysia; x) B.Sc. in Botany 
from Taiwan (1963); Research Assistant in Seed 
Technolouv at Chemara Research Station (Oil 
Palm Bra'nch) from 1963-65; at the Faculty of 
Agriculture, University of Malaya from 1965-66; 
since 1967 Technical Assistant (Botanist) in the 
Department of Chemistry. He will continue 

s t R E I M a N n 

collecting for the Phytochemical Survey of Malaya 
after the departure of J. Carrick (end 1967). He 
has a large interest in taxonomic work. 

CoLi.iCTiNG LOfAi.mis. Moslly in Sel- 
angor, Perak, Pahang, Trcngganu, and Johore 
of the states of Malaya. 

Collections. From 1968 he will make field 
collections with G. Pachiapi'an (sec there); they 
will begin their numbers as T. & P. 1 (PCSM no 
2601). Herb. Kuala Lump.; dupl. at Leyden, Kcw, 

Teruya, Z. 

Collections. Add: Also specimens in Kyoto 
Herb. Fac. Science. 

TcxJor, Carl Julius 

Add: He arrived in Java on June 25, 1842. He 
did not succeed to secure accommodation on one 
of the ships bound Un Japan, before June 1843. 
In the meantime lie was given a job on a plan- 
tation by the I). F.I. government ((/. Jaarb. Koii. 
Mij. Aanm. luinb. I, 1844. p. 15-16). 

T.G.H., sec I. G. Hartley 




Flora Malesiana 

[ser. I, vol. 8^ 

*Thonison, T. 

Collections. Java plants in Herb. Oxford 
{cf. Account by H. Newman Clokie, Oxford 1964, 
p. 117). 

*Thorne, Robert F. 

(1920, Spring Lake, N.J., U.S.A.; x) botanist, 
graduated with A.B. from Dartmouth College in 
New Hampshire (1941), awarded the M.S. (1942), 
and the Ph.D. on the flora of SW. Georgia (1949) 
at Cornell University. Subsequently to the State 
University of Iowa, later (1960) Associate Professor 
of Botany; at present Curator of the Herb, of 
Rancho Santa Ana Botanic Garden (Cal.) and 
Honor. Professor in the Claremont Graduate 
School. Though most of this earlier research has 
been in floristics, in recent years he took up the 
study of angiosperm phylogeny and geography. 
As a Fulbright Scholar he worked with Prof. 
Herbert at Brisbane, Queensland, and travelled 
widely in Australasia in 1959-60. After his return 
to the States he paid visits to Bogor, Singapore, 
etc., when en route by plane to England, where he 
would stay 5 months, working at Kew and the 
British Museum. 

Collecting localities. 1960. NE. New 
Guinea: with E. E. Henty near Lae, along Busu 
River (May 23), Markham Point and ridge above 
(24), Lae Bot. Garden (25), Oomsis Logging Area 
and Markham River (26); jointly with J. S. 
Womersley (May 31 -June 5) in the vicinity of 
Kikiepa, Wantoat Patrol Post Area, Lae Sub- 
distr., and Morobe Distr. (Finisterres); with 
Womersley (June 9-11) to Bulolo, Wau, and Edie 
Creek areas, up to c. 7000 ft ; with Womersley to 
SE. New Guinea (13-18): Port Moresby, Brown 
River, Sogeri Plateau areas, Rouna Falls. IV. 
Java: Tjibodas Mountain Gardens (July 23, with 
A. Dilmy). Singapore: Bt Timah (July 27, with 
Prof. Gilliland). 

Collections. In Australian New Guinea 
about 525 nos, viz 27437-27541 in his own series 
(coll. May 23-26), and the others numbered in the 
NGF series with J. S. Womersley. In Java and 
Singapore his own numbers 27814-27824, and 
27825-27835 respectively. The NGF numbers are 
placed at Lae etc. (see sub New Guinea Forests); 
collections under his own numbers (mostly 5 sheets 
apiece) at Lae, Brisbane, the 1st set probably going 
to State Univ. of Iowa and the others to the 
U.S. Nat. Herb, and Leyden. 

The 1st set of the Australian collection (appr. 
8000 nos) will be deposited at Iowa or U.S. Nat. 
Herb., the 2nd at Brisbane. The 1st set of the New 
Caledonia collections (>700 nos) is planned to go 
to Paris, the 2nd to Iowa or U.S. Nat. Herb., and 
the 3rd to Brisbane. 

Thunberg, Carolus Petrus 

Biographical data. Add: Act. Phytotax. 
Geobot. 2, 1933, portr. facing p. 226. 

Thung, Tjeng Hiang 

(d. 1960, Wageningen, Gld., Netherlands). 

Biographical data. Vakbl. Biol. 41, 1961, 
p. 1-3. 

*Tikau, Putan, see Forest Dept North Borneo. 
*Tingguan, Suah, see Forest Dept North Borneo. 

Tirto Atmodjo, R. 

(d. end Dec. 1903). 

Biographical data. Versl. 's-Lands Plan- 
tentuin for 1903, p. 20. 

*Tissing, M. O. 

naval surgeon who accompanied the Star 
Mountains Expeditions in Central New Guinea in 
1959 (see under C. Kalkman). He took part in the 
ascent of Mt Juliana (Sept. 9, 4640 m). He collected 
some plants in collaboration with Kalkman, 
numbered in the latter's series. 

*Togashi, Makoto 

Japanese professional plant collector. He 
collected for the 'Medical Company Herbarium 
Laboratory Takeda'. 

Collecting localities. 1961-62. Malay 
Peninsula, a.o. in Johore and Singapore. — 1966. 

Collections.^ In Kyoto Herb., Fac. Science. 
He has an intricate way of numbering, e.g. 
6222413 means 1962, 2nd month, 24th day, no 13. 

Literature. (1) J. Ohwi: 'Grasses of the 
Malay Peninsula and Singapore collected by Mr. 
M. Togasi' (Bull. Nat. Sc. Mus. 6, 1962, p. 119- 

*Tolentlno, M., see Philippine National Her- 

^Tomlinson, Phillip Barry 

B.Sc., Ph.D., Fairchild Tropical Garden, 
Florida, U.S.A. He made a trip to Papua on his 
way to Auckland. Possibly he collected plants. In 
Sept. 7955 he collected at Ipoh, Perak, Malaya. 

♦Toreu, B. N. 
collected with D. G. Frodin (see there) in 1973. 

*Toyoshima, Masami 

of the Insectivorous Plants Society, Dept of 
Biology, Nippon Dental College, Fujimi, Tokyo, 
Japan, joined an expedition to the Philippines in 
search for Nepenthes. 

Collecting localities. Philippines. 1965. 
Mindanao & Luzon (3 weeks in Aug., see S. 
KuRATA, also for Literature). 

Treub, Melchior 

Biographical data. Add:G. H. Zeijlstra: 
'Melchior Treub. Pioneer of a New Era in the 
History of the Malay Archipelago' (Amsterdam 
1959); in Dictionary of Scientific Biography, New 
York (ined.). 

Trono Jr, Gavino 

Add: Botanist, algologist in the University of 
Hawaii. Married to Miss L. Baskinas (see there). 

Try on, Henry 

Add: Collecting localities.^ In 1895- 
96 in Papua: sailing from Brisbane (Aug. 26, 1895) 
for Thursday I.; from there (Sept. 14) to Port 
Moresby, coast E of it, Kemp Welch R. (collection 
lost). Caution Bay, Port Moresby, several weeks in 
Milne Bay and Samarai, Mullen's Harbour area, 
and return to Cooktown (early Jan. 1896). 

Literature. (1) Cf. Hawaiian Plant. Rec. 55, 
1958, p. 213. 




Cyclopaedia of Collectors — Supplement II 


•Tuckwell, E. 

collected unnumbered plants on Mt Wilhelm, 
E. New Guinea (Nov. 1960). 

*Tuniau, E. A. 

collected some plants from Langkawi, Kedah 
(Aug. 5, 1962). 

•Turabull, John Wright 

(1940, England; x) took his B.Sc. (Hons) in 
Forestry (Bangor, Wales) 1962; Forestry Officer, 
South Australia Woods and Forests Department 
(1962-65); since 1965 Genetics Section, Forest 
Research Institute, Canberra, Australia. 

He made seed collections in western U.S.A. 
(1968) and in the Philippines (1969). 

CoLLECTi.NG LOCALITIES. 1969. Philippines. 
Luzon (Mar.-Apr.): provinces Ilocos Norte, Abra, 
Bontoc, Benguet, Pangasinan, Nueva Vizcaya and 
Zambales; VV. Mindoro (Apr.). 

The entire natural distributions of Pinus 
insularis and P. merkusii. 

Collections. Botanical samples and seed of 
above-mentioned Pinus species; in FRl Herb. 

L'lte«, Amoldus Johannes 

(d. 1963/64). 

Biographical data. Versl. Treub Mij. Mei 
1963-Mei 1964 (no particulars). 

•Umba, T. 

collected some 10 plants on Mt Wilhelm, E. 
New Guinea (Feb. 1970); in NGF series, Herb. Lae. 

LPNG nos 

series of the University at Port Moresby, New 

L'steri, Alfred 
Collections. Add: Also dupl. in Herb. Bot. 

Gard. Ziirich. 

•V'aldez, C, see Philippine National Herbarium. 

V'aleton Jr, Theodore 
(d. early 1962 or late 1961, Netherlands). 

•Vandenberg, Miss J. 

botanist, Div. of Botany, Lae; she collected on 
Mt Wilhelm (June 1968). 

Collections. In Herb. Lae: dupl. distributed, 
a.o. to Herb. Edinb. {New Guinea, 1968- ); ^250 
nus from Mt Wilhelm, in the NGF scries. 

Biographical data. Portr. in Fl. Mai. I, 7, 
1972, p. 285. fig. 4. 

VanoverberRh, Maurice 

Coi I I CT IONS. Add: List of new species and 
genera discovered, and those not earlier reported, 
in a paper by h. Quisumbing (sec below). 

Biographical data. Add: Act. Manila 4, 
1970, p. 25-51. 

•Vayda, Dr Andrew P. 

Assistant Professor of Anthropology, Columbia 
University, made a plant collection with his wife 
at Simbai, Madang District, Terr, of New Guinea, 

in 1963. In Herb. Lae: c. 80 nos; Pandanus dupl. 
to Dr Stone, bamboos and ferns to Dr Holttltm. 

Vecht, Jacobus van der 

Add: From 1952-55 professor of phytopathology 
(zoological part) at the Agricultural College at 
Bogor. From 1955 onwards employed in the 
Zoological Museum at Leyden; in 1962 appointed 
Extraordinary Professor of Zoological Taxonomy 

V t L D K. A M i' 

at Groningen University and from 1964-68 
Ordinary Professor in the same discipline at 
Leyden University. At present living at Putten, 
GId., Netherlands. 

•Velasquez, Dr Gregorio T. 

(1901, Calumpit. Bulacan, Philippines; x), 
phycologist, B.S. (1925) and M.S. (1931) of the 
University of the Philippines; M.A. (1937) and 
Ph.D. ( 1939) of the University of Michigan, U.S.A. 
Emeritus Professor of Botany, University of the 

Author of numerous publications on Algae. 

Collections. He collected Algae from 
numerous localities in the Philippines; in Herb. 
Manila, and distributed to Leyden, Crypt. Herb, of 
Hokkaido and Kagoshima Universities, Sing., 
Adelaide, Hawaii, Berkeley, Cambridge (Harvard), 

Bio(iR Ai'HKAL data. Amcr. Men of Science 
1949; ibid. II. Biol. .Sc. '>lh cd. I';55; ibid. Physic. 
& Hiol, Sc. 11th ed. I'>07; Philip. Men of Science 

'Veldkamp, Jan Fiedcrik 
(1941, Amsterdam, Netherlands; x), botanist. 




Flora Malesiana 

[ser. I, vol. 8^ 

educated at Leyden University, where he took his 
Ph.D. degree in 1973. Since 1967 a staff member 
of the Rijksherbarium, Leyden, in charge of the 
Malesian Gramineae. He made an expedition to 
New Guinea, after which he stayed in Indonesia 
(mainly Bogor) till Sept. 1972. 

Collecting localities. 1972. Papua & 
New Guinea: with P. F. Stevens (see also Crutt- 
well and Pullen): Tumba (May 25-29), Goroka 


(31), Goropu-complex or Mt Suckling (June 7- 
July 25),^ collecting around the base camp Mayu 
II (1750 m), at a grass plain (Pumpunipon, 2000 m) 
where — 8°C was measured, ascending to Goe 
Dendeniwa (3625 m), back to base and ascending 
Manurep or Mt Suckling (3676 m) via Goe (3580 
m). — Centr. Java: Dieng plateau (Aug. 14). — 
Bali: Batur (Aug. 2\).~W. Java: Bogor (Sept. 15). 

Collections. The New Guinea specimens 
will be deposited in Herb. Leyden, Lae, B.P. Bish. 
Mus., Canberra, Bog., the Java specimens in Herb. 
Leyden and Bog. The Malesian collections are 
numbered 5442-6040 and in the NGF and LAE 
series (with P. F. Stevens). 

Literature. (1) J. F. Veldkamp: 'A botanical 
Expedition to Mt Suckling (Goropu Mountains), 
Papua — New Guinea, 1972' (stencilled report, 
Leyden 1973, 11 pp., map). 

Veloira, Miss Nieya 

Add: Finished her M.D. in Botany in Tennessee 
University; bryologist in the Botany Dept U.P. 

Verdoorn, Frans 

Biographical data. Add: Regn. Veget. 71, 
1970, xv-xxii + 4 pp. suppl., portr. 

Vergara, Benito S. 

Add: plant physiologist, Ph.D. Chicago; on the 
staff of the I.R.I. (International Rice Institute), 
Los Bafios. 

*Verheijen, Father Jilis Antonius Josephus 

(1908, Zevenaar, Gld., Netherlands; x), a 
Roman Catholic missionary who after his clerical 
education was stationed in Flores (1935- ), 
interrupted by one and a half year of study in 
Cultural Anthropology and Linguistics at Utrecht 
University (1949-50). * 

Collecting localities. Indonesia. W. 
Flores: on the south coast in the vicinity of (M)- 
borong (0-200 m alt.), and the vicinity of Sita 
(400-600 m); in the rain-forests on the north slope 
and on the summit of the mountain range in 
Central Manggarai above kp. Mano, Robo, and 
Ruteng (1200-1950 m); in south Tjibal (N of the 
range) between 500 and 9(X) m, and at some other 
odd localities. Also on P. Roti {1969). 

Collections. Herb. Leyden: > 2908 nos in a 
poor state (nos 2379-2460 from Roti). 

The collection was made in connection with the 
preparation of a dictionary of languages and 
contains some remarkable species. 

Literature. (1) Author of papers on an- 
thropology and linguistics of the people of Flores, 
a dictionary of Manggarai language, and of some 
ornithological papers in Ardea (1961, 1964). 

Ver Huell, Quirijn Maurits Rudolf 

Biographical data. Add: Bijdr. Taal-, 
Land-, en Volkenk. 113, 1957, p. 261-273, portr. 

Vermeulen, Pieter 

Literature. (1) Add: 'Een botanische reis 
naar het Verre Oosten' (Vakbl. Biol. 38, 1958, p. 

Verschueren, Johannis Cornells 

Add: Biographical data. In Streit & 
Dindinger, Bibl. Missionum 21, 1955, p. 580, 

Versteegh, Christiaan 

Add: He left New Guinea in Dec. 1962. Since 
Oct. 1963 attached to the Plant-anatomical 
Institute of the Agricultural College at Wagen- 

Collecting localities. Add: 1954. Anggi 
Lakes with J. F. U. Zieck a.o. (Feb.).^— 1956. 
Forest area near Sausapor, NW. coast Vogelkop 
Peninsula (2nd half Oct.).— 1957. Oransbari; S. 
New Guinea: Digoel R. (Koeweh, Wage, Nanggoe, 
Getenteri, Omba, Sabekahang, Waghabang, Tanah 
Merah) (Nov. 12-22).— 1958. IV. New Guinea: 
Beriat and Wersar, near Teminaboean (Apr. 18- 
May 5); with F. A. W. Schram in Ajamaroe area 
(Ajamaroe, Semoe, Jokwer, Kamboeaja, Djitmao, 
Fan, Tehahg, Aitinjo, Erokwero, Joksoro, Awit, 
Ewai, Mos, Kami, Itaro, Koa, Tenimaboean, 
Skendi (May); Oransbari (Sept. 18); Biak /.— 

1959. Biak I. with W. Vink (Kg. Landbouw, Son, 
Bosnik, Parjerie, Siapis = Siabes) (Feb. 18-25). — 

1960. Sekoli Plain, S of Lake Sentani (Feb.- 
Mar.); P. Adi (Aug. 12-20); Mt Genofa near 
Kaimana (Aug. 23-30); Kebar Valley (Andjai, Mt 



Cyclopaedia of Collectors — Supplement II 


Watjetoni, Mt Nettoti) (Nov. 2(>-Dec. 2).— 1961. 
E part of Arfak Plain (Masni) (Feb. 24-29); 
Baliem Valley (Wiligimaan) (June).— 1962. With 
Ch. Koster to central part Arfak Mts (Minjambau, 
Mt Antop, Mt Nerimbau, Mt Lensemoi) (May 16- 
28).* — From 1957-62: surroundings of Hollandia 
in N. \ew Guinea (Noordvvijk, Kloofkamp, 
Hamadi, Berg en Dal, Renauwen, Dojo, Dozai); 
surroundings of Manokwari in IV. New Guinea 
(Maripi, Maroeni, Andai, Arfai, Saowi, Wosi, 
Fanindi, Manggoapi, Amban, Tafelberg, Borasi, 
Pasir Poetih). 

Literature. (1) J. F. U. Zieck: 'B.W. 
orientatietoumee Ransiki-Anggi-meren' (Manok- 
wari 1954, typewritten); 'De Anggimeren' (Ned. 
Nieuw Guinee 3, no 7, 1955, p. 7-10). And cf. W. 
Vink in Nova Guinea, Bot. 22, 1965, p. 490, fig. 3. 

(2) Cf. W. Vink in Nova Guinea, Bot. 22, 1965, 
p. 493^94, fig. 6. 

*Versteegh, R., see Boswezen Nieuw Guinea. 

Vertenten, Petrus 

Add: (d. 1946, Wibrijk, Belgium). 

Biographical data. In Streit & Din- 
DINGER, Bibl. Missionum 21, 1955, p. 580-581. 

*Ver?oort, Willem 

0917, Schiedam, Netherlands; x), zoologist, 
educated at Leyden University (M.Sc. 1941, Ph.D. 
1946); from 1940-^^, 1948-50, and since 1959 on 
the staff of the Rijksmuseum van Natuurlijke 
Historie at Leyden; from 1946-48 biologist in the 
employ of the Dutch Society for Whale-fishery, 
and from 1950 Teacher in Descriptive Zoology, 
Zoological Laboratory, University Leyden. In 1972 
appointed Director of the Natural History Museum 
at Leyden. In October 1973 Extra-Ordinary 

He accompanied the Dutch New Guinea Ex- 
pedition to the Sterrengebergte in 1959. Some 
scientific papers on the zoological results of the 
expedition will be published in due time. 

Collecting localities. 1959. Biak 
(Schouten I.); N. Dutch New Guinea: Hollandia; 
Centr. Dutch New Guinea: Ok Sibil and other 
localities in the Sterrengebergte, e.g. Tenmasigin 
(c. 1800 m). 

Collections. Plants collected with dr A. G. 
DE Wilde were added to the collections of C. 
Kalkman, and Dr B. O. van Zanten (see those); 
in Herb. Leyden, and Groningen. 

Vidal y Soler, Sebastian 

Collections. Add: Herb. Arn. Arbor.: 2528 
dupl. fprcs. 1958-59). 

Biographical data. Add: Fl. Mai. Bull. 
no 17. 1962, p. 898-899 (statue). 

•Vink, Willcm 

(1931, .Schiedam, Netherlands; x). botanist, 
educated at Ixydcn University (Ph.D. 1970); after 
military service Forest Botanist at Manokwari, 
West New Ciuinca, 1959 62; mid-1962 appointed 
on the staff of the Rijkshcrbarium, l.cydcn. 

Zanthoxylum vinkii Hartl. and Rhnlixlcndron 
vinkii Sleum. have been named after him. 

Collecting localities. 1959. Biak I. with 
Chr. Versteegh (Feb. 18-25); W. New Guinea: 
Sarmi hinterland (Kwansem, Bodem, Sidoearsi 
Mts, Oereb R., Bier R., Kondirdjan, mouth Tor 
R.) (May 12-June 14).— 1960. With F. A. W. 
Schram (see there) to the Wissel Lakes (Enarotali, 
Komopa, Borobaida, Motito, Moenajepa, Mt 
Monoglito, Doglia, Kebo, Mt Pekeglbaro, Kebo, 
Enarotali) (May 11-30). Terr, of Papua & .\ew 
Guinea: some collections with J. S. Womersley 

and Mrs A. N. Millar in NGF series (June 16- 
July 15), from Lae, Bulolo, Wau, Edie Creek, 
Bulolo, Pi Moresby, Rouna Falls, Idlcss Bay, 
Brown R., via Popondctta and Embc near Lae, 
Rabaul, Keravat, Rabaul, via Kavieng and Wewak 
to Madang (Gogol Valley, Mt Hagcn, Wabag, 
Kempiam, (Thimbu, Goroka, Lac. Japen I. 
(Seroei, Randawaja, Soembcrbaba, Aisao, Socm- 
berbaba, .Seroei (Sept. 20-Oct. 10).'— 1961. iV. New 
Guinea: Kebar Vallev (Andjai. Asiti, Abi R., 
Apriri R.) (Feb. 12-24); Sidci (June 5-10); Arfak 
Mts, Mt Bcribai' (Maroeni bivouac. Lake Nja- 
boeti, bivouac 1075 m, crest Mt Beribai (up to 
2100 m) (June 19-24); Biak I. (Mokmer. Oregon 
trail, Parjeri, .Saribi. Mokmer, Korim-road. 
.Sjabes) (June 28-July 6); Mapia Is (Pcgun, Bras) 
(July 7-8). -1962. W. New Guinea: Arfak Mts,* 
Anggi Lakes, together with H. O. Siiumir (Jan. 
8-29); lakfak area (P. Pamljann, I akfak K.. 
Kowap, llambar) (f'cb. 21 27); Lake Ajamaroc 
(Ajamaroc, lanah Mcrah. .Scgior. .Seta. I ubun, 
partly together witli Mrs Vink). 1959 62, In the 
surroundings of Manokwari. 196.1. Centr. Sew 
Guinea. Kubor KanKc Kxpcdiliun' with R. Pui ii n. 




Flora Malesiana 

[ser. I, vol. 8^ 

Western Highlands District (Minj, Kamang, 
Uinba, Nona-Minj Divide, Mt Kinkain, Mt Aas, 
Mt Kantz, Mt Miiyin Kolyin up to 4040 m alt.) 
(June 17-Sept. 16).— 1966. E. New Guinea: Tari 
Subdistrict Expedition with C. Kalkman (see 
there), A. N. Gillison, and D. G. Frodin, 
mountains in the Southern Highlands District; 
and in Telefomin: Subdistrict on Mt Amdutakin, 
Hindenburg Range (Sept. 9-18).*— 1968. W. New 
Guinea: Warsamson Valley, E of Sorong (May- 


Collections. BW nos in Herb. Manokwari, 
dupl. in Leyden, Lae, Brisb., Canb., Bog.; all from 
W. New Guinea and nearby islands, c. 1 300 nos. 

From the Arfak Mts (1962) c. 300 with ex- 
pedition numbers Sleumer & Vink; others with 
BW nos. Plants from the Terr, of Papua & New 
Guinea I960 in NGF series, at Lae and elsewhere; 
from 1963 and 1966 numbered Vink 16001-16583, 
and 16801-17642 respectively. 1st set in Leyden, 
2nd Lae, 3rd Canb. 

Literature. (1) W. Vink: 'Verslag van een 
tournee van de sect. Bosbotanie naar Oost. Japen 
(Aisau)'. Report 1960. 

(2) W. Vink: 'Botanical Exploration of the 
Arfak Mountains' (Nova Guinea, Bot. 22, 1965, 
p. 471-494, fig. 1-6, pi. 42-46). 

(3) W. B. Hitchcock: 'An Introduction to the 
Natural History of a New Guinea Highland 
Community' (Emu 63, 1964, p. 351-372, map). 


(4) C. Kalkman & W. Vink: 'Botanical Ex- 
ploration of the Doma Peaks and adjacent 
mountains in the Southern Highlands District, 
Territory of Papua, Australian New Guinea' 
(Report for 1966, Neth. Found. Advanc. Trop. 
Res. WOTRO, p. 50-52, 2 photogr.); 'Botanical 
Exploration in the Doma Peaks, New Guinea' 
(Blumea 18, 1970, p. 87-135, fig. l(map)-5, 
photogr. 1-25). 

Viola, Miss V. 

Add: Instructor of Botany, University of the 
Far East, Manila. 

Vischer, Wilhelm 

(d. 1960, Basel, Switzerland). 

Add: Biographical DATA. Verh. Schweiz. 
Naturf. Ges. 140, 1960, p. 253-257, portr.. 
bibliogr.; Verh. Naturf. Ges. Basel 72, 1961, p. 
358-363, portr., bibliogr.; Rev. Algol, n.s. 7, 1964, 
p. 115-118, portr., bibliogr. 

*Vlugt, P. J. van der 

visited Johore (Malay Peninsula) in Aug. 1968; 
collected living waterplants.' 

Literature. (1) P. J. van der Vlugt: 'Nog- 
maals vier nieuwe exoten' (Het Aquarium 40, 1970, 
p. 280-284, 5 photogr.). 

*Vogel, Eduard Ferdinand de 

(1942, Schiedam, Netherlands; x), biologist, 
educated at Leyden University, where he took his 
Master's degree in 1970; in Sept. 1971 he went to 
Bogor, Java, in charge of a 3-years development 
cooperation project on the study of primary forest 
tree seedlings, sponsored by NUFFIC. He founded 
a nursery for seedlings in the Bogor Botanic 

Collecting localities. 1971. SE. Borneo 
(Kalimantan Selatan) with J. Dransfield, K. 
Kartawinata, three assistants of the L.B.N., 
Bogor, and an assistant from the University of 
Bandjermasin : Proceeding from Bogor to Djaro 
near Muara Uja (Nov. 8), alluvial flat and hills 
around Djaro Dam (9-18), Batu Kumpai, lime- 
stone hill near Djaro (20), limestone hills around 
Djaro (21-22), alluvial flat around Djaro (23-24), 
proceeding to Gunung Sarempaka (25), Gunung 
Sarempaka (bivouac) (26), back to Djaro Dam 
(27), back to Bogor (Nov. 28-Dec. 5).— 1972. W. 
Java: G. Gedeh-Pangrango, above Tjibodas (Feb. 
2). S. Sumatra (with his wife I. de Vogel-Rave- 
stijn, J. Dransfield, and three assistants of 
L.B.N., Bogor): proceeding from Bogor to Sele- 
man Enim, near Muara Enim (March 7-8), 
neighbourhood of Muara Enim (9-11), proceeding 
to Bukit Seburong, near Muara Dua (12-13), 
Bukit Seburong area (bivouac) (14-16), pro- 
ceeding to Bandung Agung, along shore Danau 
Ranau (17), neighbourhood Bandung Agung and 
Balik Bukit near Kota Batu (18-19), return to 
Bogor, collecting 20 km SE of Martapura (20-21). 
W. Java: G. Gedeh-Pangrango, above Tjibodas 
(March 25); Dungus Iwul, about 10 km W of 
Kabupaten Bogor, on road Bogor-Labuan (Apr. 
10). Pulau Peutjang, Udjon Kulon N.R., as guest 
of BiOTROP course Forest Ecology : proceeding to 
Pulau Peutjang (June 19-20), Udjon Kulon 
peninsula, path of lighthouse, Pulau Peutjang, 



Cyclopaedia of Collectors — Supplement II 


along beach, around guesthouse (21), transect 1, 
behind guesthouse (22), around transect Karta- 
winata (23), along beach and on flatland (24), 
flatland and marsh (25), hills and flatland (26), 
flatland (28), back to Bogor (29-30). A^. Sumatra 
G. Leuser Reserve, Ketambe, N of Kota Tjane, 
partly with W. J. J. O. de Wilde, his wife B. de 
WiLDE-DuYFJES, H. RiJKSEN and his wife (July 
8-25): proceeding to Ketambe, on the way 
collecting in Sibolangit Botanic Gardens (July 8-9), 
G. Leuser Reserve, Ketambe, about 30 km N of 
Kota Tjane, surroundings of Alas valley (10-14), 
proceeding to G. Ketambe, in above-mentioned 
company, collecting along Lau Ketambe (15), 
ridge of G. Ketambe, 540 up to 1700 m (16), ridge 
to peaks of G. Ketambe (17-19), back to Ketambe, 
collecting along ridge and river (20), surroundings 
Alas valley (21-23), back to Bogor (24-25). 
Pulau Peutjang, Udjon Kulon Nature Reserve, 
with I. DE Vogel-Ravestijn, J. F. Veldkamp and 
his wife, and several others: proceeding to P. 
Peutjang (25-26), P. Peutjang (27-31), Udjon 
Kulon peninsula (Aug. 31 -Sept. 1), back to Bogor. 
Kalimantan Selatan, with A. Everaarts and 
assistant of L.B.N. , Bogor: Proceeding to Jaro 
(neighbourhood of Muara Uja) (Okt. 10-12), 
neighbourhood of Jaro Dam, alluvial flat and hills 
(13-14), Batu Kumpai, limestone hill (15), 
neighbourhood of Jaro Dam, alluvial flat and hills 
(16-18), proceeding to Gunung Sarempaka, 
bivouac (19), alluvial flat at foot of G. Sarempaka 
(20-24), side of G. Sarempaka up to ridge, 
limestone (22-24). alluvial flat (25), top ridge G. 
Sarempaka, bivouac, collecting on limestone 
ridges and a clay slope, up to 960 m, but unable 
to reach the top of G. Sarempaka (Oct. 26-Nov. 3), 
alluvial flat (Nov. 4-5), back to Jaro Dam (6), 
neighbourhood of Jaro Dam (9-10), back to Bogor 
(no collections (11-15). — 1973. Kalimantan Sela- 
tan: leaving Bogor (Mar. 3), proceeding to Jaro 
(5), staying 3 weeks in the same region as in 
Oct./Nov. 1972. 

Collections. > 1440 nos, 1st set Bogor, 2nd 
set at Leyden, next sets not yet sorted out. Of his 
most important collections the numbers run as 
follows: Kalimantan Selatan (1971): 700-1199, S. 
Sumatra: 1209-1349, Udjon Kulon Nature Re- 
serve: 1363-1411. N. Sumatra: 1413-1568, Kali- 
mantan Selatan (1972). 1585-2142. 

Voogd, Cornelis Nicolaas Abraham de 

(d. 1970. Wassenaar, Netherlands). 

Biographical data. FI. Mai. Bull, no 25, 
1971, p. 1879-1880. 

Vorstman, Adriana Gcertruida 

(d. 1963, Amsterdam, Netherlands). 

Add; Limnologist. The collections in Rawah 

Bening were made with Dr J. H. Cofrt (sec there). 

On the labels her name is misspelled For.stman. 
fliooRAPiMCAL DATA. Vakbl. Biol. 43, 1963, 

p. 89-91. 

•Wade, Uslic Keith 

(1940, Vancouver, B.C., Canada; x), phyto- 
sociologist, B.Sc. (1963) and M..Sc. (1965). 
University of Columbia. In summer 1963 taking 
part in an expedition to the western Canadian 
Arctic with Prof. V. S. Krajina. He started his 

postgraduate studies at the Australian University 
of Canberra, and spent a period in New Guinea 
for the phytosociological study of the alpine and 
subalpine vegetation of Mt Wilhelm under Dr D. 
Walker (see there). ^ He later returned to Canberra 
to complete his thesis for a Ph.D. 

Itinerary & Collecting localities. 
E. New Guinea (May 7965-July 1967), mostly on 

D. walker 

Mt Wilhelm, but also on Mt Giluwe, Upper 
Sirunhi and Chimbu Valley. Also visiting Bougain- 
ville (Solomons), Lake Iriste area {Papua), Mt 
Giluwe in the Southern Highlands, at Lae (2 
months herbarium work), and making short trips 
to Mt Kainde, Wau, Bulolo, and Madang. 

Collections. Herb. Lae: ANU nos 7000- 
7794; dupl. will be deposited in Canberra, Leyden, 
Kew, and Arn. Arbor. 

Literature. (1) L. K. Wade & D. N. 
McVean: *Mt Wilhelm Studies I. The alpine and 
subalpine vegetation' (Res. School Pacif. Stud. 
Dept. of Biogcogr. «& Geomorph. Publ. BG/I, 
1969. 325 pp.; Austr. Nat. Univ. Press). 

Wakefield, Norman Arthur 

(d. 1972. near Melbourne, Vict.. Australia). 

Bio(iRAPHiCAL DArA. Vict. Natuf. 90, 1973, 
p. 103-105. 

♦Walker, Donald 

(1928, Dcwsbury, Fngland; x) graduated B.Sc. 
ill Botany (lions l'M'>) at .Slic-llickl University, 
M.A. (1952) and Ph.D. (195.1) at Canibiidgc 
University. University Sub-dcpartmcnl of Quater- 
nary Research, Carnbritlgc, atui Clare College 
Cambridge (1949 W)), interrupted by military 
service in Malaya (1953-55); Professorial I'ellow 




Flora Malesiana 

[ser. I, vol. 8^ 

in Biogeography (1960-68) and Professor of 
Biogeography (1969- ), Research School of 
Pacific Studies, Australian National University, 

Collecting localities. Highlands of 
Australian New Guinea:^ Wabag-Laiagam, and 
Mt Wilhelm mainly, in 1961 (4 weeks), 1962 (5 
months), 1965 (3 months), 1967 (6 weeks). 

Collections. Made only in relation to 
ecological and pollen-analytical work. Actual 
collecting, label composition etc. is done largely 
by Mrs Patricia M. Walker. In Herb. C.S.I.R.O., 
Canberra, from where the duplicates are distrib- 
uted. About 1200 numbers, all numbered in the 
ANU series (between 1 and 1499, 5000 and 5499). 

Literature. (1) Author of: 'Stratigraphy and 
ecology of a New Guinea Highlands' swamp' (in 
Proc. UNESCO Humid Trop. Symp. Kuching 
1963, 1965); 'Vegetation of the Lake Ipea Region, 
New Guinea. L Forest, Grassland and Garden' 
(in J. Ecol. 54, 1966); 'A commentary on botanical 
data from New Guinea, Australia and New 
Zealand' (in R. Meteorol. Soc. Proc. Internat. 
Symp. on World Climate from 8000-0 B.C. 1966); 
'A reconnaissance of the non-arboreal vegetation 
of the Pindaunde catchment. Mount Wilhelm, 
New Guinea' (in J. Ecol. 56, 1968, p. 445-466, 
fig. 1-5, tab. 1-2). 

Walker, George Warren 

Collections. Add: Dupl. in Herb. Edinb. 
{ex Herb. Glasgow). 

*Walker, Patricia M., see D. Walker. 

♦Walker, Trevor George 

(1927, York, England; x), educated at Leeds 
University, B.Sc. Hons Botany (1952), Ph.D. 
(1956). From 1956-59 Temporary Assistant 
Lecturer in Botany, Leeds University, and in this 
period 1 year at the University of the West Indies 
as an Exchange Lecturer; since 1959 Lecturer in 
Botany, University of Newcastle upon Tyne. 

He made expeditions to Ceylon (1954), Jamaica 
(1958, 1962, 1963), Trinidad (1963, 1966) and to 
New Guinea (see below). 

He collected material for cytological, anatomical 
and evolutionary studies on tropical ferns. 

Collecting localities. East New Guinea. 
1964-65. With the British Museum-Newcastle upon 
Tyne Expedition. For foot patrol into Finisterre 
Mts see itinerary and literature under C. D. 
Savers, and A. C. Jermy. In 1965 around Lae, in 
the Herzog Mts, Eastern Highlands District, at 
Keglsugl and Mt Wilhelm. During 3 weeks (from 
Feb. 21) attached to a Forest Dept Survey in 
southern New Britain. 

Collections. Mainly ferns, «o.yT 7414-10285 
from New Guinea and New Britain (T 10014- 
10285) in Herb. Newcastle; dupl. will be distributed 
to various herbaria. Many living plants sent back 
to England for study. 

Wallich, Nathaniel 

Add: He laid out the first Botanic Garden in 
Singapore in 1822; it was subsequently abandoned 
during the 1830s (inf. /./. by J. Bastin, July 1970). 

Collections. At the end add: Letters re- 
lating to the establishment of the Botanical 


Garden in Singapore have been reproduced by R. 
Hanitch (J. Str. Br. R. As. Soc. 65, p. 39^8; 
reprinted J. Mai. Br. R. As. Soc. 42, 1969, p. 

Walsh-Held, Maria Ernestine 

(see Cycl. Fl. Mai. I, 1, 1950, p. 559, 606). 

(d. 1973, Geneva, Switzerland) 

Add: In March 1958 she repatriated to Switzer- 

Ward, Frank (or Francis) Kingdon 

(b. 1885, Withington, Lane, England; d. 1958, 
London, ashes buried at Grantchester, England). 

Add: Numerous plants have been named after 
him, e.g. the genus Kingdon-Wardia, and several 

Biographical data. Add: Everybody: 
Weekly, Nov. 1, 1952, with photogr.; in F. 
Kingdon- W ARD : 'Pilgrimage for Plants', 1960, p. 
11-18, incl. summary on Travels and Collections, 
and bibliogr. {I.e. p. 181-186). 

*Warent, H., see Boswezen Nieuw Guinea. 

*Waring, P. 

collected plants in Papua with Coode etc. 
Collections. Herb. Lae; dupl. Herb. Edinb. 

*Warner, John Northrup 

(1919, Los Angeles, Cal., U.S.A.; x) was 
educated in the Universities of Hawaii (B.S. 1941) 
and Minnesota (Ph.D. 1950, Plant Genetics and 
Statistics). From 1941 to present employed by the 
Experiment Station of the Hawaiian Sugar 
Planters' Association, at present Senior Geneticist. 
For his visits to New Guinea see respectively sub 
J. H. Buzacott and C. O. Grassl. 

*Wasli bin Haji Sisi, see Forest Dept Sarawak. 

Waterstradt, Johannes (John) 

Add: According to Barlow's biography (see 
below) he repatriated in 1904, taking over his 
father's market gardening business, specially in 
orchids. He returned to the East in 1908 and 1912, 
visiting Mt Kinabalu again. 

From W.'s diaries it is apparent that he took a 
passage to Australia not before 1890 or early 1891, 
and started collecting with the Germans in Ceylon 
in the middle of 1891. 

Collecting localities. (Emendated after 
Barlow, i.e. after W.'s diaries). 1891. Ceylon: 
Penang and Singapore; Labuan (end Oct.); Brunei, 
trip up the Padas R. — 1892. A^. Borneo: vicinity of 
the mouth of the Lawas R.; Mt Kinabalu (May- 
June), based at Kiau. — 1893. Back at Kiau (Jan.- 
end of Mar.).— 1894. Kudat (Jan. 23) and by boat 
to Banggi (? Banguey I.) (till end of Mar.). Philip- 
pines: Balambangan, Balabac, and Palawan. 
Return to A^. Borneo: Kudat (end of May), and 
Labuan (June 1). Expedition up the Lawas and 
Limbang Rivers {Brunei); Labuan (Oct.); return to 
Mt Kinabalu, based at Kiau again, visiting the i 
Marei Parei ridge, returning to the coast on Dec. " 
31. — 1895. Short expedition to Kinabalu (Mar.) 
and to Palawan, returning Aug. 2. — 1896. In 
March sending native collectors to Kinabalu for 



Cyclopaedia of Collectors — Supplement II 


insects, while he went to Mengalung for 2 months. — 
1899. Back in Labuan after 6 months in Europe; 
heavily involved with oil concessions and pros- 
pecting in Brunei, he arranged for collectors to go 
to Ternate, Halmaheira, Batjan, and Obi. — 1901. 
Malay Peninsula: ascent of G. Tahan (May-Nov.);* 
early Dec. back in Labuan. — 1902. Trip to Brunei 
(end of Jan.); to Sulu Is. (Jolo) via Sandakan; 
leaving (Feb. 27) for Mindanao: Zamboanga, and 
by steamer to jV. Celebes: Menado, collecting in 
that island and in Obi (21 2 months) before con- 
tinuing to Batjan, based at Labuha; leaving Batjan 
(Sept. 1), spending 6 weeks on Halmaheira and 
surrounding islands {Moluccas), and the remainder 
of the year at Saonek in Waigeo. — 1SK)3. Leaving 
Halmaheira (end of Feb.) for Ternate, and thence 
to Ambon. Because of impaired health return to 
Labuan. N. Borneo: Mt Kinabalu, Kiau (June 12- 
July 29); back to Labuan. Mindanao: Davao (till 
end of the year), travelling in the S. part of the 
island, climbing Mt Apo up to c. 6000 ft.— 1904. 
Labuan (Jan.); E. Java: Soerabaja (till end of 
March); back to Europe. — 1908. A^. Borneo: Mt 
Kinabalu.— 1912. Mt Kinabalu. 

Collections. According to Barlow the 
majority of his orchid collections before 1904 went 
to Hugh Low & Co. in London. He made 
extensively use of native collectors. 

Biographical data. H. S. Barlow, J. Mai. 
Br. R. As. See. 42, 1969, p. 115-129, 1 pi. 

*Web€r, Bruce 

American botanist educated at the University of 
Cincinnati. In June 1965 attached to the Sabah 
National Park Service and Forest Department in 
N. Borneo for two years. Collections? 

♦Weber, W. A. 

collected mosses and lichens in E. New Guinea 
under D. Walker; in the ANU series of the 
Australian National University; Herb. Canberra. 
In 1969 he accompanied Dr B. O. van Zanten 
(see there) on his trip in E. New Guinea (June- 

♦Wells, J. 

A Cadet Forest Officer who collected briefly 
(about 1957) at Bulolo, Morobe District, E. New 
Guinea; in Herb. Lae. 

Wells, T. H. 

(see Cycl. Fl. Mai. I, 1, 1950, p. 566) a farmer 
of the Childers District in Queensland, was 
commissioned by the Bureau of Sugar Experiment 
Station at Brisbane, to visit SE. New Guinea for a 
sugar-cane collecting trip, making Sogeri his point 
of departure. 

Iiimrary.' 1912. Left Brisbane (.Sept. 2); 
from Sogeri lo the Laloki River, Kemp Welch 
River and Rigo; leaving Papua in Nov. 

CoLi.r.fTioNS. Varieties of sugar-cane. Only 
few material will have been dried, if any. 

I.i ThRATURi . (1) Cj. Hawaiian Plant. Rec. 55, 
195«, p. 213; Cane Growers' Quart. Bull. Brisbane 
15. 1951, p. 39-40. 

Went, Frits Warmolt 

Add: For some years Director of the Missouri 
Botanical Ciardcn. In 1964 he accepted a professor- 


ship at Reno, conducting studies on desert plants 
and others as well. 

In Sept. 1962 he visited Mt Wilhelm in E. New 

Literature. (3) F. W. Went: 'A botanist's 
explorations in Australia and New Guinea' (Mo. 
Bot. Gard. Bull. 51, 1963, p. 10-17, 3 fig.). 

Wenzel, Chester A. 

Collections. Add: Also dupl. Philippines 
(1913) in Herb. Edinb. 

Werner, Eugen 

Collections. Add: Also dupl. in Herb. 

♦Wheeler, Josselyn 

student, working under Dr D. Walker (see 
there) and collecting plants in E. New Guinea. 

In May 1966 he worked in the Eastern Highlands. 

Collections. In Herb. Univ. Canberra, 
numbered in the ANU series (see there); con- 
siderable collection. 

♦White, S. R. J. 

made a collection of seeds, when travelling 
through Indonesia to Singapore, for W. H. 
Levvis.^ They were collected in Java: Bandung, 
Timor: Atambua, Singapore. 

Collections. Seeds of Oldenlandia corymbosa 
L. (= Hedyotis corymbosa (L.) Lamk). Not known 
whether he made other collections. The plants 
resulting from the seeds were given Lewis numbers.' 

Literature. (1) W. H. Lewis: 'Chromosome 
numbers of Oldenlandia corymbosa (Rub.) from 
Southeastern Asia' (Ann. Mo. Bot. Gard. 53, 
1966, p. 257-258). 

♦Whiteman, P. C. 

a member of ANZDEC (Asian New Zealand 
Development Consultants) team, deposited some 
specimens collected during tours (localities etc. see 
under D. W. Ivts) in Herb. Bog. 

♦Whitmore, Timothy Charles 

Add: Took his Ph.D. in 1960, and in the same 
year collected in Ecuador with the Oxford Uni- 
versity Expedition. From Sept. 1962-Sept. 1964 
Forest Botanist in the British Solomon Islands; 
from Sept. 1965-Apr. 1972 Forest Botanist under 
the Colombo Plan at Kcpong. Malaysia, specially 
in connection with the work for the Tree Flora of 
Malaya", interrupted by a stay at Cambridge and 
Kew (1968/69). and a visit to the New Hebrides 
and the British .Solomon Islands (Feb. -May 1970). 

He definitely left Malaya mid-April 1972, 
repatriating after a fortnight of palm collecting in 
Peninsular Ihailand. 

His thesis and earlier publications mainly on 
bark morphology, but auliior of numerous others 
on syslcmatics.' plant geography, conservation.* 
ecology, and miscellaneous subjects. Co-editor of 
an issue on National Parks, and chief editor of the 
'Tree l-Iora of Malaya'.* 

Whilmorcu Si I UMi R has been named after him. 

Cf>i.ii.CTiNCi i.ocai.itii-s. (cincndcd). 1957. 
Singapore with visits to SE. Jolioro and Pulai 
Mangrove Reserve (Jan. 5- Mar. 22). Malaya: 
.Sclangor, For. Res. Inst. Kepong; Kanching, 



Flora Malesiana 

[ser. I, vol. 8^ 

Serendah and Semangko For. Reserves (23-31); 
Pedis, S. Kedah and Penang (31-Apr. 8); Perak, 
Maxwell's Hill, Taiping; Bubu F. R., Kuala 
Kangsar; Lumut locality; Piah and Keladang 
Siong For. Reserves, Ipoh (9-16); Perlis (22-26); 
Selangor, F.R.I. Kepong (28-May 11); Negri 
Sembilan, Sungei Menyala F. R., Port Dickson 
(12-20); Negri Sembilan, Kuala Pilah Distr. (21- 
26); Selangor, Bt Lagong F. R. (27-June 9); 
Pahang, Kuantan Distr. (12-21); Trengganu, Bt 


Bauk F. R., Dungun (22); Kuala Trengganu Distr. 
(23-24); Johore, Mersing Distr. (July 18-Aug. 2). 
NW. Borneo, Sarawak: Kuching Distr. (Aug. 19- 
28); Brunei: Temburong Distr. (29-Sept. 14), Belait 
Distr. (Sept. 15-Oct. 4); N. Borneo: Labuan I. 
(Oct. 5-7); Lungmanis felling concession, 30 miles 
W of Sandakan (Oct. 8-12); around Sandakan 
(13-15); Kalabakan felling concession, 40 miles W 
of Tawau (16-25). Malaya (Nov. 13-20): Selangor, 
Kanching, Serendah, and Sungei Lallang For. 
Reserves. — 1962-64. In the Solomon Islands. — 
1965. Joined in Royal Society Expedition to 
Solomon Is. (July-Sept.) (c/. Corner). — 1966. 
Malay Peninsula: Pahang, Jengka F. R. & Kemasul 

F. R. (Feb.); Johore, Labis F. R. (Apr.); P. 
Penang (May); Selangor, G. Bunga Bua (May); 
Kedah & Perlis (June); Ulu Perak (July); Perak, 

G. Bubu (Aug.); Sarawak, Semengoh (Aug.); 
Perak, Slim Hills F. R. (Sept.); Perak, Kinta Hills 
(Oct.); Selangor, Genting Highlands (Nov.); 
Cameron Highlands (Dec.).— 1967. Perak. G. 
Bubu, Lumut F. R. and Pangkor I. (Feb.); 
Pahang, G. Benom (Mar.);* Pahang, Kuala 


Lompat (Apr.); W. Pahang and N. Johore (May); 
Selangor, Genting Highlands (May & June); 
Trengganu, Bt Bauk F. R. (June); Pahang, Tasek 
Bera and Palong F. R. (July); Kelantan near Gua 
Musang and Ulu Nenggiri (July); Kelantan, Ulu 
Lebir and Ulu Aring (Sept.); Selangor, Genting 
Highlands (Nov.).— 1968. Kedah, G. Inas F. R. 
(Feb.); Pahang, Taman Negara including G. 
Tahan (Feb.-Mar.);* Johore, Lenggor F. R., 
Kluang F. R. and G. Belumut (May); Kelantan, 
Kemahang F. R. (June); Trengganu, Mandi Angin 
Expedition (July 1-18); Selangor, G. Nuang 
(Aug.).— 1969. Johore, G. Ledang (Mt Ophir) 
(July); Kelantan, G. Stong (Aug.); Trengganu, 
G. Padang expedition (Sept.); Java: Tjibodas 
(Sept.); Pahang, Bt Tapah, K. Lompat and 
Maxwells Hills in Perak (Nov.); Langkawi Is 
(Dec.).— 1970. Negri Sembilan, Sg. Menyala F. R. 
(June); Pahang, Taman Negara, Ulu Sat, Ulu 
Sepia (July); Pahang, Cameron Highlands (Sept. 
and Oct.); Tapah Hills (Oct.).— 1971. Ulu Sungei 
Perak (Jan.); SW. Pahang and border Johore 
(Feb.); Pahang, Kuala Lipis to Cameron Highlands 
(May-June); Pahang, Taman Negara, and Treng- 
ganu, Ulu Sg. Trengganu (June); Trengganu, 
Kemaman, Perak and Taiping, Pahang, Fraser's 
Hill (Aug.); P. Penang (Oct.).— 1972. Ulu Sg. 
Trengganu (Feb.); S. Kelantan, G. Rabong (Mar.). 

Small trips with few collections have been 
omitted; above-mentioned trips have partly been 
made with Dransfield,* Shah, or Cockburn. 

Collections. Add: Collections 7957, num- 
bered TCW 1-686, distributed to Sing., Cambr., 
and Leyden (when fertile). General Malaysian 
collections (1965-72), mainly trees, numbered in 
the new FRI series (1-1000, 3001-4950, 8501-9000, 
12001-13000, 15001-16000, 20001-?) Herb. Ke- 
pong, principally distributed to Kew, Leyden, 
Sing., Arn. Arbor., Bog., Cambr. An important 
palm collection at Kepong, with virtually no dupl. 
In Malaya he especially collected in under-col- 
lected or/and hard to visit places. 

Collections from Borneo to Herb. Kuching and 
Sand, too; few numbers collected jointly with 
J. A. R. Anderson (see there) numbered in the 
Sarawak series; from Brunei joint collections with 
P. S. ASHTON, numbered BRUN 389-700, distri- 
buted by AsHTON (see there). 

In the Solomons (1962-64) about 4000 collections 
were made; during the Roy. Soc. Exp. (1965) 
numbered in the RSS series, in 1970 with TCW 
nos. Collections from Ecuador, New Hebrides, 
B.S.I.P. (1970) and Thailand in the TCW series. 

Literature. (1) Papers, but also a book: 
'Palms of Malaya' (Oxford Univ. Press 1973). 

(2) (with E. Soepadmo & A. Ho): 'National 
Parks of Malaysia' (special issue of Mai. Nat. J. 
24, 1971, p. 111-262). 

(3) In which besides author of numerous families 
(Publ. by Longman, London, 1972- ). 

(4) T. C. Whitmore: 'An outline of the Forest 
Zones on northeast Gunong Benom' (Bull. Br. 
Mus. (Nat. Hist.) 23, no 2, 1972, p. 11-15). See 
also 'Introduction' by Lord Medway, I.e. p. 3-7, 
2 maps, and paper on bryophytes and filmy ferns 
by Anne Johnson, I.e. p. 17-20, and photographs 
by Lord Medway. 

(5) J. Dransfield & T. C. Whitmore: 'Palm 
hunting in Malaya's National Park' (Principes 13, 



Cyclopaedia of Collectors — Supplement II 


1969, p. 83-98, 16 fig.); 'A Podostemacea new to 
Malaya, Indotristicha malayana' (Blumea 18, 

1970, p. 152-155, pi. l.fig. 1). 

•Wieringa, Wabe 

(1904, Rauwerd, Fr., Netherlands; 1965, 
Leyden, Netherlands) who worked for a short 
period at Wageningen, was attached to the technical 
staff (finally Head) of the Rijksherbarium at 
Leyden from Dec. 1928 onwards. From July 1949- 
Jan. 1956 he was placed at the disposal of the 
Buitenzorg Herbarium (interrupted by a leave in 
1952), in which period he distributed thousands of 
duplicates to the more important herbaria. 

Collections. He collected a few plants with 
J. H. Kern (see there) in the vicinity of Bogor, 
W. Java, numbered in the latter's series. 

Biographical data. Blumea 13, 1966, p. 
175-176, portr. 

Wiesenthal, Franz 

Add: Biographical data. In Streit & 
DiNDlNGER, Bibl. Missionum 21, 1955, p. 533-534, 

Wilde, Andre Gerard de 

Add: (b. 1925), M.D., Ph.D. medicine. Later on 
the staff of the Anatomical-Embryological Labor- 
atory of Leyden University, joined the Dutch 
New Guinea Expedition to the Sterrengehergte as a 
Physical Anthropologist in 1959. Since 1963 
Professor and Director of Anatomy and Embryol- 
ogy, University of Groningen. 

Collecting localities. 1959. Dutch Centr. 
New Guinea: Sibil Valley, Hollandia-Sentani (N. 
coast), Tsjop and Ok Bon Valley. 

Collections. Some plants (incl. mosses) with 
the collection of B. O. van Zanten, and C. 
Kalkman (see there), in Herb. Leyden. 

Biographical data. 'Who's Who in Science 
in Europe' 1, 1967, p. 414. 

•Wilde, Jan Jacobus Friedrich Egmond de 

(1932, Heemskerk, N.H., Netherlands; x) 
graduated in tropical forestry at Wageningen 
Agricultural College in 1958. Subsequently in the 
army for his number and in May 1959 sent to Biak 
near Dutch New Guinea as a lieutenant in the 
reserve. After return he joined the Laboratory for 
Plant Taxonomy and Geography at Wageningen, 
working on African Meliaceae. 

Collecting localities. 1959-60. Biak, N. 
of W. New Guinea. In Feb. 1960 at Manokwari, 
Oransbari etc. in NW. New Guinea. 

Collections. Nos 1-1071 collected at Cote 
d'lvoirc French W. Africa. In Malavsia, Biak no.s 
1150-1249 (before Feb. 1960), 1258-59. in New 
Guinea nos 1250-57 collected by Prof. Dr J. F. 
Kofji-S at tnarotali (Wissclmcrcn); Herb. Wage- 
nintfen. 76 dupl. (incl. single specimens) from 
Malaysia in I/erh. Leyden. Living orchids and 
seeds are cultivated at Wageningen. 

•Wilde, Willcm Jan Jacobus Oswald dc 

(\'Ji(), Heemskerk, Netherlands: x), botanist, 
educated at leyden University (Ph.D. 1971). 
From 1960-1961 Assistant of the Rijksherbarium, 
Leyden; in 1962 working on the Carmaui (Carai- 
bcan Marine Biological Institute), Curasao; 

1963-1966 staff member Agriculture University 
Wageningen (while in Africa for International 
Technical Assistance, The Hague); since 1966 staff 
member Rijksherbarium, Leyden. 

From his stay in the Cameroons (1963) onwards 
he invariably collected together with his wife, 
nee Duyfjes, Brigitta Emma Elisabeth (1936, 
Bandung, Indonesia; x), botanist, educated at 
Leyden University; presently at Agricultural 
University Wageningen, working on North 
African Allium. 

W. J. J. O. de WILDE 

Author of taxonomic papers on Najadaceae, 
Saraca (Caesalpiniaceae) and Passifloraceae (thesis 
on the genus Adenid), and of reports on the 
Netherlands Biological Expedition to Turkey 

Collecting localities. 1972. Indonesia. 
N. Sumatra: Atjeh. Exploration Gunung Leuser 
Nature Reserves:' Ketambc area (Orang Utan 
Orphanage) (r . 300 m) ( May 1 5-June 1 6) ; climbing 
G. Bandahara (17-28). large 'blang' c. 2800 m 
(25); Kctambe area (June 29-July 5); Pcmatang 
Siantar, Prapat (Lake loba) and G. Batu Lopang 
(F of Prapal) (July 8); Kctambe area (9-14); 
climbing Ci. Kctambe (2200 m) (15-20); Kctambe 
area (July 2 1 -Aug. I); expedition into the interior, 
SW of CJ. Kctambe (up to c. 2700 m), also climbing 
again G. Kctambe (r. 2200m) (3-18); Kctambe 
area (Aug. 19-Scpt. 3). 

Colli crioNS. Sumatra collections in Herb. 
Leyden (G. I. cuscr nos 12000 LMK8. 135(4-14456; 
Pcmatang Siantar. Prapat. (i. Hatu lopang. nos 
I348'> 135f)3); dupl. in Herb. Hog. Plants from 
Turkey. 19.59 (2154 /;.'v) also in I c\dcn. 

African collections ( 1963 6()) all in Wageningen, 
;• 10.000; numbered from I onwards. 




Flora Malesiana 

[ser. I, vol. 8^ 

Literature. (1) A report will be published in 
Fl. Mai. Bull, and in Mai. Nat. J. 

Biographical data. Curr. vitae in Thesis 
Leyden, 1971, p. 3^. 

Wiljes-Hissink, Elisabeth Adrienne de 

Collections. Add: In Herb. Wageningen: 
35 nos collected in Ambon and Ceram (Seram). 


*Willeinse, Reinierus Hendrikus 

(1938, Haarlem, Netherlands; x) schoolteacher 
at a MAVO (highschool) at Velzen, collected about 
10 plants, mainly Labia tae, in E. Java and Bali in 
mid-July 7970; in Herb. Leyden. 

Williams, Robert Statham 

Collections. Add: Also dupl. in Herb. 
Edinb. (1905). 

*Willis, James Hamlyn 

(1910, Oakleigh, Vict., Australia; x), botanist, 
B.S. University of Melbourne (1939). In 1972 he 
retired as Director of the Melbourne Herbarium. 

Collecting localities. 7970. E. New 
Guinea: Mts Wilhelm and Giluwe (June/July). 

Collections. Herb. Melbourne: ±500 phan- 
erogams, bryophytes, lichens and fungi from New 

Wind, Rutgert 
(d. 1964, The Hague, Netherlands). 

*Wing, James A., see Forest Dept North Borneo. 

Winkler, Hans 

Biographical data. Add: Mitt. Staatsinst. 
Allg. Bot. Hamburg 11, 1957, p. 7-8. 


Winterbottoni, James Edward 

Add: From 1834 onwards he made almost 
annually tours in Europe. From 1846-49 he 
travelled in company of Capt. Strachey (later 
Sir Richard) to India, Further India, China, 
Kashmir, Tibet, Nepal, and the Indian Archi- 
pelago. Early 1854 he started his tour to the Near 

Itinerary. Leaving England (Jan. 4, 1846) 
for Bombay. As it proved too late for a trip to the 
Himalaya, he determined to go on to Java (ar- 
riving June 20), and after making excursions about 
the island, returned at Batavia on July 24; by 
steamer (27) to Singapore (arrival Aug. 1). With 
the China mail to Hongkong and up the coast to 
Shanghai, returning by the same route to Singa- 
pore, proceeding to India and the Himalaya; 
home (about April 75-^9) by way of Egypt, Malta, 
and Gibraltar. 

Collections. Add: Also in Herb. Edinb. 
from Java (1846). 

Biographical data. Add: Lond. J. Bot. 6, 
1854, p. 345-349; Bonplandia 2, 1854, Nov. 15, p. 

*Winters, Harold Franklin 

(1913, Nowata, Oklahoma, U.S.A.; x) v\as 
educated at Oklahoma State University at Still- 
water (B.Sc. horticulture 1940), at Ohio State 
University, Columbus (M.Sc. horticulture 1942); 
graduate studies in Botany at the University of 
Maryland, College Park (1962-67). Assistant 
Orchard and Nursery Inspector for State of 
Oklahoma (1942-43); Horticulturist, Fed. Exp. 
Station U.S. Dept of Agric, Mayaguez, Puerto 
Rico (1943-56); from 1956 with the New Crops 
Research Branch, Plant Science Div., Agric. 
Research Service, U.S. Dept of Agric, Beltsville, 

During the New Guinea Expedition 1970 he 
was assisted by Dr Joseph J. Higgins (see there), 
collecting materials of possible ornamental value. 
The expedition was sponsored jointly by the 
Agricultural Research Service of the U.S. Dept of 
Agric. and Longwood Gardens, Kenneth Square, 

Itinerary. 7970. Australian New Guinea. 
Travelling by way of Hawaii, New Caledonia, 
and Brisbane, to Lae (Jan. 17-18); at Wau, Mt 
Kaindi, Edie Creek, Meri Creek (19-22); Patep 
village (24-26); Aiyura and Kassam Pass (Feb. 2- 
5); Markham River Valley near Lae (9-11); 
Mindik and Oreganum villages, Huon Peninsula 
(12-17); Botanic Garden Lae (19-24); Goroka 
(26-27); Chimbu country near Kundiawa, Mt 
Ekuam and Mt Hagen (Mar. 6); Mt Hagen area, 
Mt Kum, Baiyer R., Wabag Rd and Wahgi 
Valley (7-13); Lae (14-21); Port Moresby area. 
Brown R., Rouna Falls and Sogeri (22-31). 
Philippines, Luzon: Mt Makiling and cultivated 
plants from Univ. of the Philippines (Apr. 2-5); 
Bot. Garden Singapore (7); visit to the Institute 
for Horticultural Research at Djakarta, and the 
Botanic Gardens at Bogor and Tjibodas, in iV. 
Java, Indonesia (9-13). 

Collections. A new series. Winters & 
Higgins, was started, 873 nos, amongst which 
several Ericaceae. Rhododendron species identified 



Cyclopaedia of Collectors — Supplement II 


by Dr H. Sleumer. In Herb. U.S. Nat. Arbor. 
Wash.; dupl. of Ericaceae in Herb. Leyden. 

Biographical data. Amer. Men of Science 
11th ed. vol. 6, 1967, p. 5923. 

♦Wirawan, Nengah 

(1941, Tabanan, Bali, Indonesia; x) graduated 
from the College of Agriculture at Tjiawi, Bogor 
(B.Sc. Botany in 1962); Assistant Botanist of the 
Herbarium Bogoriense; 1965-67 Head of the 
Mountain Garden Tjibodas, and in those years 
part-time student in biology at the Bandang 
Institute of Technology; since May 1967 full-time 

Collecting localities. 7960. W. Java: 
Peutjang I. and Udjong Kulon Nature Reserve 
(2 weeks with Kostermans).— 1961. Sumbawa 
with Kostermans (see there) (Apr. 8-May 25); 
W. Java: Mt Hondje near Tamandjaja in Banten 
(= Bantam) (Dec.-Jan. 1962, 10 days).— 1963. 
Tjigenter and Tjilintjing, Udjong Kulon Nature 
Reserve (Apr. 13-27); Udjong Kulon Nature 
Reserve: Mt Pajung,^ trail from Tjibunar — San- 
giang Sirah (Dec. 25-Jan. 22, 1964).— 1964. 
Peutjang I. (Jan. 23-29); Bali: southern slope of 
Mt Batukau (Tabanan), from Pura Luhur, near 
Wongaja Gede, and summit Mt Batukau (July 
18-21). — 1965. Flores with Kostermans (see 
there): Waingapu (Simiba), near the harbour and 
the village Kembeniru (Apr. 4-5), Ende, Mt 
Wongge (-7), Mborong, S. coast W. Flores (8), 
stay at Kissol and on Mt Ndeki (9-14), Wae Rana 
(15), to Mbero and Nangarawa (16-17), Mbengan 
F. R. (19), Wae Rana-Kissol-Ruteng-Mano (20- 
23), N. slope .Mt Ranaka-Mandosawu complex 
(24-30), Mt (Potjo)Likang, S of Ruteng (May 2-3), 
Ruteng-Wae Mao Base, near Rana Mese (4), 
vicinity of Rana Mese (5-10), Wae Mao-Ruteng 
(11), road Ruteng-Pongkor and Mt (Potjo) Reok, 
S of Ruteng (13-14), Ruteng-Rio, N. coast W. 
Flores (18), Ponto II F. R. (19-20), vicinity of Reo 
(21-24); leaving Reo for Surabaja (26). 

Collections. About 500 nos and 969 joint 
numbers with Kostermans (K. & W. 1-969) from 
the Flores Expedition. Mostly in 6-10 sets, 
deposited in Herb. Bog., Leyden, Kew, Sing., 
Paris, Wash., Leningr., etc. 

Literature. (1) Wirawan : 'Beberapa tjatafan 
mcngenai keadaan vegetasi G. Pajung, Udjung- 
kulon' (Notes on the vegetation of Mt Pajung, 
Udjungkulon) (Laporan Seminar Biologi Pertama 
Tjiawi (Bogor) 1965. p. 247-257). 

Wit, Hcndrik Cornelis Dirk de 

Add: Onwards of Nov. 1953 Lecturer, later 
(1959) Professor of Botany at Wagcningon. He has 
specialized in African botany and in aquatic plants. 

Crudia denitii Kostermans has been named after 

Witsen, Nicolaas C. 

Add : He also had plants drawn in Japan. China, 
and Africa (cf. F. De Haan. Priangan, I, 1910, 
Pcrs. p. 282-284). 

Woerden, Corncillc Leon I-ambcrt Henri van 

rd. 1970. Utrecht. Netherlands). Add: Re- 
patriated in 1952; settled at Grocnckan near 
Utrecht, and in the employ of the Gcnctical 


Institute of the University at Utrecht from 1953- 

Biographical data. F1. Mai. Bull, no 25, 
1971, p. 1880. 

*Wolff, Torben Lunn 

(1919, Copenhagen, Denmark; x) was educated 
at the University of Copenhagen; Cand. mag. 
(zoology) 1947. He was a member of the Atlantide 
Expedition to West Africa 1945^6, the Galathea 


Expedition 1950-52, and the Danish Rcnnell 
Expedition 1951.' 

With the Noona Dan Expedition (see there for 
Itinerary, Collections, etc.Y he collected some 
botanical specimens in Yalon {Mtis.sau, Rism. 
Arch.) and Renncll (Solomon Is.) in 1962. In 1965 
and 1966 some more specimens in the Solomons. 

Litiraturi:. (1) Author of 'The Natural 
History of Renncll Island' (1955). 

(2) T. Woi.fe: 'The Noona Dan Expedition' 
(Nature 198. 1963. p. 1044-1045). 

Womersley, John Spencer 

Colli rTiN(i loi Ai iTiis. Add: Jn the fol- 
lowing years several liKal trips were made; in Nov. 
/V.5y short visit to VVext New Guinea. — 19()6. Kanni 
Valley with L. L. Hintv; Wau area, foothills of 
Giluwc, S. Highlands with K. R. W(k)||.iams; 
Okapa area. \\. Highland Dislr. with ("(K)I)|.— He 
will have certainly occasionally collected later. 

Literature. (2) Aild: with J. M. M< Adam: 
'The Forests and Forest Coiulilions in the Icr- 
rilorics of Papu.i and New (iuiiica' (prep, for llic 
Hrit. Common\^calth lor. Conf. in Austr.. 1957, 
Port Moresby). 



Flora Malesiana 

[ser. I, vol. 81 

Wong, Y. K., see Forest Dept North Borneo. 

Wood, Devillo D. 
Add: In 1969 he lived in California. 

Wood, Geoffrey Howorth Spencer 

Biographical data. Add: Proc. Linn. Soc. 
Lond. 169th sess., 1958, p. 38-40; Gard. Bull. 
Sing. 17, 1960, p. 498-501. 

♦Woods, M. G. 

collected in the KL series of the Phytochemical 
Survey of the Federation of Malaya, Kuala 
Lumpur, in S. Pahang, in March J959. 

*Woods, Patrick (Paddy) James Blythe 

(1932, Ballinderry, Co. Antrim, N. Ireland; x) 
attended a course for student gardeners at the 
Royal Botanic Garden, Edinburgh (1953-56) and 
was awarded the Diploma in Horticulture (DHE); 
joined the staff as a Scientific Assistant to B. L. 
Burtt; at present Experimental Officer, cur- 
rently working with the latter on Old World 
Gesneriaceae particularly on the genus Aeschyn- 
anthus, and on New Guinea Orchidaceae. In 1962 
he accompanied Burtt to Malaya and Sarawak. 
His New Guinea trip was financed by Mr Geoffrey 
Elworthy, an Australian, who wishes to en- 
courage taxonomic work on Orchidaceae. 

Collecting localities. 1962. Malaya and 
Sarawak, see Burtt's itinerary. Terr, of New 
Guinea & Papua: between Bulolo and Edie Creek 
(Oct. 3-13); Loloke (= Laloki) Gorge (17); 
Cape Rodney to Doma Village in the Musa Mts, 
Centr. Owen Stanley Range (20-24); Doma Village 
area and surrounding mist-forest ridges (26-Nov. 
25). — 1968. Trip to New Guinea with Mr Michael 
Black and C. E. Ridsdale (see there), also 
collecting in Java and Malay Peninsula. 

Collections. Herb. Edinb.: mostly Gesneria- 
ceae and Orchidaceae. A 3(X) numbers living 
material were collected. Dupl. herb. spec, will be 
distributed to Lae, Leyden, etc. 

*WoolIiams, Keith Richard 

(1940, Chester, England; x), graduate in 
Horticulture, Roy. Bot. Gard. Kew (1963); 
currently Curator, Botanic Garden, Lae, Terr, of 
New Guinea. 

Collecting localities. 1963. Sabah ( = 
North Borneo), live plants; Talibu, Southern 
Highlands, New Guinea, with J. S. Womersley. — 
1968. Mt Wilhelm in New Guinea, living collections. 

Collections. Mostly living plants, but dried 
material with Womersley in Herb. Lae; dupl. also 
in Edinb. 

*Worobai, see Boswezen Nieuw Guinea. 

Wray, Leonard 

Collections. Add: Also dupl. in Herb. 
Edinb. {ex Calc. and Kew). 

♦Wright, K. G. 

collected in the NGF series at Sawette sawmill 
near Salamau, Papua (1959); Herb. Lae. 

Wyatt-Smith, John 

Add: He made an exploration of Mt Tahan in 
Feb. 1960, accompanied by B. A. Mitchell, 
Wong Yew Kwan, and some others. 


He retired early in April 1963 under the Malay- 
anisation Scheme and accepted an appointment 
under FAO sponsorship at Ibadan University in 
Nigeria. At present settled in England. 

Literature. (2) Add: 'A Survey of the peat 
swamp research plots in naturally regenerated 
forest' (Mai. Res. Pamphl. no 39, 1961, 55 pp., 
map); 'Ecological Studies on Malayan Forests' I 
{Ibid, no 52, Dec. 1966, 20 pp., 6 fig., 33 tab., 26 
app. tab.). 

Wycherley, Paul Renoden 

Add: (1928, Bromley, Kent, England; x) was 
educated at the University College, London 
(1946-52), B.Sc. (Hons) Botany (1949), Ph.D. 
(1952); in 1951/52 guest student at Wageningen 
Agricultural College in the Netherlands. In 1953 
Botanist, and from 1965-71 Head, Botany Division 
of the Rubber Research Institute of Malaya. 
Sometime President Malayan Nature Society and 
Chairman Terrestrial Conservation Section, Inter- 
national Biological Programme, West Malaysia. 
Intensive travel throughout Great Britain and 
West Malaysia; extensive travel in Thailand, East 
Malaysia, Australia, Japan, Canada, Central and 
South America, and Europe. An ardent advocate 
for and author on nature conservation in Malaysia, 
especially the Malay Peninsula. In 1972 Director of 
King's Park and Botanic Garden, Perth, W. 

Collecting localities. Mostly in plan- 
tations to study the weed floral A few collections 
fro.m forest and limestone hills, especially Sungei 
Buloh and Batu Caves, Selangor respectively. 

Collections. Few specimens in Herb. Rubb. 
Res. Inst. Malaya, Sing, and Kew. 

Literature. (1) Author of 'Identification of 
plants on Malay Rubber Estates' (Plant. Bull. 
Rubber Res. Inst. Malaya 89, 1967, p. 55-65, 8 pi. 
(65-72)); 'Ground covers in plantation crops' 
(Symp. Ecol. Res. Humid Trop. Veget. Kuching 
1963, 1965 p 94-119, 6 fig., 11 tab.); 'Vegetation 
of Rubber Plantations' (J. Rubb. Res. Inst. 
Malaya 16, 1960, p. 87); 'The Spread of Croton 
hirtus' (Proc. Humid Trop. Symp. Tjiawi 1958, 
1961, p. 279-280, 2 fig.); 'Ridley and Batu Caves' 
(Mai. Nat. J. 25, 1972, p. 22-37, pi. 6-10, incl. 
Check list flora); and numerous other papers. 


collected in Sarawak onwards of 1957; Herb. 
Kuching; dupl. a.o. in Herb. Edinb. 

*Yamada, Kindi 

collected Sonneratia at Bandjermasin, SE. 
Borneo (Sept. 10, 1942). 

*Yasue, Yasunobu 

Entomologist, graduated in the Faculty of 
Agriculture, Kyoto University, 1941. Associate 
Professor of the Ohara Institute for Agricultural 
Biology, Okayama University, Kurashiki, Japan. 

Member of the Kyoto University West Irian 
Expedition, Aug. 1963-Mar. 1964. 

Lived at Surabaya before 1940. Collected a few 
plants, very fragmentary. 


is cited by Ohwi (in MS), as collector of a 
Digitaria in Celebes; no date, nor locality. 



Cyclopaedia of Collectors — Supplement II 


Yvan, Melchior 

Literature. (2) Add: C. Montagne: 'Note 
sur quelques plantes cryptogames recueillies en 
Chine pendant le sejour qu'y fit notre ambassade, 
et communiquees par M. le docteur Ivan' (Rev. 
Bot. Duchartre 2, 1846-47, p. 363-365). 

f •Zahir bin Yusoff, Mohammed 

Assistant of the Forest Research Institute at 
Kepong, collected 50 numbers in Pahang (G. 
Benom etc.), Malaya, in Apr. 1967. 

♦Zanten, Bernard Otto van 

(1927, Oudemolen, Dr., Netherlands; x), 
bryologist, educated at Groningen University 
(Ph.D. 1959). After having taken his doctor's degree 
he joined the Dutch New Guinea Expedition to the 
Sterrengebergte, and subsequently paid a 3-months 
visit to New Zealand. At present on the staff of the 
Botanical Laboratory of Groningen University, 
working on his moss collection. In 1969 he made a 
half-year bryological collecting trip to Australia, 
New Zealand, Guadalcanal, New Guinea, Luzon, 
and Japan, sponsored by WOTRO. 

Collecting localities. 1959 (Apr.-Sept.). 
Former Dutch New Guinea} N. coast: HoUandia 
and Ifar; S. New Guinea; at Tanah Merah and 
Kaliwin on Digoel R.; Ajerok; on the Kau R. at 
Bian-Katem, Wombon, Koekoeboen, Anoendjam- 
dit, Aremko, Oemkoeboen, and Oejambiep; still 
farther to the north at Amiol and Katem, Songgan 
and Arimkop; Sibil Valley; Mt Antares (1300- 
3400 m). — 1968. Bryological exploration (June- 
Dec.):* E. New Guinea: from Port Moresby 
exploring (June 8-13) Brown R. area, Musgrave 
R., Sogeri Plateau, Siritumu Lake; from Lae in 
Suru R. area (15-16); Wau (Mac Adam Memorial 
Park, Mt Kaindi) (17-19); Goroka (Marafunga, 
Daulo pass) (21-23); Mt Wilhelm (June 25-July 
10); Mt Hagen (Baiyer R., Jimmy R.) (July II- 
13); Lae (Situm, 16). Accompanied by D. N. 
MacVean and W. A. Weber (see those). Subse- 
quently in Australia, New Zealand and the British 
Solomons, returning to E. New Guinea: Goroka 
(Daulo Pass) (Oct. 13-15), GembogI (16), Mt 
Hagen (18-22), Mt Giluwe (23-27), Mt Wilhelm 
(Oct. 29-Nov. 4), and Owen Stanley Range 
(Efogi) (Nov. 10-12), with R. Johns. With Mrs 
VAN Zanten-Tuntelder in the Philippines: 
Luzon: Mt Makiling (Nov. 19), Baguio (Mt 
Cabuyo, Mt S.Thomas, Asin) (20-21), Bontoc(Mt 
Data, Mt Polis, Sagada) ill-lA), Baguio (Mt 
Cabuyo, 25), and in Japan (Nov. 3()-Dcc. 13). 

Collections. About 670 New Guinea nos, 
1959, of which the greater part was collected in 
Sibil Valley and on Mt Antaris. About H of the 
material consists of Musci,^ the remaining part of 
Hepaticae, Fungi. Lichens, and Atgac. Only few 
phanerogams, collected at Tanah Merah, in Sibil, 
and the Antares. His collection includes some 
plants collected by Dr W. VtRVfiORT and Dr A. G. 
DP. Wilde (sec those), and in collaboration with C. 
Kalkman. In Herb. Leyden; at present Musci and 
Hepaticae in Groningen for revision, after which 
the 1st SCI V ill go to Ixydcn too. while a dupl. set 
will remain in Hcrh. Groningen. The phanerogams 
arc separately numbered. 

In New Zealand in 1959 about 600 nos were 
collected of which some 200 phanerogams; in 

Herb. Leyden: lichens and 1 set of phanerogams; 
another set in Groningen, and the 3rd set in his 
private herbarium. The mosses will remain at 
Groningen, but a dupl. set will be sent to Leyden 
after identification. 

The 1969 collection consists almost exclusively 
of bryophytes, of which those from the Philippines 
and Japan are collected with his wife, Mrs H. H. 
van Zanten-Tuntelder. A few Hymenophyllums 
were brought home and some phanerogams from 
Guadalcanal (Solomon Is.). 

Literature. (1) B. O. van Zanten: 'Mosses 
of the Star Mountains Expedition, Central New 
Guinea' (Nova Guinea, Bot. no 16, 1964, p. 263- 
368, pi. 22-36; for localities and dates cf. p. 264- 

(2) Cf. complete itinerary in Fl. Mai. Bull, no 
24, 1969, p. 1788. B. O. van Zanten: 'Verslag 
van een verzameltocht naar Australisch Nieuw- 
Guinea, Australia, Nieuw-Zeeland, Guadalcanal, 
Luzon, en Japan, 5 juni-17 dec. 1968 (1970) 66 pp., 
mimeogr., folio. 

*Zeano, see Forest Dept North Borneo. 

Zeeuw, Carl de 

(1912, East Lansing, Mich., U.S.A.; x) received 
an arts education and forestry training in Michigan 
State College; subsequently to Syracuse for work 
in wood anatomy under Dr H. P. Brown at the 
New York State College of Forestry (M.S.). 
During the war he served with the U.S. Army Air 
Force, being stationed in Australia in 1942 and 
thereafter in New Guinea in the Port Moresby 
area, then inland from Buna Gona at Dobodura in 
Papua, and finally in the Markham valley in the 
Mandated Territory of New Guinea. 

Upon return from service he resumed work at 
Syracuse on the Ph.D. in the investigation of the 
reaction of wood anatomy and physical behaviour; 
at present Professor of Wood Products Engineer- 
ing at Syracuse. 

Collections. Mainly woody plants, partly 
from the Northern Territory of Australia (15), 
the others (37) from New Guinea (1943-45). In 
Herb. Brisbane (dried coll. -*- wood samples). 
Herb. Melbourne (wood samples in the For. Prod. 
Lab. C.S.I. R.O.), and at N.Y. State College (wood 
samples). The herbarium material of the last lot is 
still in his hands and in due time will be deposited 
in the Am. Arbor. Fern collections (about 15) and 
some other herbaceous plants in Herb. Brisbane. 

Zeijlstra, Hcin Hidde 

(d. 1961. Haarlem. Netherlands). 

Zieck, J. F. U. 

Correct for J. F. H. Zieck (sec Suppl. Cvcl. Fl. 
Mai. I, 5. 1958. p. cccxxxvi). In 1962 he left New 
Guinea. In 1968 working for the I orest Products 
Research Centre, Boroko, Port Moresby, Papua. 

Author of several mimeographed internal 
reports on the forests of New Guinea. 

ZnllinKcr, Hcinrich 

Coi.i.Ec rioNs. Add: Some of his ferns from 
Java, arc also cited as Sporlfdir. Java. Evidently 
at least fern duplicates in Herb. Sporiidir (sec 

Biographic Ai. data. Add: Pctcrm. Gcogr. 
Mitt. 3. 1857, p. 210; ibid. 4. 1858. p. 39. 




As in the Cyclopaedia and the 1st Supplement, a selection has been made of geographical names (mostly 
islands, lakes, mountains, rivers). The larger islands, e.g. Sumatra, Java, Borneo, New Guinea, Luzon, 
will only exceptionally be found in the index as the result would have been a host of numerals behind 
them, difficult to handle and most of the references being hardly worthwhile; for a short survey of the 
larger Malesian units see p. (3). The Solomon Islands have been omitted as falling beyond the scope of 
Flora Malesiana. 

Names of ships, expeditions, and persons are sometimes included. 

The abbreviations placed in italics behind the names are the following: 

d = district 
/ = island(s) 
/ = lake(s) 

m = mountain 

p = place 
pen. = peninsula 
r = river 
s = ship 

Reference. Reference is made to the parenthesized Arabian pagination, instead of to the Roman 
numerals. The letters a and b following the page numbers indicate the left and right column respectively 
of the page. 

Aas m 102a 

Abi r 101b 

Abilala m 85a 

Adelbert Range 80b 

Adi / 13b, 52b, 57a, 67a 

Admiral Gardner s 88a 

Adzare Range 82b 

Agung m 29a 

Aifat r 67a, 91a 

Ajamaroe /101b 

Alas r 103a 

Albert Edward m 36b, 80b, 82b 

Amboaki r 91a 

Amboina, see Ambon 

Ambon / 58b, 93a, 105a, 108a 

Ambua m 52b 

Amdutakin m 52b, 102a 

Amorwange m 82b 

Anadi r 39a 

Anak Takun m 75a, 83b 

Andrassy m 47b, 65b 

Anggi / 39a, 76b, 91a/b, 94b, 

100b, 101b 
Antares m 52b, Ilia 
Antop m 101a 
Api m 22b 
Api Benarat m 23a 
Apo m 10b, 40a, 54a, 70a, 79b, 

80b, 88b, 105a 
Apo Batu Buli Range 72a 
Apoy m 40a 
Apriri r 101b 
Archbold / 28b 
Archbold Expeditions 16b, 24b, 

Arfak Mts 38b, 52b, 57a, 91a, 

Aroa r 78a 
Asmat region 13b 
Astrolabe Range 53a, 78a, 87a 
Atjeh 30b, 68a/b, 107b 
Aunde /82b, 91a 
Aur / 64a 

Babuyanes / 78b, 82a 

Bahiri r 78a 

Baiaio r 9b 

Baiyer r 18a, 39a, 108b, Ilia 

Bako Nat. Park lla/b, 21a, 26b, 

30a, 38b, 48a, 49b, 53a, 67b, 

70b, 78b, 89a, 92b 
Baku, see Bako 
Balabac /71a, 104b 
Balambangan / 104b 
Bali / 14a, 29a, 56a, 57a, 58b, 

68b, 93a, 97a, 108a, 109a 
Bali(e)m Valley 25b, 42b, 57b, 

60b, 87a, 101a 
Baling m 92a 
Balleh /• 73a 
Baluran m 45a 
Bami r 39a 
Banahao m 73b 
Bandahara m 107b 
Bangeta m 82b 
Banguey / 104b 
Baram r 10a, 22b, 37a, 53a 
Barigi /• 45b 
Barija r 20b 
Barker, Miss R. 92a 
Basilan / 39b, 73b, 79b 
Batan / 12b, 26a, 39b, 42a, 78b 
Batanes / 26a, 78b, 82a 
Batanta / 13b 
Batjan / 105a 
Batu Brinchang m 59a 
Batu Lawi m 72a 
Batu Lopang m 107b 
Batu Tibang m 10a 
Batukau ni 109a 
Batulanteh m 57a 
Batur / 68b 
Batur m 100a 
Bauk m 106b 
Baung m 29a 
Bawean / 45a 
B(e)lumut m 23b, 30a, 39b, 84b, 

89a, 106b 
Bena r 92b 
Bencoolen d 30b 
Benom m 51a, 106a 
Bensbach r 80b 
Beremban m 70b 
Beribai m 101b 
Berumbun m 22b 
Berumput m 13a, 19a, 92b 
Besar m 84a 



Biak / 39a, 57b, 67a, 70b, 87b, 

100b, lOla/b, 107a 
Biaru r 78a 
Bier A- 101b 
Biliran / 26a 
Billiton / 45a 
Biota m 82b 

Bismarck Mts 23a, 50b, 96a/b 
Bisschop, G. 75a 
Blumut, see Belumut 
Bohayan / 47b 
Bohol / 56b 
Bongao /71a 
Bonite s 38a 
Bosavi m 85a 
Brangkat m 89a 
Bremer, Capt. 81b 
Brinchang m 22b, 69a, 70b 
British Museum— Newcastle 

upon Tyne Exp. 50b, 78a, 

84b, 104a 
Bromo m 68b, 73a 
Brong m 89a 
Brown r 26a, 39a, 41b, 70b, 78a, 

80a, 87a, 98a, 101b, Ilia 
Bubu m 22b, 48a, 88b, 106a 
Bujong Malacca m 9b 
Bumbu r 70b 
Bunga r 41b 
Bunga Bua m 106a 
Bunguran / 46b 
Burep r 41b 
Burias / 56b 
Burke, L. R. 60b 
Bus(s)u r 12a, 70b, 82b, 98a 
Butibum r 41b 
Buzu, see Bus(s)u 

Cabuyo m Ilia 

Cagayan Sulu / 56b 

Calamangan / 56b 

Calayan / 78b 

Cambridge Kinabalu Exp. 92a 

Cameron Highlands 9a, 10b, 
15b, 22b, 30a, 39b, 41a, 48a, 
53b, 69b, 70a/b, 76a, 78b, 
90a, 91b, 106a/b 



Select index — 2nd Supplement Cyclopaedia 


Camiguin / 26a, 42a, 70a 

Canlaon m 39b, 70a 

Capul / 56b 

Carstensz Mts 25b, 29b, 46a 

Catanduanes / 70a 

Cebu / 56b 

Celebes / 50b, 59b, 65a, 84a, 

91b, 105a 
Ceram / 58b, 93a, 108a 
Chabang Tiga m 84a 
Challenger s 27a 
Chimbu Vallev 16a, 38b, 75a, 

91a, 95a, 103b 
Chin See Chung 84a 
Chini m 23b 
Christmas / 10b, 67a 
Clav r 39a 

Colijn, Dr A. H. 29b 
Concord m 40a 
Conflict group 38b 
Coquille s 30b 
Corregidor / 35a 
Credner /71a 

Crocker Range 9b, 23b, 67b 
Cromwell Mts 45b 
Crownprince Range 82b 
C.S.I.R.O. — Rijksherbarium 

Exp. Kubor Range 78a, 101b 
Cyclops Mts 56b, 82b, 90b 

Dafonsero m 91a 

Dammi / 64b 

Danau ... see 34a 

Danfu r 25a 

Data m 63b, 73b, 90b, 95a, Ilia 

Deagan / 56b 

Dempo m 73a 

D'Entrecasteaux / 38b 

Destacado / 56b 

Dickson m 42a 

Dieng m 68b, 100a 

Digoel r 100b, II la 

Djambi d 30b 

Doma Peaks 36a, 38b, 52b, 78a 

Doole p 59a 

Dua /31a 

Duke of York / 67a 

Dyaul /71a 

Ekuam m 108b 
Elandora m 16b, 50b 
Emira / 25a 
Endeavour s 12b, 25a 
Engelhart, M. 60b 
Enggom m 82b 
Erap r 39a 
Eravc r 78a 

Faita r 90b 

Fakfak r 101b 

Fara r 21a 

Finistcrrc Mts 43a. 50b, 75a, 

78a. 84b. 98a. 104a 
Flore? / 57b, 61b, 86b, 100b, 

My r 66b. 74a. 80b 
I rcybourg Pass 85a 

Gading m 19a, 22b, 89a, 92b 
Gaharu m 19a 

Gajah m 72b 
Galathea Exp. 70b 
Garatun m 27a 
Gamier, Marietta 36a 
Gebeh / 77b 
Gedeh-Pangrango m 14b, 30b, 

43b, 49b, 56a, 57b, 58a, 70a, 

71b, 75b, 102b 
Geelvink Bay 87b 
Geger Bintang m 30b 
Geli Mutu m 61b 
Genofa m 100b 
Genting Highlands 106a 
Gigantanga / 56b 
Giluwe m 25a, 63a, 78a, 80b, 

87a, 103b, 108a, 109b, Ilia 
Gogol r 39a, 66b, 101b 
Goldie r 78a 
Golo / 56b 
Gombo(c)k m 49a 
Goodenough / Ua 
Guimaras / 82a 
Guiting-guiting m 79b 
Gwamongga m 91b 

Haga r 25b 

Halmaheira / 105a 

Handeulum Sisi / 90a 

Harcourt j' 81b 

Herbert r 66b 

Hermit /71a 

Herzog Mts 50b, 85a, 104a 

Hijau m 46b, 69a, 89a 

Hinde, A. 92a 

Hindenburg Range 52b, 102a 

Hindenburg-Dap Range 36b 

Homoraon / 56b 

Hondje m 57b, 109a 

Hose Mts 10a, 11a, 19a 

Hunstein Range 45b 

Hydrographers Range 20b, 78a 

Idjen m 61a, 68b 

Iglit m 63b 

Inasm 18b, 106b 

Iran r 92b 

Iraya m 12b, 42a, 78b 

Isarog m 73b 

Ischon r 91a 

Islands in Milne Bay 38b 

Itbayat / 78b 

Jabau r 91a 

Jacobsen, G. 92a 

Jahn, Mr 57b 

Jalimo r 84b, 90b 

Jang m 45a 

Japen ; 57a, 87b 

Jasar m 22b, 70b 

Java / I la. 76b 

Jclakoi m 18b 

Jcrai /;; 48a. Ma. 83b 

Jim(m)i or Jimmy Valley 41a, 

64a. 66b, 82a, Ilia 
Job / 87b 
Jolo/3Ia. 56b 
Juliana m 98b 

Kahanov, Or 57b 
Kairuli m 16b. 41b, 51b. 91a, 
103b. 108b. Ilia 


Kaironk r 18a 

Kajang m 1 la 

Kalimantan Selatan d 102b, 

Kana m 47a 

Kangean Islands 45a 

Kantz m 102a 

Kanyi m 92b 

Kar Kar / 66b, 80b 

Karimondjawa / 45a 

Kaur Ilia 

Kaugel r 1 6a 

Kebar Valley 87b, 91a, 101b 

Kedah Peak 64b 

Kelabit Highlands 10a 

Kelai r 57b 

Kelobo / 52a 

Kemaboe r 26a 

Kemiri m 68b 

Kemp Welch r 70b, 105a 

Kerewa m 52b 

Kerigomna m 39b 

Kerinci or Kerintji m 30b, 34a 

Ketambe m 103a, 107b 

Keyan m 47a 

Kidayap m 58b 

Kidul m 56a 

Kikori r 85a 

Kinabalu m 9a/b, 10a, 15b, 22b, 
26b, 37a, 47a/b, 49b, 53a/b, 
56b, 65b, 67b, 71b, 74a, 84a, 
89a, 91b, 92a/b, I04b, 105a 

King, Capt. P. P. 81a 

Kinkain m 78a, 102a 

Kiriwina / 36b 

Kode /83b 

Koebre m 9 1 b 

Kofiau / 60b 

Komodo / 67a, 83b 

Korongl r 75a 

Koronon m 79b 

Kosim Kardin, M. 34a 

Krabo m 57a 

Krakatau / 45a 

Kratke Mts 16b, 41b. 50b 

Krotky s 74b 

Kubor Range 78a, lOlb 

Kujabu r 91a 

Kukukuku ranges 87a 

Kum m 108b 

Kummc m 82b 

Kumusi r 20b 

Kuscmun ni 91a 

Kutai Reserve lOa, 30b. 93a 

Kutubu / 78a, 87a 

Kwapcna, Nava 36a 

Kyoto Univ. Biol. Exp. to SE. 
Asia 49a. 89a 

Kyoto Univ. W. Irian Exp. I lOb 

I abuan / 65b, 68a. 82b, 104b. 

!05a, I06a 
I.agaip r K7a 
Lagong in 53a 
Lajang r 92b 
Laka'an ni 53b. 86b 
LakAn. sec Laka'an 
I.akci / 89a 
I.ala z 21a 

I.aloki r 39a. 41b. 78a, 105a 
Lambert. A. B. 81b 



Flora Malesiana 

[ser. I, vol. 8^ 

Lambir m lib, 19a, 37a, 53a 

Lampong Distr. 30b, 50a, 67a 

Langkawi / 9b, 18b, 22a/b, 38a, 
41a, 51a, 54b, 83b, 95b, 99a, 

Latimodjong Mts 31b 

Laut / 46b 

Lawas Distr. 92b, 104b 

Lawes m 39a 

Lawit m 23 b 

Lebir m 23b 

Ledang m 83b, I06b 

Legaspi m 58b 

Lei r 52b 

Lensemoi m 101a 

Leronr41b, 82b 

Leuser m 103a, 107b 

Leyte / 26a, 37a, 39b, 56b, 70a 

Liicang m 109a 

Limbang Distr. 92b, 104b 

limestone localities 18b, 22b, 
23a, 29a, 36a/b, 48a, 49b, 
54b, 57b, 59a, 67b, 78b, 80b, 
83b, 84a, 102b, 103a, 110b 

Lipuun / 26a 

Lomblem (= Lomblen) / 61b, 

Lombok / 97a 

Long /■ 66b 

Lord Duncan s 88a 

Lou /■ 89b 

Low, Hugh 62a, 105a 

Lowry, J. B. 92a 

Lubi m 39b 

Luf /71a 

Lumaku w 71b 

Lumut Hills 37a 

Luzon / 70a 

Mackenzie, Colin 60b 

Madoera /61a 

Maja m 23a 

Makanoi m 91a 

Makatarem m 78b 

Makiling m 70a, 73b, 75a, 79b, 

82a, 108b, Ilia 
Malabar m 75b 
Malengelo / 36a 
Mamad, Mr 57b 
Mambare r 20b 
Mandi Angin m 23b, 106b 
Manurep m 100a 
Manus i 36b, 71a 
Mapia / 101b 
Maquiiing, see Makiling 
Marei Parei Ridge 104b 
Markham r 12a, 16b, 39a, 70b, 

82b, 84b, 98a, 108b 
Marklo / 36a 
Masbate / 56b 
Mat(t)ang m 30a, 49b, 53a, 67b, 

84a, 91b, 92b 
Matsubara, M. 44b 
Matupi m 82b 
Maxwells Hills 106b 
Mayon m 70a, 73b 
Medamit r 92b 
Medang m 48a 
Mendi r 16a 
Mentawa m 22b 


Meos Noem / 57a, 87b 

Meratus m 33a 

Mersing m 73a 

Meru Betiri Reserve 45a 

Merurong Plateau 17b 

Mesenuk m 91a 

Mesilau r 91b 

Michael m 16b, 24b 

Milyin Kolyin m 102a 

Mindanao / 58b, 63a, 70a, 73b, 

79b, 80b, 95a, 98b, 105a 
Mindoro / 56b, 58b, 63b, 93b, 

Misool / 76a 
Mitchell, B. A. 110a 
Moller s 53a 
Mongremotti m 91a 
Monoglito m 101b 
Morehead r 78a 
Morotai / 57b 
Motupore / 36b 
Mullankassan / 68a 
Mulu m 10a, 19a, 22b, 47a 
Murik / 66b 
Murray / 66b 
Murray Pass 80b, 82b 
Murud m 10a, 19a, 71b 
Musa Mts 32a, UOa 
Musgrave r 36b, 41b, Ilia 
Mussau / 23a, 71a, 109b 
Mutis m 86b 

Naersen, K. van 61a 

Nago /71a 

Naho / 85a 

Naho r 84b 

Namance / 25a 

Naranjo / 56b 

Natuna / 33a, 46b 

Ndeki m 57b, 109a 

Ne m 52b 

Negros / 39b, 56b, 70a, 73b 

Nerimbau m 101a 

Nettoti w 91a, 101a 

New Britain / 25a, 26a, 35b, 

36b, 39a, 43a, 50b, 55a, 61a, 

66b, 70b, 71a, 74a/b, 80a, 

82b, 85a, 95b, 104a 
New Guinea Border Exp. 38a, 

43a, 68b, 80a, 93a 
New Hanover / 25a, 71a 
New Ireland / 24b, 25a, 31b, 

36b, 50b, 52a, 53b, 66b, 71a, 

84a, 95b 
Njaboeti /101b 
Njapa m 57b 
Noona Dan Exp. 23a, 29b, 56a, 

71a, 72b, 109b 
Nuang m 106b 
Nunukan / 74a/b 
Nusa / 36b 
Nusa Baru / 56a 
Nusa Kambangan / 68b, 93a 
Nusalik / 36b 
Nuttall, Thomas 15b 

Obi / 105a 
Obree m 23a 
Oereb r 87b, 101b 
Oiamadawa'a /n 11a 

Okapa r 38b 
OUerenshaw, David 36a 
Oriomo r 63a 
Orion Mts 52b 
Otto m9a, 16b, 24b 
Owen Stanley Range 21a, 23a, 
32a, 53a, 78a, 87a, 110a 

Padang m 106b 

Padang Lawas 49a 

Padar / 83b 

Padas r 104b 

Pajung m 57b, 109a 

Palawan / 26a, 39b, 56b, 65a, 

71a, 73b, 79b, 104b 
Panay / 79b 
Pandjang / 101b 
Pangicor / 106a 
Pangrango, see Gedeh-Pang- 

Panti m 20a, 23b, 42a, 49a, 84a 
Papahag /71a 
Papandajan m 58a, 75b 
Pasu m 37a 
Patep r 41b 
Pekeglbaro m 101b 
Pelagus r 92b 
Pemanggil / 18b 
Penang / 20a, 22b, 33a, 38a, 

48a, 49a, 54b, 58a, 69b, 70a, 

73a, 84a, 95b, 104b, 106a/b 
Penck m 36a. 
Penibukan Ridge 71b 
Penrissen m 49b 
Peutjang / 22b, 35a, 57a, 90a, 

93a, 102b, 103a, 109a 
Philippine — Hawaii Exp. 56b 
Piora m 20b, 43a 
Poe m 13a, 19a 
Polis w 73b, 90b, Ilia 
Pondok m 84a 
Pondok II m 89a 
Pongani r 78a 
Prahau ( = Tangkoeban Prahoe) 

m 56a 
Preussische Expedition 79b 
Prince Albert Range 78a 
Prince Alexander Range 80b 
Prinseneiland / 45a, 57a 
Puan m Ala. 
Pueh Range 92b 
Pulai m 18b, 20a, 49a, 83b 
Pulau = island, see under 

second word 
Pulog m 23b, 50a, 79b, 81a. 

Pulusari m 15a 

Rabong m 89a, 106b 

Raja m 32a 

Rajang, see Rejang 

Ramu r 25a, 36a, 39a, 77b, 80b, 

84b, 109b 
Rana Mese / 57b, 109a 
Ranaka m 109a 
Ranoe Upas / 34a 
Rapet m 84a 
Rara m 90b 



Select index — 2nd Supplement Cyclopaedia 


Rattlesnake s 62b 

Raveni m 90b 

Rawang ... see 34a 

Rawlinson Range 20b, 45b, 82b 

Raya m 73a, 83b 

Reedtz Thott m 53b 

Rejang r 49b, 74b 

Reok m 109a 

Rindja / 83b 

Rio Ampalit r 61b 

Romblon / 40a, 79b 

Rossel / 43 a 

Rote (Roti) / 86b, 100b 

Royal Society Expeditions 22b, 

26a/b, 44b, 84a, 89a, 92b, 

Rijksen, H. 103a 
Rijksherbarium New Guinea 

Exp. 90b 

Sabal Tapang r 92b 

Sabtan(g) / 26a, 78b 

Sakar / 47b 

Salahutu m 58b 

Saiak m 30b, 43b, 90a 

Salawaket, see Sarawaket 

Salim m 47b 

Samar ;" 40a, 63b, 70a, 79b 

Samperolat m 57a 

St. Cruz / 71a 

St. Joseph r 85a 

Santo T(h)omas m 73b, 90b, 

Santubong m 37a 
Sapamoro /71a 
Sarawaket (Saruwaket) m 10b, 

20b, 41b, 45b, 82b 
Sarempaka m 33a, 102b, 103a 
Saru-mot /n 91b 
Saruwaket, see Sarawaket 
Satuma / 18b 
Schrader Range 18a 
Scratchley ni 25a 
Sebat Kechil r 92b 
Sedili Kechil r 23b 
Sega r 57b 
Segaham Range 49b 
Selabor m lOa 
Sclang m 92b 
Selingan / 23b 
Sembilan / 46b 
Scnang / 18b 
Scnjavin .v 76b 
Scnsencmcs m 91b 
Sentani / 57b, 70b, 9Ia, lOOb, 

Separation Point 65a 
Scpik r 28b. 3Va. 44a, 45b, 66b, 

78a/b, 80b, 84b 
Serangoon r 58a 
Sewic m 36b 
Sheridan, I)r J. 92a 
Shungol m 42a 
Siasi /' 56b 
Sibium Range 78a 
Sibutu / 56b 
Sibuyan / 40a, 79b 
Sidocarsi Mts 87b, 101 b 
Sierra Madrc Range 50a 
Silon m Mb 

Simbai r 23a 

Simpson Massif 27a, 53a, 78a, 

Singapore / 20a, 47b, 58a, 104b 
Sipingan / 34a 
Siritumu /Ilia 
Situ Bagendit / 58a 
Situ Gunung / 34a 
Skeat Exp. 40a 
Slim Hills 106a 
Snake r 39a 
Sohoton /// 40a, 63b 
Solor / 61b 
Star Mts 43a, 52b, 69b, 80b, 

98b, 101a, 107a 
Stong/M 18b, 106b 
Strickland r 66b 
Strong m 25a, 55a, 97a 
Subis in 37a 
Suckling m lla, 36b, 60a, 78a, 

95b, 100a 
Sujarta, Mr 57a 
Sulphur s 44b 
Sulu/31a, 64b, 105a 
Sumba / 49a 
Sumbawa / 33b, 49a, 57a/b, 

58b, 76b, 109a 
Supiori Island 67a 
Suru r 11 la 
Swart Valley 13b 

Taal Volcano 73b 

Tabanan m 109a 

Tabanio r 49a 

Tabar / 52a 

Tabauwan / 47b 

Tabayoc m 79b 

Tafa m 36b 

Tahan m 30a, 69b, 93a, 105a, 

106b, 110a 
Takun m 32b, 67b 
Talawe m 36a 
Tale r 87a 
Talomo m 80b 
Tamrau Range 91a 
Tanggamus m 50a 
Tangis m 35b, 36a 
Tangkuban Prahu m 30b, 56a 
Tapah hills 106b 
Tapis m 18b, 23b 
Tapul / 56b 

Tari Subdistr. Exp. 102a 
Taskul / 25a 
Tataan / 64b 
Tau Range 73a, 89a 
Tauri r 39a 
Tawar / 34a 
Tawau Hills lOa 
Tawitawi / 51b, 64b, 71a 
Ichak r 87a 
Fcbu m 88b 
Tcnibrok m 91b 
Fcmpurong m 9b, 72b 
Fcnajan r 93a 
'Icnggcr m 30b 
'Fcnom rn 67b 
Tcraja ni 47a 
Tcrnatc / lib, 105a 
Thursday / 98b 
1 icao / 56b 

Timah m 44a, 49a, 53a, 58a, 

67b, 70a, 93a, 98a 
Timbun Mata / 23b 
Timor / 10b, 14a, 28b, 34b, 44b, 

53b, 56b, 59b, 65b, 84b, 86b, 

91b, 92b, 105b 
Tinjar r 37a 
Tioman / 38a, 84a 
Toba/68b, 107b 
Tobi //I 91a 
Tokiri Range 91a 
Tor r 101b 
Torricelli Mts 28b 
Trist / 43a 
Trobriand / 36b, 38b 
Trusmadi m 65b, 7 lb 
Tuan Sket m 89a 

Udjong Kulon pen. 20a, 22b, 
30b, 31a, 39b, 45a, 57b, 58b, 
76b, 90a, 93a, 102b, 103a, 

Ulu Kali m 36b, 39b 

Umi r 82b 

Undup r 9b 

Ungaran m 61a 

Uranie s 38a 

Ursula / 56b, 71a 

USDA Sugar Cane Exp. 39a 

Vanapa r 78a, 82b 
Vega s 55a 
Victoria m 21a, 79b 
Vikerman, Commander 61a 
Vulcan / 82b 

Wadimana m 53a, 78a, 87a 

Wahgi Valley 16a, 78a, 108b 

Waigco(u) / 60b, 82b, 105a 

Walker m 67b 

Wamena r 87a 

Wamsuf Massif 91a 

Warida r 39a 

Warjori r 91b 

Warrapuri r 82b 

Warsamson r 102a 

Watjctoni /;; 1 01 a 

Waumi r 91a 

Wckari r 91a 

Wharton Range 82b 

Wilhclm ;;; 10a, 12a, 15b, 16b, 
32a. 34a. 46a. 50b, 63a. 66b, 
75a. 80b. 82b. 84b. 91a. 94b, 
95b. 99a. 103b. 104a, 105b. 
I08a. llOa. Ilia 

Wiliiclmina //; 87a 

WisscI l.akcs 39a. 87b, 101 b 

Wondiwoi Mts 57a. 87b 

Wong Yew Kwan 1 lOa 

Wongge m IWii 

Worcester .v 74a 

Wosi r 39a 

Vanicb, Andrew 36a 
Yellow r 66b 
Yodda r 21a 
Yule / 78a 


Zendilerong r 26a 




HYPERICACEAE^ (N. K. B. Robson, London) 

Trees, shrubs or perennial or annual herbs. Leaves simple, opposite and decussate 
(Mai. spp.), entire (Mai. spp.). sessile to shortly petioled, often with ± translucent 
and sometimes black or red glandular dots and/or lines. Stipules 0. Inflorescences 
terminal and sometimes axillary, very rarely axillary only, cymose to thyrsoid 
or rarely racemose, bracteate at least initially, l-'rv-flowered. Flowers bisexual, 
actinomorphic, homostylous or heterodistylous. Sepals 5 (Mai. spp.), free or 
± united, imbricate, entire or with margin variously divided and often glandular, 
lamina glandular like the leaves, usually with greater proportion of glands linear 
rather than punctiform, persistent (Mai. spp.). Petals 5 (Mai. spp.). free, imbricate 
(contorted), alternisepalous, entire or with margin variously divided and often 
glandular, lamina usually glandular like the leaves, sometimes with nectariferous 
basal appendage, glabrous (Mai. spp.), caducous or persistent. Stamen fascicles 
5 (Mai. spp.), epipetalous, free or variously united, each with 1-cv) stamens; 
filaments variously united or sometimes apparently free, the free part usually 
slender; anthers 2-thecal, dorsifixed, often with gland terminating connective. 
Staminodial fascicles 3 or (Mai. spp.), when present alternating with stamen 
fascicles. Ovary 1, superior, 5-3-celled or 1 -celled with 5-2 parietal placentas; 
styles 5-3 (2), free or — united, ± slender; stigma punctiform to capitate; ovules 
00-2 on each placenta (Mai. spp.), anatropous, horizontal or ascending. Fruit 
capsular (Mai. spp.), dehiscing septicidally or loculicidally. Seeds cv-1 on each 
placenta, sometimes winged or carinate; embryo cylindric, straight or curved, 
with cotyledons longer to shorter than hypocotyl; endosperm absent. 

Distribution. There are 7 genera with c. 550 spp., cosmopolitan except for Arctic regions 
and most of Polynesia, but only Hypericum and Triadeniim occur outside the tropics and im- 
mediately adjacent areas. Of the three tribes, the Vismieae (3 genera) occur in Africa (including 
Madagascar) and America, the Cratoxyleae (3 genera) in Madagascar, Indo-Malesia, E. Asia 
and NE. America, and the Hypericeae {Hypericum) throughout most of the range of the family 
except for most lowland tropical areas. In Malesia only two genera are present: Cratoxylum 
Bl. and Hypericum L. 

Ecology. Apart from Hypericum and Triadenum, the genera are confined to lowlands or 
intermediate altitudes in the tropics and subtropics, in ± mesophytic habitats. Hypericum 
tends to be a high altitude mesophytic genus in the tropics; but some species occasionally 
grow at low altitudes. In temperate regions it ranges widely in altitude and habitat, but never 
occurs under extremely arid conditions. Many species of Hypericaceae are tolerant of a wide 
range of soil conditions, but others (especially some herbaceous species of Hypericum) are 
restricted to particular soil types. 

Pollination. Flowers o^ Hypericum are nearly all open-pollinated, being visited only for 
pollen by e.g. Syrphid flies and Bomhus spp. Specialized pollination with nectar secretion and 
sometimes dimorphic hcterostyly has evolved twice in Hypericum and also occurs in Cratoxylum 
and Eliaea (sec below, under Morphology). 

Dispersal . Seeds of the Hypericeae and Cratoxyleae are small and sometimes have wing-like 
expansions of the testa which would tend to promote wind disjicrsal. Those without such an 
aid are normally dispHirscd by gravity; but the seeds of some species of wet habitats may be 
carried in mud on the feet of wading birds. Birds or other animals arc instrumcnla! in the 
dispersal of those few species of Hypericum (in four distinct parts of the j;cnus) in which the 

(I) As will appear in the treatment, Hypericaceae do not deserve family rank and should be 
viewed as a subfamily of Guttiferae. It is merely for convenience that this name is maintained 


Flora Malesiana [ser. I, vol. 8^ 

normally capsular fruit has become ± baccate, as well as those of the Vismieae, where the 
fruit is always baccate or drupaceous. 

Morphology. The three tribes of the Hypericaceae can be distinguished by several floral 
characters, as follows: 
Vismieae: Perianth 5-merous; petals adaxially pubescent; staminodial fascicles 5; stamen 

fascicles 5; ovary 5-merous; fruit baccate or drupaceous. 
Cratoxyleae: Perianth 5-merous; petals glabrous; staminodial fascicles 3; stamen fascicles 3 

{i.e. 2 +2 + 1); ovary 3-merous; fruit capsular. 
Hypericeae: Perianth 5-4-merous; petals glabrous; staminodial fascicles absent or very rarely 3; 

stamen fascicles 5-4 (free or variously grouped or united); ovary 5-2-merous; fruit capsular 

or rarely baccate. 

The grouping of the stamens has attracted much attention. (For a discussion of this question 
see RoBSON, Trans. Bot. Soc. Edinb. 41, 1972, 365-383, and works cited therein). Evidence 
from morphology, vascular anatomy and ontogeny indicates that in this family as well as the 
Guttiferae, the androecium basically comprises two diplostemonous whorls of stamen fascicles. 
In the Hypericaceae the episepalous fascicles are sterile or absent, whereas the epipetalous 
ones are fertile and may be free {Vismieae, Hypericum pro parte), united 2+2 + 1 {Crat- 
oxyleae, Hypericum pro parte) or all united {Hypericum pro parte). The individual filaments of 
each fascicle may be united for over i^ of their length or less; or they may arise independently 
from the receptacle, so that the androecium appears to be afascicular. The number of stamens 
in each fascicle varies from 80-100 to 1 in Hypericum, plants with an androecium in the latter 
state typifying the genus Sarothra L., which Linnaeus placed in his group Pentandria. These, 
however, cannot be recognized as distinct from Hypericum. 

Some species of Cratoxylum, and some specimens of the monospecific genus Eliaea, have an 
appendage at the base of the petal ('petal scale') which encloses nectariferous tissue {cf. Hoch- 
REUTiNER, C. R. Soc. Phys. Hist. Nat. Geneve 35, 1918, 82-85), and some of these exhibit 
dimorphic heterostyly, indicating a trend towards specialized insect pollination. This pollination 
syndrome also occurs exceptionally in Hypericum (see Robson, I.e.). It is associated with stiff" 
erect sepals, resulting in an eff"ectively tubular corolla. In such flowers, Hochreutiner {I.e.) 
has shown that the sterile episepalous stamen fascicles ('hypogynous scales') may act like 
lodicules by swelling and thereby expanding the perianth whorls. 

Anatomy. Research on the anatomy of the Hypericaceae has been summarized in Metcalfe 
& Chalk (Anat. Dicot. 1, 1950, 165-169), while a more detailed discussion of floral and vege- 
tative anatomy of the genera Eliaea and Cratoxylum will be found in P. Baas (Blumea 18, 1970, 
369-391). Some of the most important anatomical papers for systematic purposes concern the 
nature and distribution of the secretory system that is found throughout the Hypericaceae and 
Guttiferae. Owing to a photosensitive reaction induced by hypericin ('hypericism'), the bio- 
chemistry of this and related substances, as well as the clinical details, have been much studied 
(review in J. M. Watt & M. G. Breyer-Branderwijk, Med. & Poison PI. Southern Africa, ed, 
2, 1962, 495). The most recent summaries of the biochemical studies on hypericin and its distri- 
bution in Hypericum, both morphologically and systematically, are by C. Mathis & G. Ouris- 
SON (Phytochemistry 2, 1963, 157-171) and C. Mathis (Etude Chimiotaxonomique du genre 
Hypericum L., These Fac. Pharm. Univ. Strasbourg. 1963). The form and distribution of 'black' 
glands has proved to be of taxonomic importance in Hypericum, while in Cratoxylum the form 
of the glands in the petals is of sectional significance. Other recent papers concerning anatomical 
topics in the Hypericaceae are few, but that by E. K. Schonfield (Mem. N.Y. Bot. Gard. 18, 1, 
1968), on petiole anatomy in the Guttiferae and related families, may be mentioned. 

Taxonomy. The tribes Vismieae, Cratoxyleae and Hypericeae are frequently treated as the 
subfamily Hypericoideae of the Guttiferae, from which they differ by no one constant character, 
and so the rank of subfamily may well be the most appropriate one. The combination of the 
following characters, however, make it immediately recognizable : 

Flowers bisexual. Antesepalous stamen fascicles sterile or absent; epipetalous fascicles free 
or variously united; stamens with filaments slender, free or partly united, and anthers small, 
dehiscing longitudinally. Ovary with styles 2-5, ± elongate, free or partly or rarely wholly 
united and placentae 2-5, axile to parietal, each with l-~ ovules. Fruit capsular or baccate or 

1974] Hypericaceae (Robson) 3 

drupaceous. Seeds exarillate; embryo with cotyledons free, not incrassate, and hypocotyl 
— slender. Germination epigeal. Leaves usually with venation ± reticulate; glandular canals 
often interrupted or replaced by globular lacunae ('punctate glands'), often coloured dark red 
or black by hypericin. 

Cytology. The basic chromosome number for the Hypericeae {i.e. Hypericum) appears to 
be n = 12, from which there is a descending series along several separate evolutionary lines to 
n = 7 or, possibly, 6 (N. K. B. Robson & W. P. Adams, Brittonia 20, 1968, 95-106). Tetraploids 
occur on all the base numbers; but the only higher degrees of 'ploidy' recorded (pentaploidy and 
hexaploidy in H. perforatum L. and its hybrids) are associated with a partially apomictic breeding 
system. Only four species in the other two tribes have known chromosome numbers; two 
unspecified species of Vismia {Vistnieae) have n = 10, whereas in the Cratoxyleae, Cratoxylum 
formosum (Jack) Dyer has n = 7 and Triadenum virginicum (L.) Rafin. n = 19. Structural 
hybridity occurs in two American species of Hypericum, H. punctatum Lamk and H. mitchel- 
iianum R\T)B. (Robson & Adams, I.e.). 

Chemotaxonomy. Hypericaceae were treated as Guttiferae-Hypericoideae in my 'Chemo- 
taxonomie der Pflanzen' 4 (1966) 223, to which the reader is referred for references of work 
published before 1965. 

Our chemical knowledge of Hypericaceae is based largely on Hypericum. A great variety of 
polyphenolic compounds seems to be characteristic of Hypericum and related genera. Among 
them catechins, leucoanthocyanins including leucodelphinidin, chlorogenic acids, flavonols and 
condensed tannins very often occur in large amounts. The quercetin glycosides rutin, quercitrin 
and hyperin are very common in Hypericaceae; hyperin (= quercetin-3-galactoside) was 
isolated in 1938 from Hypericum perforatum and named after this plant. Most characteristic, 
however, are anthraquinonoid and xanthonoid pigments. They are mainly located in schizo- 
genous cavities which are present in roots, stems, leaves and flowers. Two photodynamically 
active emodin-type naphthodianthrones, hypericin and pseudohypericin, seem to occur in all 
members of the sections Euhypericum sensu R. Keller, Carnpylosporus and Campylopus 
(monotypic; H. cerastoides(^ rhodopeum) only), but not in other sections of the genus Hypericum. 
Hypericin occurs probably also in roots of Psorospernutm guineense but seems to be lacking in 
species previously placed in Ascyrum. Celebixanthone was isolated from the bark of Cratoxylum 
sumatranum ( = celehicum) and Haronga (= Harungaua) madagascariensis contains anthra- 
quinones, anthrones and xanthones. Chrysophanol, physcion, madagascin, madagascinanthron, 
harunganin, haronginanthron and euxanthone were isolated from its bark. Very recently 2,7'- 
biemodyl, a new type of dimeric anthraquinones, was extracted from its leaves (D. G. Buckley 
et at. Austr. J. Chem. 25, 1972, 843). Maculaxanthone, a complex xanthone derivative has been 
isolated from roots of Hypericum maculatum (P. Arends, Tetrahedron Letters 1969: 4893) and 
uliginosin-A and -B were extracted from whole plants of the Mexican Hypericum uliginosum; 
both compounds are isoprenylated and acylated phloroglucin-dcrived phenols with strong anti- 
biotic activity (H. L.Taylor &R. M. Brooker, Lloydia32, 1969,217). Eliaeaarticulata coninins 
quercitrin, leucocyanidin and condensed tannins; anthraquinones are absent but the xanthone 
mangiferin is present in its leaves (H. Jacquemin et al. PI. Med. Phytotherapic 3, 1969, 196). 
Mangifcrin occurs also in leaves of Hypericum humifusum (Ph. Lebreton & M.-P. BoucHEZ, 
PI. Med. Phytotherapic 1, 1967, 188). 

Species of Hypericum do also contain essential oils stored in schizogcnous cavities. A special 
feature of these oils is the presence of appreciable amounts of aliphatic compounds like 2- 
mcthyloctane, nonane, undecane, octanal and decanal. 

Free tritcrpcnes seem to be common in roots and barks. Friedclin and bctulinic acid were 
isolated from the bark of Harungaua madagascariensis and recently bctulinic acid was shown 
to be a constituent of rootbark of Hypericum inodorum Mill. ( datum Air.) and //. andro- 
.laemuni L. (K. R. HARdRLAVis et al. Phytochcmistry 7, 1968, 331). Saponins {i.e. glycosylated 
triterpcncs) and alkaloids seem to occur rarely in Hypericaceae. 

Chemically Hypericaceae arc closely related to Ciuttiferae. The latter produce highly charac- 
teristic pigments in their resins, olcorcsins or 'laticcs'. The pigments arc complex polyisoprenyl- 
atcd and acylated compounds derived from phloroglucin (the so-called coumarin-type nco- 
flavonoids), bcnzophcnoncs and xanthones. Besides, they produce biflavonoids in wood and 

Flora Malesiana [ser. I, vol. 8^ 

leaves. This group of plant constituents was believed some years ago to be practically restricted 
to gymnosperms. Constituents like the uliginosins, euxanthone, mangiferin, celebixanthone and 
maculaxanthone connect Hypericaceae chemically intimately with Giittiferae. 

The same is valid for the preponderance of friedelan-type and kipan-type (lupeol, betulin, 
betulinic acid) compounds among triterpenes and for the patterns of simple phcnolics. 

From the phytochemical point of view there is absolutely no need to separate Hypericaceae 
from Giittiferae ( = Chisiaceae), i.e. to postulate family rank for Hypericoideae. — R. Hegnauer. 

Embryology. Embryo sac development in Hypericum is of the Polygonum type. Endosperm 
development is at first free-nuclear, and a chalazal cyst is produced at the 8- or 16-nucleate 
stage. This is later absorbed in the rest of the endosperm, which then becomes cellular (c/. 
A. N. Rao, Phytomorphology 7, 1957, 36-45, for details and some earlier references). Apospor- 
ous embryos occur in over 90% of the seeds of H. perforatum L. (K. L. Noack, Z. Indukt. 
Abstamm. Vererb. 76, 1939, 569-601). In Triadenum the embryo sac development resembles 
that o^ Hypericum, and apospory has been reported (O. Myers, Amer. J. Bot. 51, 1964, 664). 


1. Trees. Flowers red, pink or white. Leaves without red or black glands. Stamen fascicles alternating 
with hypogynous scales. Capsule loculicidally 3-valvcd. Seeds winged .... 1. Cratoxylum 

L Herbs or shrubs. Flowers yellow, often tinged red. Leaves with translucent glands and sometimes 
with black or red glands. Hypogynous scales (in Mai.) absent. Capsule (in Mai.) septicidal. Seeds 
(in Mai.) not or scarcely winged 2. Hypericum 


Blume, Verh. Bat. Gen. 9 (1823) 174; Bijdr. (1825) 144; Mus. Bot. Lugd. Bat. 2 
(1852) 15; Corner, Gard. Bull. S. S. 10 (1939) 21; Gogelein, Blumea 15 (1967) 
453. — Elodea (non Juss., nee MiCHX, nee Elodes Adans.) Jack, Mai. Misc. 2 
(1822) 21. — Hornsehuehia {non Nees) Blume, Cat. Gew. Btzg (1823) 15; ef. 
Steen. Bull. Jard. Bot. Btzg III, 17 (1948) 459. — Hyperieum seet. Tridesmos 
Choisy in DC. Prod. 1 (1824) 546. — Tridesmis {non Lour.) Spach, Suit. Buff. 5 
(1836) 358; Ann. Sc. Nat. II, 5 (1836) 351; Bl. Mus. Bot. Lugd. Bat. 2 (1852) 
18. — Ancistrolobus Spach, Suit. Buff. 5 (1836) 360; Ann. Sc. Nat. II, 5 (1836) 
352. — Fig. 1-10. 

Trees or shrubs, glabrous (in Mai.). Bark at base of stem exuding yellow resinous 
sap, hardening black. Branchlets compressed, 4(-6?)-lined, with interpetiolar 
scars linear, closed (continuous and straight or curved upward) or open (inter- 
rupted and curved downward) (fig. 1). Leaves opposite or rarely subopposite, entire, 
sessile or petioled, often with fine translucent ('pale') glandular dots, without black 
or red glands. Inflorescences terminal, racemiform, paniculate to axillary racemes 
or single flowers. Flowers (except staminodial fascicles and gynoecium) 5-merous, 
sometimes heterodistylous. Sepals quincuncial-imbricate, coriaceous, persistent, 
with longitudinal pale or black glandular lines or dots, sometimes elongating in 
fruit. Petals deep crimson to pink or white, sometimes tinged orange or green, with 
reddish or black glandular dots or lines (fig. le), sometimes with nectariferous 
scale-like, basal appendage (fig. le, 6f, 8e-f), caducous to subpersistent. Stamen 
fascicles united 2+2 + 1, single one epipetalous and double ones episepalous, 
glabrous, caducous, each with cv) stamens; filaments crimson to white, slender, 
united for over half their length; anthers crimson to white, ± shortly oblong to 
rhombic, dorsifixed, sometimes with 1-2 brown resiniferous glands terminating the 
connective. Staminodial fascicles 3, alternating with the fascicles, varying in shape 
and size (fig. 6g, i, j, 8g, i). (Flowers with 4-5-merous inner whorls occasionally 
occur). Ovary incompletely 3-celled; styles 3, free, ± slender, stigmas small, ± 
capitate. Ovules oo~4 on the basal half of each placenta, erect or ascending. Fruit 
a loculicidal, 3-valved, ± ligneous capsule, with slightly prominent longitudinal 


Hypericaceae (Robson) 

Fig. I . a-d. Scheme of development of terminal bud in Cratoxylum, showing the origin of the inter 
petiolar scar; in this case the latter is 'continuous'. — C. sumatramim (Jack) Bl. ssp. sumatianum. e. 

Petal from inside, appendage absent, x 3. 

vittae. Seeds r<j-A on each woody placenta (columella), cylindric to ovoid, imbricate; 
testa not sculptured; winged unilaterally or all round; embryo cylindric, straight, 
with cotyledons longer than the hypocotyl. 

Distr. 6 species, occurring in continental Asia from India (east of 92°E) east to S. China (south of 
c. 24''N and including Hong Kong and Hainan) and throughout Malesia, but not in the Moluccas and 
New Guinea. Fig. 2. 

Ecol. In various habitats, from dense primary or secondary forests to grassland, and on various soils, 
occurring from sea-level to c. 1000 m or rarely to 1800 m. 

Morph. The bodies described above as staminodial fascicles have been the subject of much investi- 
gation. Morphological and anatomical evidence indicates that they are sterile members of the cpiscpalous 
androecial whorl (see Robson, Trans. Bot. Soc. Edinb. 41, 1972, 375, 381); but, whereas the five fertile 
members of the epipetalous whorl are still present (four of them united in pairs to form double fascicles 
opposite sepals), the three sterile members of the episepalous whorl arc ail single organs i.e. two members 
of this whorl do not develop. 

Hcterodistyly occurs regularly in sect. Trulesnios and C. cochiiicliincmis and may be an occasional 
feature of other species. It is also present in the closely related genus Eliaea. This tendency towards 
specialized insect pollination is accompanied by the development of nectariferous petal appendages 
(absent in sect. Cratoxylum). In addition, the flowers of those species in which hcterodistyly is most 
pronounced tend to be pseudo-tubular, i.e. to have erect rigid sepals which prevent the petals from 
spreading. In these species (at least), the sterile fascicles probably act like the lodiculcs of grasses by 
swelling and thus aiding the expansion of the sepals and petals from the bud (see HocuRiauiNi.R, C. R. 
Soc. Phys. Hist. Nat. Geneve 35, 1918. 85). 

Uses. Species oi sect. Cratoxylum and Tridesmos have hard, heavy wood. C. sumatramim is good for 
making charcoal and implements. C. cochincliinense has fairly durable wooil. but is not used much for 
timber. Species o^ sect. Tridesmos, on the other hand, give a llcxible durable timber, which is sometimes 
used by Indonesians for construction. The species of sect. IsopteryKium have rather soft, red wood that 
splits badly and blunts tools on account of its high silica content. The Dayak of Indonesian Mornco, 
however, make drums of old hollow trunks of C. arhorescens. 

Ht YNh (Nutt. PI. 1927, 1080) stated that the bark at the base of the trunk of C./ormosum produces a 
resinous exudate (at first golden yellow, then red, finally black) that is used as a remedy for scabies and 
leg wounds. 

Notes. Cratoxylum is closely related to, but distinct from the Madagascan monotypic genus Lliaea 
Cambkss., according to GfXii;uiN (1967) and Haas (Hlumca 18. 1970. 369 391). It is intermediate in 
distribution between that genus and Triadenum Raiin., a herbaceous genus that has been wrongly 
included in Hypericum but appears to have been derived from Cratoxylum. Triadrnum has an I-. Asia-l-. 
North American disjunct distribution (Assam, Taiwan. Japan, Korea. Siberia eastern U.S.A. and 
r .mada). 

This account is based largely on the revision of the genus by CiociELF.iN (1967). 


Flora Malesiana 

[ser. I, vol. 8^ 


1. Leaves usually deciduous, with nerves ± curved, distinct, uniting or not but not forming an intra- 

marginal vein. Seeds unilaterally winged. Inflorescence terminal and/or axillary. Petals with or 

without nectary scale, with punctate or linear glands. 

2. Petals without nectary scale, with linear glands. Inflorescence terminal and sometimes axillary. 

Staminodial fascicles (if well developed) recurved or cucullate. Leaves with nerves not arched or 

uniting. Sect. Cratoxylum. 

3. Inflorescence paniculate, many-flowered, terminal and sometimes also axillary. Leaves very rarely 

glaucous beneath. Columella '/i to nearly \i as long as capsule 1. C. sumatranum 

3. Inflorescence of I-5-flowered cymules, terminal and axillary or axillary only. Leaves nearly always 
glaucous beneath. Columella basal 2. C. cochinchinense 

2. Petals with nectary scale, with punctate glands. Inflorescence axillary on old or the base of young 
shoots. Staminodial fascicles (if well developed) never recurved. Leaves with nerves arched, uniting. 
Sect. Tridesmos (Choisy) Dyer. 

4. Capsule 4-6 mm 0. Seeds (7-)12-17 per cell, obovoid-cylindric, 2-A mm wide, with wing narrow 
at the base. Leaves 3li-14 cm long, oblong to elliptic or more rarely obovate . 3. C. formosum 

4. Capsule 2-4(-5!/^) mm 0. Seeds 4-6 per cell, cylindric to ovoid-cylindric, \\i-2 mm wide, with wing 

not narrowing at the base. Leaves 2-9 mm long, elliptic to obovate 4. C. maingayi 

1. Leaves evergreen, with nerves straight, obscure, uniting to form an intramarginal vein. Seeds winged 

all round. Inflorescence terminal. Petals with minute nectary scale and punctate glands. Sect. Iso- 

pterygium Engl. 

5. Seeds 10-13 per cell. Leaves acute to acuminate, 2-4 times as long as broad, obovate-oblong to 

obovate-oblanceolate or elliptic, not or scarcely papillose beneath, not yellow or orange beneath. 

5. C. arborescens 
5. Seeds 4-7 per cell. Leaves rounded to obtuse-apiculate or rarely acute, usually 2 times as long as 
broad, elliptic, papillose-glaucous in the vein alveoli and usually yellow to orange beneath. 

6. C. glaucum 

1. Cratoxylum sumatranum (Jack) Blume, Mus. 
Bot. Lugd. Bat. 2 (1852) 16; MiQ. Fl. Ind. Bat. 1, 2 
(1859) 516; Suppl. (1861) 194; Corner, Card. Bull. 
S. S. 10 (1939) 35; Gogelein, Blumea 15 (1967) 
459. — Elodea sumatrana Jack, Mai. Misc. 2 
(1822) 22. — Hornschuchia hypericina Bl. Cat. 
Gew. Btzg (1823) 15. — C. hornschuchii Bl. Verh. 
Bat. Gen. 9 (1823) 174; Korth. Verh. Nat. Gesch. 
Bot. (1842) 178; Bl. Mus. Bot. Lugd. Bat. 2 
(1852) 15; MiQ. Fl. Ind. Bat. 1, 2 (1859) 516; K. &. 
V. Bijdr. 5 (1900) 134; Gagnep. Not. Syst. 1 
(1909) 20. — C. clandestinum Bl. Mus. Bot. Lugd. 
Bat. 2 (1852) 15; MiQ. Fl. Ind. Bat. 1, 2 (1859) 
516; K. & V. Bijdr. 5 (1900) 132; Gagnep. Not. 

Fig. 2. Range of the genus Cratoxylum. 

Syst. 1 (1909) 20; Back. Schoolfl. Java (1911) 
87; Engl, in E. & P. Nat. Pfl. Fam. ed. 2, 21 
(1925) 183. — C. racemosum Bl. Mus. Bot. Lugd. 
Bat. 2 (1852) 16; Miq. Fl. Ind. Bat. 1, 2 (1859) 
516; K. & V. Bijdr. 5 (1900) 135; Back. Schoolfl. 
Java (1911) 87; Engl, in E. & P. Nat. Pfl. Fam. 
ed. 2, 21 (1925) 183; Back. & Bakh./. Fl. Java 1 
(1963) 383. — C. celebicum Bl. Mus. Bot. Lugd. 
Bat. 2 (1852) 16; Miq. Fl. Ind. Bat. 1, 2 (1859) 
516; Merr. Philip. J. Sc. 4 (1909) Bot. 293; En. 
Philip. 3 (1923) 76; Meuer, Bot. Newsbull. Herb. 
For. Dept. Sandakan 7 (1967) 64. — Ancistrolobus 
floribundus TuRCZ. Bull. Soc. Nat. Mosc. 31 
(1858) 382. — C. arboreum Elmer, Leafl. Philip. 
Bot. 8 (1919) 3084. — C. hypericinum (Bl.) Merr. 
En. Born. (1921) 392; Back. & Bakh./. Fl. Java 
1 (1963) 383. — Fig. 3a-d. 

Tree or shrub, to 35 m tall, deciduous to sub- 
deciduous, glabrous; bark fissured, peeling in 
strips, dark brown; young shoots with interpetiolar 
scar continuous. Leaves sessile or with petiole up 
to 1 5 mm, 4-1 8 by 2-7 cm, elliptic to ovate-oblong, 
apex rounded to cuspidate, base subcordate or 
rounded to attenuate, herbaceous to chartaceous, 
sometimes glaucous beneath. Inflorescence a 
foliate panicle, often large; pedicels \^A-5 mm. 
Flowers homostylous. Sepals 3-6^/^ by 3-4!^ mm. 
Petals dark or brownish red to brick red, pale 
green at base, 4—9 by lM-4mm. Stamen fascicles 
2\4-l^A mm long, with stamens rt congested, c. 
120 per fascicle; anther gland absent. Staminodial 
fascicles (if well developed) yellow, up to 3 mm 
long, flattened, oblong to obovate, cucullate. 
Ovary l'/^-3 mm long; styles H^-3 mm. Capsule 
7-10 by 3-5 mm, c. 1-3 times as long as the 
sepals, cylindric, with columella basal to half as 
long as capsule. Seeds 3-10 per loculus, 5-7H by 
l'/2-2mm, oblanceolate to oblong. 

Distr. SE. Asia and Malesia: Sumatra, Java, 


Hypericaceae (Robson) 


Lesser Sunda Is. (Bali to Sumbawa), Borneo, 
Celebes, and the Philippines. Fig. 4. 

Note. C. suniatranum comprises 3 subspecies, 
one of which (ssp. neriifoliiitn (Kurz) Gogelein) 
occurs from Bengal to Thailand and Laos. It 
differs from the Flora Malesiana subspecies in 
having leaves always sessile to subsessile and 
sometimes glaucous beneath, always relatively 
few-flowered terminal and axillary panicles, petals 
bright red to brick red, and sepals in fruit 8-9 by 
4-5 mm, almost as long as the capsule. In contrast, 
ssp. suniatranum and blancoi have leaves sessile or 
petiolate (to 15 mm), usually relatively many- 
flowered terminal (and sometimes axillary) 
panicles, petals dark red to brownish red, and 
sepals in fruit 3-7 by 2^ mm, up to U as long as 
the capsule. 


1. Leaves oblong to lanceolate, l-AV^ times as 
long as broad, apex acuminate to cuspidate. 

ssp. sumatranum 

1. Leaves elliptic to elliptic-oblong, VA-lM times 
as long as broad, apex rounded or, more rarely, 
acute to shortly acuminate. . . ssp. blancoi 

ssp. sumatranum. — Fig. 3a-c. 

Leaves oblong to ovate-lanceolate or lanceolate, 
2-A\i times as long as broad, 5-18 by 2-5 cm, apex 
acute to cuspidate, base rounded to attenuate. 
Panicle variable in size. Capsule 7-12 by 3-5 mm, 
with sepals up to H as long. Seeds 3-10 per loculus. 

Distr. Malesia: Sumatra, Java, Lesser Sunda 
Is. (Bali, Lombok, Sumbawa), Borneo, Celebes, 
Philippines (Luzon, Mindoro, Samar, Leyte, 
Panay, Siargao). Fig. 4. 

Ecol. In primary or secondary forest, open 
woodland or grassland (Philippines) on well- 
drained soils of various types, 20O-80O(-1200 )m. 
Fl. Nov. -May (Borneo), July-Dec. (Java); fr. 
March-Sept. (Borneo), Dec.-April( Java, Sumatra). 

Vern. Sumatra: garinggang (P. Simalur), 
garunggang, kemutun; Java: maron(g) (Salak), 
rinjung gide (Sund.), wuluan (Central Java); 
Borneo: garonggang, girungang, sirungan (N. 
Borneo), laka-laka, s^rungan-mampat (Brunei), 
lingan, mampat (W. Kutai); Celebes: kaju arang, 
sisio pule (Malili); Philippines: many names, cf. 
Merrill 1923: 76. 

Note. This rather variable subspecies is 
represented in various islands by slightly different 
races which, although distinguishable locally, 
cannot be recognized or keyed out as taxonomic 

ssp. blancoi (Bl.), Blumca 15 (1967) 
463. — Hypericum olympicum (non I..) Blanco, 
Fl. Filip. (1837) 613. — C. blancoi Bl. Mus. Bot. 
Lugd. Bat. 2 (1852) 17; CiAONF.p. Not. Syst. 1 
(1909) 21; Mf.rr. En. Philip. 3 (1923) 75. — 
Ancistrolohus micradenius Tt RCZ. Bull. Soc. Nat. 
Mosc. 31 (1858) 382. - C. hlancoivar. apiculatum 
Mfrr. i'hilip. J. Sc. 4 (1909) Bot. 294. — C. 
punciuluium Elmlr ex Mirr. F.n. F'hilip. 3 (1923) 
76, nontrn, in sched. FIr. .3d. 

Leaves ovate or rarely elliptic-oblong to obovatc, 
]'/i-2*A times as long as broad, 4 14 by 3!^- 7 cm. 
apex rounded to acute or shortly acuminate, base 


Fig. 3. Leaf shapes, x \4. a-c. Cratoxylum suma- 
tranum (Jack) Bl. ssp. sumatranum, d. ssp. 
blancoi (Bl.) Gog., e-f. C. cochinchinense (Lour.) 

broadly to narrowly cuneate. Panicles variable in 
size, often large. Capsule 8-12 by 3-4 mm, with 
sepals over half as long. Seeds 6(-7) per loculus. 

Distr. Malesia: Philippines (Luzon, Catandu- 
anes, Panay, Guimaras, Ncgros, Bohol, Leytc, 
Samar, Palawan, Mindanao, Basilan). Fig. 4. 

Ecol. "In thickets and secondary forests at low 
altitudes" (Mfrrii i ). 

Vern. I'hilippincs: many names, cf. Merrill 
1923: 76. 

Note. Ssp. blancoi often intergrades with ssp. 
sumatranum where their respective distributions 

2. Cratoxylum rochinchlncnsfd oiiR.) Bi uMr, Mus. 
Bot. Lund. Bat. 2 (1852) 17; Mirr. i:n. Philip. 3 
(1923) 77; Hurk. Diet. (1935) 678; Cornir. (iard. 
Bull.S.S. IO(l939).34;SMvniiis,((»mm, Sarawak 
Trees (1965) 69; CJtKiUMN. Blumca 15(1967)463. 
— Hypericum chinensc (nun Oshk k nrr 1.) Kftz. 
Obscrv. 5 (1789) 27. Hypericum cochinchinense 
Lour. Fl. Coch. (1790)472; Cornfr. (iard. Bull. 

Flora Malesiana 

[ser. I, vol. 8"^ 

Fig. 4. Range of Cratoxylum sumatranum (Jack) 

Bl. ; delineated ssp. sumatranum, dotted ssp. 

blancoi (Bl.) Gog., triangles ssp. ueriifolium 

(KuRz) Gog. 

S. S. 10 (1939) 26. — Hypericum petiolatum (non 
L.) Lour. F1. Coch. (1790) 472. — Hypericum 
biflorum Lamk, Encycl. 4 (1797) 170; Hook. & 
Arn. Bot. Beechey Voy. (1833) 173. — Vismia 
cochinchinense (Lour.) Spreng. Syst. Veg. 3 
(1826) 350. — Hypericum carneum Wall. Cat. 
(1828) no 4820, nomen. — Hypericum pulchellum 
Wall. Cat. (1828) no 4821, nomen. — Hypericum 
horridum Wall. Cat. (1828) no 4822, nomen. — 
Elodea pulchella Loud. Hort. Brit. Suppl. 1 (1832) 
587, nomen. — Ancistrolobus ligustrinus Spach, 
Suit. Buff. 5 (1836) 361 ; Ann. Sc. Nat. II, 5 (1836) 
352, t. 6. — Ancistrolobus sp., Wight, 111. 1 (1840) 
111. — C. polyanthum Korth. Verb. Nat. Gesch. 
Bot. (1842) 175, t. 36; Bl. Mus. Bot. Lugd. Bat. 
2 (1852) 16; Kurz, J. As. Soc. Beng. 43, ii (1874) 
85, incl. var. genuinum et var. carneum KuRz; 
Dyer, F1. Br. Ind. 1 (1874) 257, incl. var. ligustrinum 
(Spach) Dyer et var. wightii Dyer; Kurz, F1. 
Burma 1 (1877) 84; Pierre, For. Fl. Coch. 1 
(1882) t. 50; Forb. & Hemsl. J. Linn. Soc. Bot. 
23 (1886) 72; King, J. As. Soc. Beng. 59, ii (1890) 
145; BoERL. Cat. Hort. Bog. (1901) 62, incl. var. 
macrocarpum Boerl. ; Gagnep. Not. Syst. 1 (1909) 
19; Fl. Gen. I.-C. 1 (1910) 290; Gamble, Fl. Pres. 
Madras 1 (1915) 71; Ridl. Fl. Mai. Pen. 1 (1922) 
152; Craib, Fl. Siam. En. 1 (1925) 112; Engl, in 
E. & P. Nat. Pfl. Fam. ed. 2, 21 (1925) 183; 
Hundley & Ko Ko, List Trees etc. Burma ed. 3 
(1961) 19; S.MYTHiES, Comm. Sarawak Trees 
(1965) 68. — Ancistrolobus carneus Voigt, Hort. 
Calc. Cat. (1845) 89, nomen. — Elodea chinensis 
(Retz.) Hance in Hook. Lond. J. Bot. (1848) 
472. — C. petiolatum (LouR.) Bl. Mus. Bot. Lugd. 
Bat. 2 (1852) 17. — C. ligustrinum (Spach) Bl. 
I.e. 16; Merr. En. Philip. 3 (1923) 77; Trans. Am. 
Phil. Sec. 24 (1935) 268; Burk. Diet. (1935) 678; 
Corner, Card. Bull. S. S. 10 (1939) 29, 34. — C. 
myrtifolium Bl. Mus. Bot. Lugd. Bat. 2 (1852) 
17. — C. wightii Bl. I.e. 18; Corner, Gard. Bull. 
S. S. 10 (1939) 24. — Elodea sp.. Griff. Notul. 4 

(1854) 569. — Ancistrolobus brevipes Turcz. Bull. 
Soc. Nat. Mosc. 31 (1858) 383. — C. lanceolatum 
MiQ. Fl. Ind. Bat. Suppl. {\U\) 500. — Stalagmites 
erosipetala MiQ. J. Bot. Neerl. 1 (1861) 126. — C. 
biflorum (Lamk) Turcz. Bull. Soc. Nat. Mosc. 36 
(1863) 580. — C. chinense (Retz.) Merr. Philip. 
J. Sc. 4 (1909) Bot. 292. — C. hypoleuca Elmer, 
Leafl. Philip. Bot. 5 (1913) 1787. — Fig. 3e-f. 

Tree or shrub, to c. 30 m tall, deciduous, 
glabrous; bark smooth, pale or scaly, grey-brown; 
young shoots with interpetiolar scar continuous or 
interrupted. Leaves with petiole 2-5 mm, 3-10'/^ by 
1-4 cm, elliptic to ovate-lanceolate or lanceolate, 
apex obtuse to acutely acuminate, base broadly 
cuneate to attenuate, herbaceous to chartaceous, 
nearly always glaucous beneath. Inflorescence of 
1-5-flowered axillary and usually terminal cymules; 
pedicels 1-2 mm. Flowers heterodistylous. Sepals 
5-7 by 2-5 mm. Petals dark red to pink or orange, 
5-10 by 2'/2-5 mm. Stamen fascicles 4-8 mm long, 
with stamens ± congested, c. 45-55 per fascicle; 
anther gland sometimes present. Staminodial 
fascicles (if developed) yellow, up to 3 mm long, 
flattened, oblong to obovate, cucullate. Ovary 
2-3 mm long; styles 1-3 mm. Capsule 8-12 by 
4—5 mm, c. \-l\4 times as long as sepals, ellipsoid- 
cylindric, with columella basal. Seeds (5-)6-8 per 
loculus, 6-8 by 2-3 mm, oblanceolate to elliptic or 

Distr. Burma, China (south of 23°N), Hong- 
kong, Hainan, Indo-China, Thailand, in Malesia: 
Sumatra, Malaya, Borneo, Philippines (Palawan, 
Culion). Fig. 5. 

Ecol. In primary or secondary forest, open 
woodland, grassland and river banks on usually 
well-drained soils of various types, 0-500 m (to 
1200 m in China). throughout the year. 

Vern. Sumatra: mulu (Lampongs), kaju lulus, 
lelulus (Palembang), tnengidjang (Barika); Malaya: 
derum selunchor, selunchus (Perak), kaju arang, 
kemuntong, pelawan (Kelantan), belawan padang. 

Fig. 5. Range of Cratoxylum cochinchinense 
(Lour.) Bl., coinciding with the range of sect. 


Hypericaceae (Robson) 

k^lochus (Pahang), beluchus (Negri Sernbilan), 
serapat; Borneo: mampat, maradjalang, inulun 
(W. Borneo), baditk-baduk, geronggang, mogizon, 
pela^\an-pelawan (Brunei), selangan-biabas, taik- 
aA:a/7(Selangan Is.). 

Note. The variation in C. cochinchinense 
ranges between the following extremes : 

(1) 'C. polyanthum var. wight ii' (Dyer). Shrub or 
small tree; bark rough, grey-brown; inter- 
petiolar scar usually interrupted. Leaves 
sometimes subopposite; lamina apex usually 
obtuse, base broadly to narrowly cuneate, 
seldom separating from midrib. Stamen 
fascicle stalks slender, over M of the total 
fascicle length (in flowers with staminode 
fascicles present). 

(2) 'C. polyanthum var. ligustrinum' (Dyer). 
Becoming large tree; bark smooth, pale; 
interpetiolar scar continuous. Leaves opposite; 
lamina apex acuminate, base attenuate, often 
slightly separating from midrib to produce 
minute appendages. Stamen fascicle stalks 
often compressed, short. 

3. Cratoxylum formosum (Jack) Dyer, F1. Br. Ind. 
1 (1874) 258; Kurz, FI. Burma 1 (1877) 84; 
Pierre, For. Fl. Coch. I (1882) t. 51 ; King. J. As. 
Soc. Beng. 59, ii (1890) 147; K. & V. Bijdr. 5 
(1900) 137; Gagnep. Not. Syst. 1 (1909) 288; Fl. 
Gen. L-C. 1 (1909) 288; Back. Schoolfl. Java 
(1911) 88; RiDL. FI. Mai. Pen. 1 (1922) 153, f. 16; 
Craib, Fl. Siam. En. 1 (1925) 111; Engl, in E. & 
P. Nat. Pfl. Fam. ed. 2, 21 (1925) 184; Hend. Gard. 
Bull. S. S. 4 (1928) 222; Corner, Gard. Bull. S. S. 
10 (1939) 34; Wavside Trees Mai. (1940) 325, t. 69; 
Back. & Bakh. / FI. Java 1 (1963) 363; Meijer, 
Bot. NewsbuII. For. Dept. Sandakan 7 (1967) 
64; GoGELEiN, Blumea 15 (1967) 467. — Elodea 
formosa Jack, Mai. Misc. 2 (1 822) 24. — Hypericum 
biflorum Uion Lamk) Choisy in DC. Prod. 1 (1824) 
546. — Tridesmis ochnoides Spach, Suit. Buff. 5 
(1836) 359; Ann. Sc. Nat. H, 5 (1836) 351, t. 4A; 
Bl. Mus. Bot. Lugd. Bat. 2 (1852) 18. — Tridesmis 
jackii Spach, Ann. Sc. Nat. II. 5 (1836) 352; 
Turcz. Bull. Soc. Nat. Mosc. 31 (1858) 384. — 
Hypericum aegiptium (non H. aegypticum L.) 
Blanco, Fl. Filip. (1837) 615. — Tridesmis 
formosa (Jack) Korth. Verb. Nat. Gesch. Bot. 
(1843) 197, t. 37. — C. penladelphum Turcz. Bull. 
Soc. Nat. Mosc. 36 (1863) 580; K. & V. Bijdr. 5 
(1900) 139. — Fig. 6a-l. 

Tree to 35 m tall, deciduous, glabrous, or 
partly pubescent; trunk spiny at base, bark grey, 
coarsely fissured, scaly; young shoots glabrous or 
pubescent, with interpetiolar scar interrupted. 
Leaves with petiole 5-15 mm, 3^-14 by 1-7 cm, 
elliptic or sometimes obovatc to oblong or 
lanceolate or ovate, apex acute or shortly acumin- 
ate to rounded, broadly cuneate to rounded, 
herbaceous to chartaccous. sometimes glaucous 
beneath. Inflorescence of 1-6-flowcrcd cymulcs in 
lower axils of current shoots or. if shoot fails to 
develop, apparently axillary on older stems; 
pedicels 3-!0(-15)mm. Flowers hcterodistylous. 
Sepals 4-7 by 2-4 mm. Petals white or pink to red 
or (very rarely) purplish. 7-17 by 3 -7 mm; 
nectary scale 2-4 mm. rounded, entire or sub- 
entire. Stamen fascicles 5-l4mm. with stamens 
relatively lax, 20 per fascicle; anther gland present, 

purple, or absent. Staminodial fascicles (if well 
developed) orange-red, trigonous, linguiform, 
attenuate to truncate, not cucullate. Ovary 2-A14 
mm long; styles 2-8 mm. Capsule 10-16 by 4-6 mm, 
three times as long as sepals, ellipsoid, with colum- 
ella — half as long as capsule. Seeds (7-) 12-1 7 per 
loculus, 6-7l2 by 2-4 mm, oblanceolate. 

Distr. Hainan, South Vietnam, Cambodia, S. 
Thailand, S. Andaman Is., in Malesia: Sumatra, 
Malaya, Banka, Java, Borneo, Celebes (incl. P. 
Buton), Philippines (Luzon, Polillo, Mindoro. 
Sibuyan, Guimaras, Negros, Busuanga, Culion, 
Palawan, Mindanao). Fig. 7. 

Ecol. In primary or old secondary forest, hill 
slopes, river margins or swamps, on sand or clay 
soils. 0-600(-1200) m. Fl. March-July (Borneo) or 
later (Sumatra, Banka) or earlier (Malaya). 

Vern. Sumatra: kebutul, kemutul (Lampongs), 
kembutul, tembutun (Palembang), mampat (Palera- 
bang, Riouw), ampet, mampet, temutun (Banka); 
Malaya: derom (Kedah, K. Lumpur), betina, 
gerunggang, mumpat (Perak), kemuntong, mampat 
(Kelantan), mampat (K. Lumpur), drum (P. 
Penang); Java: rembang putun; Borneo: butun, 
kasat baku, mulun (S. & E. Borneo), pelawan 
(Brunei), biabas tahun, geronggang, kalambunan, 
mcngkutan, obah bukit, serungan (N. Borneo); 
Philippines: mango-gong, marangguub, salingagon 

Note. C formosum comprises two subspecies 
which scarcely overlap in distribution. Only ssp. 
formosum occurs in the Flora area. It differs from 
ssp. prunifiorum (Kurz) Gogelein (S. China and 
Burma to S. Thailand) in being completely 
glabrous with leaves elliptic to oblong or rarely 
lanceolate (not oblong to lanceolate) and the 
anther connective eglandular (not glandular). 

4. Cratoxvlum maingavi Dyer. Fl. Br. Ind. I 
(1874) 258; King, J. As. Soc. Beng. 59, ii (1890) 
147; Ann. R. Bot. Gard. Calc. 5 (1896) 138, t. 165; 
Gagnep. Not. Syst. 1 (1909) 19; Ridl. Fl. Mai. 
Pen. 1 (1922) 154; Burk. Diet. (1935) 679; 
Corner. Gard. Bull. S. S. 10 (1939) 35; Wayside 
Trees Mai. (1940) 327; Gogelein. Blumea 15 
(1967) 470. — C. harmandii Pierre. For. Fl. Coch, 
1 (1882) t. 53; Gagnep. Not. Syst. 1 (1909) 18; 
Fl. G6n. I.-C. 1 (1909) 288. — C. formosum var. 
thorclii Gagnep. Not. Syst. 1 (1909) 19; Fl. G^n. 
I.-C. 1 (1909) 289. — C. cochinchinense var. 
calcareum Ridl. Kew Bull. (1938) 115. — C. 
acuminatum Merr. Pap. Mich. Ac. Sc. 23 (1938) 
185. — C. subglaucum Merr. I.e. 186. — C. 
parviflorum Merr. J. Arn. Arb. 19 (1938) 56. — C. 
thorclii Pierre ex Gagnep. Fl. G*n. I.-C. Suppl. I 
(1943)252. — FIr. 6m-o. 

Shrub or small tree, to 20 (rarely 35) m tall, 
deciduous or partly deciduous, glabrous; bark 
narrowly fissured with small scales, dark greyish- 
brown; young shoots with interpetiolar scar 
interrupted. Leaves with petiole 3 7 mm. 2-9 by 
\-A\i cm. elliptic to oblong, apex shortly acumin- 
ate to rounded, base cuneate to subattcnuate. 
chartaccous. not glaucous beneath. Inflorescence 
of 1-4-flowcrcd cymulcs in lower (or sometimes 
also upper) axils of current shoots or. if shoot fails 
to develop, apparently axillary o.i older stems; 
pedicels 2-6(-8) mm. Flowers hcterodistylous. 
Sepals 2*A-5 by 1-3 mm. Petals white or pink. 


Flora Malesiana 

[ser. I, vol. 8^ 

Fig. 6. Sect. Tridesmos. — Cratoxylum formosum (Jack) Dyer ssp. formosum a Fertile twig apex, 
XM b. leaf, xVz, c. node, showing open interpetiolar scar, enlarged, d. short-styled flower, x 2, e. petal, 
from inside, x4, f. base of same, showing appendage, x8, g. gemtals and staminodial fascicles ot 
short-styled flower, x6, h. pistil of long-styled flower, x6, i. staminodial fascicle x6, 1. seed, a nne 
vein running submarginally through wing towards top of seed proper, x4. -C. formosum ssp. pruni- 
florum (KuRz) GoG. j. Staminodial fascicle, x6, k. anther, showmg connective glands, x2U. — c. 
maingayi Dyer. m-n. Seeds, x4, o. cross-section of capsule, schematic («' ^-^' ' J^Y^^.'^^ rJ^^^ 
Braskamp s.n., dd. 12-11-1927, d-h Magnen, Gourgand, Chatillon dd. 20-10-1909, y-A: Henry 
10687, m-n SF 36465, o Rahmat Si Boeea 7907). 


Hypericaceae (Robson) 


Fig. 7. Range of sect. Tridesmos; delineated 

Cratoxylum formosum (Jack) Dyer ssp.formosum, 

triangles ditto ssp. pruniflorum (KuRZ) Gog., 

dotted C. maingayi Dyer. 

6-12 by 2-5 mm; nectary scale 2-3 mm, truncate, 
undulate-denticulate. Stamen fascicles (5?-)7-9 
mm, with stamens relatively lax, nearly always <20 
per fascicle; anther gland present. Staminodial 
fascicles trigonous (?), linguiform, attenuate, not 
cucullate. Ovary 2-3 mm long; styles 1-5 mm. 
Capsule (6-J9-15 by 3-4(-5;^) mm, c. 2'A times as 
long as sepals, fusiform, with columella ±?^ as 
long as capsule. Seeds 5-6 per loculus, 6-6li by 
VA-2 mm, oblong to oblanceolate-oblong. 

Distr. Burma, Cambodia, Central Vietnam, in 
Malesia: Sumatra (Payakumbu, Asahan, Riouw, 
Palembang), Malaya (Kedah, P. Penang, Johore, 
Singapore), Borneo (Sarawak, only near Kuching). 
Fig. 7. 

Ecol. In lowland forest and on limestone hills, 
(6O-)30O-8(X) m. Fl. March-May (Malaya), Oct. 

Vcrn. Sumatra: sSmapat (West), p^matang 
(Palembang), kaju bonbon, kaju si haras (Asahan); 
Malaya: dirom bukit (Kedah), mampat (Johore); 
Borneo: girunggang (Sarawak). 

Note. The discontinuous populations of C. 
maingayi tend to be morphologically distinguish- 
able. They all differ from C. formosum, however, in 
the form and number of the seeds, the number of 
stamens per fascicle, the size of the leaves and the 
capsule shape. 

5. Cratoxylum arborcscens (Vami.) Blume, Mus. 
Bot. Lugd. Bat. 2 (1852) 17; Kurz, J. As. Soc. 
Beng. 43, ii (1874) 83; Dyfr, Fl. Br. Ind. 1 (1874) 
258; KuRZ, Fl. Burma 1 (1877) 85; Kinc;. J. As. 
Soc. Bcng. 59, ii (1890) 146, incl. var. miquclii 
Kino; Ridl. Fl. Mai. Pen. 1 (1922) 153; Fm,!,. in 
E. & P. Nat. F>n. Jam. cd. 2. 21 (1925) 184; 
Foxw. Mai. For. Rcc. 3 (1927) 138. tt.; Hinij. 
Card. Bull. S. S. 4 (1928) 223; Bdrk. Diet. (1935) 
678; Corner, Gard. Bull. S. S. 10 (1939) 22. 31, 
34; Wayside Trees Mai. (1940) 325; Munoi i y &. 
Ko Ko. List Trees etc. Burma cd. 3 (1961) 19; 

Meijer, Bot. Newsbull. For. Dept. Sandakan 7 
(1967) 64; Gogelein, Blumea 15 (1967) 471. — 
Hypericum arborescens Vahl, Symb. 2 (1791) 86, 
t. 43. — Vismia arborescens (Vahl) Choisy, Prod. 
Monogr. Hyper. (1821) 36; in DC. Prod. 1 (1824) 
543. — Hypericum coccineum Wall. Cat. (1828) 
no 4823, nomen. — Ancistrolobus glaucescens 
TuRcz. Bull. Soc. Nat. Mosc. 31 (1858) 383. — C. 
cuneatum MiQ. Fl. Ind. Bat. 1, 2 (1859) 517; 
Gagnep. Not. Syst. 1 (1909) 21. — Fig. 8. 

Tree, up to 45 m tall, evergreen, glabrous; bark 
scaly, reddish; young shoots with interpetiolar 
scar continuous. Leaves with petiole 5-10 mm, 
5-16 by 2-6 cm, obovate-oblong to obovate- 
oblanceolate or elliptic {1-A times longer than 
broad), apex acute to cuspidate, base cuneate to 
attenuate, usually coriaceous, not or scarcely 
glaucous or papillose beneath. Inflorescence a 
many-flowered terminal pyramidal panicle; pedi- 
cels lH-3 mm. Flowers homostylous. Sepals VA-() 
by 2-A\i mm. Petals deep red or very rarely 
orange or white, 4\i-l by 2! 2-5 mm; nectary scale 
up to 1 mm, often deeply laciniate. Stamen 
fascicles 4-5 mm, with stamens relatively congested, 
30-40 per fascicle; anther gland absent? Staminod- 
ial fascicles (if well developed) greenish yellow, 
flattened, obovate, cucullate. Ovary 1 ! 2-2 mm 
long; styles lH-3 mm. Capsule 7-9 by c. 4 mm, c. 
\Vi times as long as sepals, cylindric, with columella 
half as long as capsule. Seeds 10-18 per loculus, c. 
5 by 0.8 mm, narrowly oblong. 

Distr. Burma (Tenasserim), in Malesia: 
Sumatra, Malaya (except the NE. part), Borneo. 
Fig. 10. 

Ecol. Primary forest, dipterocarp forest, open 
forest, peat- and swamp-forest and fresh-water 
swamps on sand or sandy loam, 0-900 m or higher 
(to 1800 m on Mt Kinabalu). 

Vern. Sumatra: geronggang, gronggang; lede 
(Atjeh), kalat (Riouw), ampet, idat (Banka), 
kaju dori (Billiton), kaju si pare-pare (Asahan); 
Borneo: gerongan, gdrunggang, serungan (N. 
Borneo, Brunei). 

Note. ''Var. miquelii' is an extreme form with 
slender petioles and thin lanceolate-elliptic leaf- 
laminae, which is connected to the typical form by 
many intermediates. 

6. Cratoxylum glaucum Korth. Verb. Nat. Gesch. 
Bot. (1842) 176; Bl. Mus. Bot. Lugd. Bat. 2 (1852) 
17; MiQ. Fl. Ind. Bat. 1, 2 (1859) 517; Hall./. 
Ann. Jard. Bot. Btzg 14 (1897) 26; Gagnf.p. Not. 
Syst. 1 (1909) 21 ;Go(iELEiN. Blumea 15(1967)473. 
— C. microphvllum MiQ. I"l. Ind. Bat. Suppl. 1 
(1861) 500; Ridl. Fl. Mai. Pen. 1 (1922) 153; 
Smythies, Comm. Sarawak Trees (1965) 69. — C. 
polvstachvum Ti'RCZ. Bull. Soc. Nat. Mosc. 36 
(1863) 579. —C />rocm/w DiHLS, Bot. Jahrb. 
(1926) 31 1. — Cratoxylum A Corner, Gard. Bull. 
S. S. 10(1939)32. — Fig. 9. 

Tree, up to 25 m tall, evergreen, glabrous; bark 
scaly (?); young shoots with interpetiolar scar 
continuous. I. caves with petiole 2-3 mm. 2-10 by 
I 5 cm. elliptic (mostly less than twice as long as 
wide), apex acute or obtuse with often blackish 
mucro to rounded, base broadly to narrowly 
cuncalc, coriaceous, densely greyish to orangc- 
ycllow papillose-glaucous beneath. Inflorescence u 
few- to many-flowered terminal, cylindric to 


Flora Malesiana 

[ser. I, vol. 8^ 

Fig. 8. Cratoxylum arborescens (Vahl) Bl. a. Flowering twig, XM, b. venation of leaf, c. upper node 
of a, enlarged, showing continuous petiolar scar, d. flower, x 2, e. petal from inside, with appendage, 
x4, f. petal appendage, x8, g. genitals with small staminodial fascicles, x6, h. staminal phalange 
from outside, i. large, well-developed staminodial fascicle, x6, j. young anther, filament with swollen 
cells, x20, k. young fruit, x2, 1. dehisced capsule, x2, m. seed, winged all around, a vein running 
from margin with a loop through wing towards the centrally situated seed proper, x 4 {a-h, j SAN 

23821, / Hamid 6383, k-m Griffith 839). 


Hypericaceae (Robson) 


Fig. 9. Cratoxylum glaucum Korto. (Photogr. Father A. Elsener. Kalimantcn. Sanggau, 1961). 


Flora Malesiana 

[ser. I, vol. 8^ 

pyramidal panicle; pedicels 1-3 mm. Flowers 
homostyious. Sepals 3-6 by 2!^-5 mm. Petals deep 
red to crimson, 6-8 by 4-5 mm; nectary scale up 
to 1 mm (usually shorter), shallowly denticulate. 
Stamen fascicles c. 5 mm, with stamens relatively 
congested, 30-40 in each fascicle; anther gland 
absent. Staminodial fascicles (if well developed) 
VAmm, flattened, oblong, cucullate. Ovary 114- 
2]A mm long; styles 2!/^-3 mm. Capsule iVi-lO by 
3-4 mm, 2'/^-3 times as long as sepals, cylindric, 
with columella ±[4 as long as capsule. Seeds 
(4-) 6-8 per loculus, 5-5'A by 1 mm, narrowly 

Distr. Malesia: Malaya (E. Johore: Mt Ophir, 
G. Arong near Mersing), E. Sumatran Is. (Lingga, 
Banka, Billiton, Karimata), Natunals. (Bunguran), 
Borneo (west: Sarawak, Brunei; central: Sanggau, 
Kenepai; south-east, also P. Lampei). Fig. 10. 

Ecol. Mainly on podsolised sands and kerangas 
(heath woodland, raised beaches, etc.), also in 
swampy or peaty areas (fresh-water swamps, peat 
swamp, peat forest), 0-100 m, rarely higher (600 m 
on Mt Maros, Banka, 1000 m and above on Mt 
Ophir in Johore). 

Vern. E. Sumatran Is.: edat, idatjdet (Banka), 
grunggang (Billiton); Borneo: edat, kaju longgang 
(Pontianak), geronggang, gerimgan (Samarinda, 
Sampit, Sarawak), serongan (Brunei). 

Fig. 10. Range of sect. Isopterygiiim; delineated 
Cratoxylum arborescens (Vahl) Bl., dotted C. 
glaucum KoRTH. 


LiNNE, Gen. PI. ed. 5 (1754) 341 ; Sp. PI. (1753) 783; R. Keller in E. & P. Nat. 
Pfl. Fam. ed. 2, 21 (1925) 175, pro parte excl. sect. IVElodea; Y. Kimura in Nakai 
& Honda, Nova Fl. Jap. 10 (1951) 108; Robson, Blumea 20 (1972) 251-274.— 
Ascyrum L. Gen. PI. ed. 5 (1754) 342; Sp. PI. (1753) 787; Engl, in E. & P. Nat. 
Pfl. Fam. ed. 2, 21 (1925) \lA.—Sarothra L. Gen. PI. ed. 5 (1754) 133; Sp. PI. 
(1753) 272; Y. Kimura in Nakai & Honda, Nova Fl. Jap. 10 (1951) 229. — 
Brathys MuTis ex L./. Suppl. PI. (1781) 43; Spach, Hist. Nat. Veg. Phan. 5 (1836) 
446; Ann. Sc. Nat. II, Bot. 5 (1836) 356; Miq. Fl. Ind. Bat. 1, 2 (1859) 513. — 
Norysca Spach, Hist. Nat. Veg. Phan. 5 (1836) 426; Ann. Sc. Nat. II, Bot. 5 
(1836) 363; Y. Kimura in Nakai & Honda, Nova Fl. Jap. 10 (1951) 96. — Taka- 
sagoya Y. Kimura, Bot. Mag. Tokyo 50 (1936) 498; in Nakai & Honda, Nova Fl. 
Jap. 10(1951)85. 

Small trees, shrubs or perennial or annual herbs, glabrous (in Mai.). Bark 
not exuding resinous sap. Branchlets terete or 2-4-lined or -angled. Leaves op- 
posite (rarely whorled), entire (rarely gland-fringed in extra-Mai. spp.), sessile or 
shortly petioled, with translucent ('pale') glands containing essential oils and 
sometimes black or red glands containing hypericin or pseudohypericin. Inflores- 
cences terminal, cymose, dichasial or monochasial. Flowers (sometimes except the 
gynoecium) 5(-4)-merous, homostyious (in Mai.). Sepals quincuncial or rarely 
decussate, coriaceous to chartaceous, persistent (in Mai.), glandular like the 
leaves. Petals yellow, often tinged red, glandular like the leaves, without nectarif- 
erous appendage (in Mai.), glabrous, caducous or persistent. Stamen fascicles 
epipetalous, free or variously united (2 + 1 + 1 + 1 or 2 + 2 + 1 or (5) or (4)) 
and then with double ones episepalous, glabrous, caducous or persistent, each with 
1-C.60 stamens; filaments yellow, slender, usually united only towards the base or 
apparently free ; anthers yellow or reddish, shortly oblong, dorsifixed or apparently 


Hypericaceae (Robson) 


Fig. 1 1 . Distribution of Hypericum in Malesia. Figures above the hyphen indicate the number of 
endemic species in each island or island group, that below the hyphen the total number of species. 

basifixed {sect. Takasagoyd), with amber or black gland terminating the con- 
nective. Staminodial fascicles absent (or very rarely 3, alternating with 2 + 2 + 1 
stamen fascicles in extra-Mai. spp.). Ovary 5-3-celled or 1 -celled with 5-3(-2) 
parietal placentas; styles 5-3(-2), free or ± united, ± slender; stigma small, flat 
or ± capitate. Ovules <>o-2 on each placenta, horizontal. Fruit a septicidal 5-3(-2)- 
valved ± coriaceous capsule (rarely tardily dehiscent or ± baccate in extra-Mai. 
spp.), with + prominent vittae in the valves. Seeds oo-l on each placenta, curvcd- 
cylindric to ellipsoid, sometimes carinate with a membranous wing; testa variously 
sculptured; embryo cylindric, straight or curved, with cotyledons usually shorter 
than the hypocotyl. 

Distr. About 400 spp. throughout most of the tropical and temperate zones, but absent from some 
lowland tropical areas {e.g. the Amazon basin) and rare in Australasia (2 native species, here .spp. 14 
and 15), in Malesia 13 spp. Fig. 1 1. 

Ecol. In forest margins, grasslands, marshes or among rocks; rare in general in tropical lowland, but 
occurring in Malesia from sea-level in Sumatra to 3400 m in New Guinea. 

Notes. Hypericum, as recognised here, is identical with Engler's Hypcriccae except for Hypericum 
sect. Elodea (Juss.) CfioiSY, which, as the genus Triadenum Rafin., belongs in the Cralu.xylcae and 
appears to be a herbaceous derivative of Cratoxylum. 

The geographical relationships of the Malcsian species arc interesting. Sect. Ascyreia (Sumatra to SW. 
Celebes) also occurs in south continental Asia, Taiwan, Ceylon, Turkey, and Socotra. Sect. Takasagoya 
(Philippines) is otherwise confined to Taiwan. Sect. Hypericum (Philippines, Sabah, Sumatra) is pan- 
boreal in distribution; but the Philippine species is found elsewhere only in Taiwan anti Japan, whereas 
the species of .Sabah and Sumatra is otherwise eastern Himalayan in distribution. One of the two species 
oi sect, liralhys, which also occurs from Japan to Ceylon and in Australia, New Zealand and Hawaii, is 
closely related to a species of southeastern U.S.A., whereas the other has a mainly southern distribution 
(Australia, Now Zailand, New Caledonia, New Cjuinea) with outliers in South and Hast Asia (eastern 
Himalayas, Vietnam, Taiwan) and allinities with western South America. The allinilios of the remaining 
section, sect. Humifusoidcum, lie in quite a dilferent direction, namely with Airica. I ho New (iuinca 
species form a closely related group which appears to Ik derived from sect. ( iiiupyli)spt>rus of Africa, 
Madagascar and the Mascarene Islands. In aiUiilion, however, there are three species of sect. Ilumi- 
fusnidcum in Africa and Madagascar (//. luitalcnse W(K)I) A. I-.vans, //. wilmsii U. KiiiiK and H. 
pcplidifolium A. Ricii.), which are apparently derivatives, in turn, of (his New Ciuinea group. The 
relationships therefore are Africa -► New Guinea -► Africa. This New Guinea group also appears to 

16 Flora Malesiana [ser. I, vol. 8^ 

have given rise to two other species, H. pulogense Merr. (Philippines) and H. beccarii N. Robson (Java, 

Hybrids of Hypericum species have been found from time to time in nature, but only between rather 
closely related species. Artificial hybrids are not always easy to produce, but some 'wide' crosses have 
been successful. The resultant plants, however, are usually weak and have always proved sterile. Some 
artificial crosses between closely related species, however, have thrived, and one or two are well-known 
garden plants. In general, apart from the hybrids involving H. perforatum L., which is largely pseudo- 
gamous and produces n and 2n gametes, crosses are likely to be successful only between species with 
the same chromosome number. 


1. Petals and stamens caducous. Stamen fascicles, styles and ovary loculi 5. Broad-leaved shrubs, 

without black glands. 
2. Styles free or partially united. Anthers dorsifixed. Inflorescence up to 3 nodes long. Sect. Ascyreia 
3. Styles free. Leaves laxly or scarcely reticulate beneath. 

4. Sepals acute to subacute. Petals (20-)25-45 mm long. Branches erect or arching, not frondose. 

1. H. leschenaultii 

4. Sepals rounded. Petals 9-18 mm long. Branches arching, sometimes frondose. . 2. H. uralum 

3. Styles united almost to the apex. Leaves densely reticulate beneath. . . . 3. H. monogynum 

2. Styles completely united. Anthers apparently basifixed. Inflorescence up to 14 nodes long. Sect. 

Takasagoya (Y. Kimura) N. Robson 4. H. geminifljorum 

1. Petals and stamens persistent. Stamen fascicles 5-3 (/.e. 2 + 2 + 1) or stamens apparently afasciculate. 
Styles 3-5. Ovary loculi 1-4. ± Ericoid shrubs or shrublets or herbs, often with black glands on 
various parts of the plant. 
5. Capsule longitudinally vittate (scarcely so in H. beccarii ssp. steenisii). Placentation axile to parietal. 
Black glands usually present. Petals pale yellow (flavus) to golden yellow (aureus). Shrubs or herbs. 
6. Plant an erect or ± straggling shrub or woody herb. Styles 3-5, equalling or shorter than the 
ovary. Placentation axile to parietal. Sect. Humifusoideum R. Keller. 
7. Flowers solitary, not in dichasial cymes nor with shoots in the axils of the uppermost leaf pair. 
8. Leaves cuneate at the base, linear to elliptic or ovate-lanceolate, rarely broader, with laminar 
pale glands almost always linear towards the leaf base. 
9. Leaves spreading or ascending, 8-12 mm long, flat. Styles 2H-3 mm long. . . 5. H. sewense 
9. Leaves imbricate, ± appressed, 2-9 mm long, flat or incurved. Styles 114-214 mm long. 
10. Leaves flat, ovate-lanceolate to narrowly elliptic-oblong, with laminar glands mostly inter- 
rupted to punctate. Placentation axile 6. H. macgregorii 

10. Leaves incurved, narrowly elliptic or lanceolate-elliptic to linear with laminar glands mostly 

linear. Placentation parietal 7. H. saruwagedicum 

8. Leaves rounded to cordate at the base, narrowly ovate or ovate- triangular or elliptic to subcircular, 
with laminar pale glands linear towards the leaf base only or wholly punctate. 

9. H. papuanum 
7. Flowers in regular or irregular dichasial cymes or, if solitary, then with shoots in the axils of 

the uppermost leaf pair. 
11. Flowers always solitary with shoots in the axils of the uppermost leaf pair; leaves with laminar 

pale glands linear, often with parallel rows of dots. Styles 3 8. H. bifurcatum 

11. Flowers in regular or irregularly dichasial cymes; leaves with laminar pale glands linear or 

punctate, without parallel rows of dots. Styles 3-5 9. H. papuanum 

6. Plant a perennial or annual herb or, if suffrutescent, then styles 1.4-1.7 times as long as the ovary. 
Styles 3 (4). Placentation axile. 
12. Stem intemodes 2-4(-6)-lined, at least when young. Stamen fascicles 3-5 or stamens not obviously 
in fascicles. Flowers solitary or with shoots in uppermost leaf axils. Suffrutescent or wiry 
herbs. Sect. Humifusoideum R. Keller. 
13. Plant a suffrutescent herb with erect or ascending stems. Leaves sessile. Flowers 2-3 cm 0. 

10. H. pulogense 

13. Plant a perennial herb with diffuse wiry branches. Leaves petioled. Flowers ^i-1 cm 0. 

11. H. beccarii 
12. Stem internodes terete. Stamen fascicles 3. Flowers in dichasial or monochasial cymes. Tufted 
or straggling perennial or annual herbs. Sect. Hypericum. 

14. Sepal apex rounded. Petiole absent or up to 0.7 mm long. Perennial. 12. H. pseudopetiolatum 
14. Sepal apex acute. Petiole 1-3 mm long. Annual 13. H. petiolulatum 

5. Capsule smooth, not vittate. Placentation parietal. Black glands absent. Petals usually apricot 

yellow (armeniacus) to orange (aurantiacus), sometimes red-tinged; small herbs with quadrangular 

stems. Sect. Brathys (MuTis ex L. /.) Choisy. 

15. Petals c. 1.3 times as long as sepals. Sepals lanceolate to narrowly elliptic, acute or subacute. 

Stamens 30-50. Leaves mostly ovate-lanceolate to linear-lanceolate or narrowly oblong. Erect 

perennial or annual 14. H. gramineum 

15. Petals c. 0.85-1 (-1.3) times as long as sepals. Sepals narrowly oblong to elliptic or obovate, acute 
to rounded. Stamens 5-30. Leaves very variable in shape. Erect to prostrate annual (always?) 

15. H. japonicum 


Hypericaceae (Robson) 


Fig. 12. Hypericum leschenaultii Choisy, West Java, Mt Pangrango, summit, 3000 m (Photogr. Docters 

Van Leeuwen). 

1. Hypericum leschenaultii Ciioisy in DC. Prod. 1 
(1824) 545; in Zoll. Syst. Vcr/. (1854) 151; V\. 
Jav. (1 858) 6; Guili.emin in Dclcsscrt, Ic. Scl. PI. 
3(1817) 17. t. 27; Back. Schdolfl. Java (1911) 87; 
Kh.i.ik in i:. & P. Nat. Pfl. lam. cd. 2. 21 (1925) 
176; H'KiiK. C'andolica 2 (1925) 435. incl. var. 
Ivpicum H(k:mr., /. clalior Hocmr. el var. coria- 
(I'um {\U.) Hcx:iiR.; Siaim. Hot. Mag. (1926) t. 
9160; FJ<Krr. v. Leki/win, Irop. Natuiir 16 (1927) 
115 f. 23. 24; Mirk. Contr. Arn. Arb. 8 (1934) 
107; Sri IN, Mull. Jard. Hot. Hl/t? III. 13 (1934) 219; 
lUfK. & »AKM./. 11. Java 1 (1963) 382; Rohson. 
J. R, Hon. Soc. 95 (1970) 489; Shin. Mt M. Java 
(1972) t 23: 6. - - H. jdvanicum Hi.. Hijdr. I (1825) 
141; Hassk. Cat. Hort. Bog. (1844) 213; Mor. 


Syst. Verz. (1846) 25; Choisy in Zoll. Syst. Verz. 
(1854) 151. -- //. cnriaccttm Bi,. Bijdr. 1 (1825) 
142. -- //. trijloniin Bi.. I.e. 142, 143. incl. lar. 
anfuistalum Bi..; IIassk. Cat. Hort. Bog. (1844) 
213; Mor. Syst. Verz. (1846) 25; (Burhiikii), The 
Ciardcn 23 (1883) 158, t. 376; KuiiUR in li. & P. 
Nat. I'll. l-am. cd. 2. 21 (1925) 176, f. 73. — //. 
m-rvosum (iioisY in Zoll. Syst. Verz. (1854) 152, 
non I). Don. 1825. notn. Uln'if. Nuriwra 

lc\rhtii(iiiliii (i'Moisy) Bi . Mus. Bol. I.iigil. Bat. 2 
(1856) 24; Mk;. II. Ind. 1. 2 (1859) 515; V. 
KiMiKA m Nakai & Honda. 11. Jap. 10 
(|'>51)98. • Norvscd jamnica {\\\ .) Bi . Mus. Bot. 
I.ugd. Hal. 2 (1856) 24; Mio. II. Ind. Bat. 1, 2 
(1859) 514. Norysca curiacea (Bi.) Bi.. Mus. 


Flora Malesiana 

[ser. I, vol. 8^ 

Bot. Lugd. Bat 2 (1856) 24; MiQ. Fl. Ind. Bat. 1, 2 
(1859) 515. — H. nervosa (Choisy) Miq. I.e. — H. 
hookerianum var. leschenaultii (Choisy) Dyer, Fl. 
Br. Ind. 1 (1874) 254, quoad typum; Ridl. J. Fed. 
Mai. Stat. Mus. 8, 4 (1917) 17. — H. patulum 
(ssp.) 6 leschenaultii (Choisy) O.K. et (ssp.) /? 
variabile O.K. Rev. Gen. PI. 1 (1891) 60. — H. 
hookerianum {non W. & A.) Buysman, Flora 107 
(1914) 358. — Norysca hookeriana var. leschen- 
aultii (Choisy) Y. KiMURA in Hara, Fl. E. Himal. 
(1966) 210, quoad typum. — Fig. 12. 

Shrub or treelet, M-lM m, branchlets 4-lined and 
flattened when young, becoming terete. Leaves 
subsessile or up to 7 mm petiolate, 2\i-2> by 1-3M 
cm, lanceolate to narrowly ovate or rarely ovate, 
apex acute to rounded-apiculate, base broadly 
cuneate to rounded, margin often reflexed, glaucous 
beneath; 4 main lateral veins, with lax reticulate 
venation; glands all pale, shortly striate and 
punctate. Inflorescence l-3(-10, rarely up to 18)- 
flowered, terminal (1-2 nodes), corymbose. 
Flowers (3'/^-)4-7 cm 0, flat or slightly concave; 
buds ± broadly ovoid, acute or subacute. Sepals 
1-10 by 2-8 mm, free, imbricate, narrowly oblong 
or narrowly elliptic or rarely ovate to oblanceolate, 
very acute to subacute, entire, midrib sometimes 
visible below and sometimes incrassate above; 
spreading or recurved in flower and fruit; glands 
all pale, linear. Petals deep golden yellow, (2-)2;-^- 
4M by lVf-3 cm, broadly obovate to obovate- 
circular, entire, deciduous; apiculus ± distinct, 
acute to obtuse; glands all pale, linear. Stamen 
fascicles 5, 9-12 mm long, c. Vs as long as petals, 
each with c. 80 stamens, caducous; anthers 
yellow, gland amber. Ovary 6-9 mm, ovoid or — 
narrowly ovoid-conic to narrowly ellipsoid, 
acuminate; styles 5, 3'/i-7 mm, about half as long 
as ovary, erect or suberect, outcurved near apex, 
free; stigma small; placentas 5, axile, with central 
lacuna. Capsule 1-2 cm, narrowly ellipsoid or 
narrowly ovoid-conic. Seeds dark orange-brown to 
reddish-brown, 0.8-1 mm, cylindric, sometimes 
curved, scarcely carinate, shallowly and minutely 
linear-foveolate to reticulate. 

Distr. Malesia: Sumatra, Java, Lesser Sunda 
Is. (Bali, Lombok, Flores), SW. Celebes (Mts 
Bonthain & Latimodjong). Fig. 13. 

Fig. 13. Range of Hypericum leschenaultii Choisy. 

Ecol. Grassy slopes, thickets, open woodland, 
1500-3300 m. 

Note. The form on Mt Kerintji (Sumatra) has 
more prominent main lateral leaf veins, larger 
flowers and broader sepals than the remainder of 
the species. In this it comes closest to the nearest 
relative of H. leschenaultii, a hitherto undescribed 
species from Thailand that differs in having 
elliptic-oblong leaves, with an intramarginal vein, 
more prominent secondary lateral veins and an 
apiculate apex, and ovate-triangular sepals. 

2. Hypericum uralum Buch.-Ham. ex D. Don, Bot. 
Mag. (1823) t. 2375. — H. patulum (non Thunb.) 
Wall. Cat. (1831) no 4809; Dyer, Fl. Br. Ind. 1 
(1874) 254; Steen. Bull. Jard. Bot. Btzg III, 13 
(1934) 219; Gagnep. Fl. Gen. I.-C. Suppl. 1 (1943) 
248; Hundley & Ko Ko, List trees etc. Burma ed. 
3 (1961) 19. — Norysca urala (Buch.-Ham. ex D. 
Don) K. Koch, Hort. Dendrol. (1853) 66, no 3; 
Y. Kimura in Nakai & Honda, Nova Fl. Jap. 10 
(1951) 102. — H. patulum var. uralum (Buch.- 
Ham. ex D. Don) Koehn-e, Deut. Dendrol. (1893) 
415. — H. garrettii var. ovatum Craib, Fl. Siam. 
En. 1 (1925) 111. 

Bushy shrub, 1.2-1.8 m (in Sumatra), with 
arching, sometimes frondose branches; branchlets 
4-lined or 4-angled when young, eventually becom- 
ing 2-lined or terete. Leaves subsessile or up to 
H mm petiolate, l-AM by ^A-lM cm, lanceolate to 
ovate, apex acute to rounded-apiculate, base 
narrowly or rarely broadly cuneate, very glaucous 
below; 3 main lateral veins, with scarcely visible 
lax reticulate venation; glands all pale, shortly 
striate towards midrib, otherwise punctate. 
Inflorescence l-3(-10)-flowered, terminal (1-2 
nodes), corymbose, and often also lateral, racemi- 
form. Flowers 11/^-3 cm 0, ± concave; buds 
broadly ovoid to globose, obtuse to rounded. 
Sepals VA-9 by (l-)2-6\i mm, free, ± imbricate, 
oblong or elliptic to obovate-spathulate, rounded, 
entire, midrib invisible or almost so; spreading or 
ascending in flower, ascending in fruit; glands all 
pale, linear. Petals bright yellow to golden yellow, 
9-18 by 5-12 mm, broadly obovate to obovate- 
circular, entire, caducous; apiculus distinct to 
obscure, ± rounded; glands all pale, linear. 
Stamen fascicles 5, 4-6(-8) mm long, c. !4-'/i as 
long as petals, each with 40-60 stamens, caducous ; 
anthers bright yellow to orange-yellow, gland 
amber. Ovary 3-5 mm, ovoid to globose; styles 5, 
2H-4'/^ mm, (0.6-)0.7-0.9 times as long as ovary, 
partially or wholly outcurving, free; stigma 
narrowly capitate; placentae 5, axile. Capsule 
7-11 mm, subglobose to globose. Seeds dark 
brown, c. M mm, cylindric-ellipsoid, not curved, 
slightly carinate, shallowly linear-reticulate. 

Distr. Thailand, China (Yunnan, Szechuan), 
Tibet, Burma, Khasia and Himalaya from Assam 
to Kashmir, in Malesia: northern half of Sumatra 
(Mt Kerintji northward to Mt Telong in the Gajo 

Ecol . Grassy or rocky slopes, pastures, thickets, 
open woodland and montane forest, 1700-3300 m. 

Note. The Sumatran form of H. uralum tends 
to be more luxuriant and to have larger flowers 
than those elsewhere; but its more erect habit is 
matched by some Nepal specimens. The form in 
cultivation in Europe, dwarfer, with arching 


Hypericaceae (Robson) 


frondose branches and small flowers, is apparently 
confined to the Himalaya; but the variation be- 
tween these extreme forms appears continuous. 

3. Hypericum monog>'num Linne, Sp. PI. ed. 2 
(1763) 1107; Mill. Card. Diet. ed. 8 (1768) /jo 11; 
Bl. Bijdr. 1 (1825) 141. — H. chinense Linne, 
Syst. Nat. ed. 10, 2 (1759) 1184, non Osbeck, 
Dagbok Ostind. Resa (1757) 244; Amoen. Acad. 
8 (1785) 323; Curtis, Bot. Mag. 10 (1796) t. 334; 
Choisy, Prod. Monogr. Hyper. (1821) 40; in DC. 
Prod. 1 (1824) 545; in Zoll. Syst. Verz. (1854) 150; 
Pi. Jav. (1858) 5; Hassk. PI. Jav. Rar. (1848) 278; 
FoRB. & Hemsl. J. Linn. Soc. Bot. 23 (1886) 72; 
Back. Schoolfl. Java (1911) 87; Hayata, Ic. PI. 
Forraos. 1 (1911) 78; Keller in E. & P. Nat. Pfl. 
Fam. ed. 2, 21 (1925) 176; Hand.-Mazz. Symb. 
Sin. 7 (1931) 401; Back. & Bakh. /. Fl. Java 1 
(1963) 382; Robson, J. R. Hort. Soc. 95 (1970) 
489. — H. aureum Lour. Fl. Coch. (1790) 472. — 
Norysca chinensis (L.) Spach, Hist. Nat. Veg. Phan. 
5 (1836) 427; Ann. Sc. Nat. H, Bot. 5 (1836) 364; 
Bl. Mus. Bot. Lugd. Bat. 2 (1856) 22; Miq. Fl. 
Ind. Bat. 1, 2 (1859) 514; Kimura in Nakai & 
Honda. Nova Fl. Jap. 10 (1951) 103. — Norysca 
aurea (Lour.) Bl. Mus. Bot. Lugd. Bat. 2 (1856) 
22. — Norvsca punctata Bl. I.e. 23. — H. chinense 
ssp. obtusi'folium O.K. Rev. Gen. PI. 1 (1891) 60. 

Bushy shrub, '/2-1.3 m, with spreading branches; 
branchlets 2(-4)-lined when young, eventually 
becoming terete. Leaves sessile or with petiole up 
to I'/i mm, 2-4'/2 by 1-154 cm, elliptic or oblong to 
obianceolate, apex obtuse or minutely apiculate 
to rounded, base cuneate to rounded or subcordate 
paler below; 4-6 main lateral veins, with intra- 
marginal vein and conspicuous dense reticulate 
venation; glands all pale, punctate. Inflorescence 
\-c. 15-flowered, terminal (1-3 nodes), corymbose. 
Flowers 3-5 cm 0, plane or convex; buds ovoid, 
subacute to acute. Sepals AVi-XO by lH-3 mm, 
free, imbricate, narrowly elliptic or narrowly 
oblong to lanceolate, acute or more rarely obtuse 
to rounded, entire, midrib invisible or almost so; 
spreading or ascending in flower and fruit; glands 
ail pale, linear. Petals golden-yellow to lemon- 
yellow, 2-3 by 1^-1'/^ cm, obovate, entire, 
caducous; apiculus variable, acute to rounded or 
absent; glands all pale, linear. Stamen fascicles 5, 
18-28 mm long, almost equalling petals, each with 
25-35 stamens, caducous; anthers bright yellow, 
gland amber. Ovary 2'/i-4 mm, broadly ovoid to 
subglobose; styles 5, 12-18 mm, c. VA-S times as 
long as the ovary, united almost to the apex; 
stigma small; placentas 5, axile. Capsule 6-10 mm, 
broadly ovoid or ovoid-conic to subglobose. Seeds 
dark reddish-brown, cylindric, curved, narrowly 
carinatc, shallowly linear-reticulate to lincar- 

Distr. SE. China, Taiwan. Introduced to 
England in 1753 and now cultivated in many parts 
of the world, including Java and Celebes. 

Ecol. Only as a cultivated ornamental, largely 
in estates in hill stations up to c. 18(X) m. Not quite 
hardy in temperate areas. 

Notes. The plant cultivated in Malcsia, to 
which the above synonymy and description refer, 
is the type form, with smaller, obtuse to rounded 
leaves and smaller flowers, which is native to 
Kwangtung and adjacent parts of China. 1 he plant 

with larger, acute leaves and larger, more numer- 
ous flowers, known as H. salicifolium Sieb. &. 
Zucc, H. chinense var. salicifolium (Sieb. & 
Zucc.) Choisy, H. chinense ssp. salicifolium (Sieb. 
& Zucc.) O. K., etc., appears to intergrade with 
the typical one in China, so that segregation, even 
as a variety, is not possible. 

Unfortunately the epithet chinense cannot be 

4. Hypericum geminiflorum Hemsl. Ann. Bot. 9 
(1895) 144; Lev. Bull. Soc. Bot. Fr. 54 (1908) 590; 
Hayata, Ic. PI. Formos. 1 (1911) 76; ibid. 3 (1913) 
41. — H. trinervium Hemsl. Ann. Bot. 9 (1895) 
144; Lev. Bull. Soc. Bot. Fr. 54 (1908) 590; 
Hayata, Ic. PI. Formos. 1 (1911) 79. — H. 
loheri Merr. Philip. J. Sc. 4 (1909) Bot. 294; En. 
Philip. 3 (1923) 75. — H. acutisepalum Hayata, J. 
Coll. Sc. Tokyo 30, 1 (1911) 308; Ic. PI. Formos. I 
(1911) 77, t. 15. — 1H. pustulosum Keller in E. 
& P. Nat. Pfl. Fam. ed. 2, 21 (1925) 176. — H. 
lackeyi Elm. Leafl. Philip. Bot. 9 (1934) 3190. — 
Takasagoya geminiflora (Hemsl.) Y. Kimura, Bot. 
Mag. Tokyo 50 (1936) 501, f. 2; in Nakai &. 
Honda, Nova Fl. Jap. 10 (1951) 93, t. 40. — 
Takasagoya acutisepala (Hayata) Y. Kimura, Bot. 
Mag. Tokyo 50 (1936) 501, f. 3 j-o; in Nakai & 
Honda, Nova Fl. Jap. 10 (1951) 92. — Takasagoya 
trinervia (Hemsl.) Y. Kimura, Bot. Mag. Tokyo 
50 (1936) 503; in Nakai & Honda, Nova Fl. Jap. 

Lax shrub, M-\\im, branches long, often 
pendulous then ascending near apex; branchlets 
i-lined when young, eventually becoming terete. 
Leaves subsessile, 2-A^A by 0.6-2.2 cm, oblong to 
elliptic or ovate-oblong, apex subacute to rounded, 
usually apiculate, base broadly to narrowly cune- 
ate, paler below; c. 1 main lateral veins, uniting to 
form strong intramarginal vein, the reticulate 
venation not or scarcely visible; glands all pale, 
punctate. Inflorescence 1 -flowered, terminal and 
on solitary or paired short shoots from up to 14 
nodes below, more rarely axillary shoots 2(-3)- 
flowered with flowers pedunculate. Flowers 2-3 cm 
0, plane; buds ovoid, acute or subacute. Sepals 
{\-)\li-2Hby l-\'A(-2) mm, free or connate at the 
base, broadly ovate or triangular or oblong- 
lanceolate or subcircular, subacute to rounded, 
entire, midrib invisible; spreading in flower, 
ascending in fruit; glands all pale, linear and 
punctate. Petals bright yellow, 9-15 by 5-7 mm, 
narrowly obovate, entire, caducous; apiculus 
absent; glands all pale, linear. Stamen fascicles 5, 
6-10 mm long, c. ?s as long as petals, each with 
6-11 stamens, caducous; anthers bright yellow, 
gland amber. Ovary 2h-3',» mm. narrowly ellip- 
soid; styles 5, 4-6(-7) mm, c. 1.3-2 times as long 
as ovary, completely united; stigmas small; 
placentas 5, axile with small central lacuna. 
Capsule 8-1 1 mm, narrowly cylindric to narrowly 
cylindric-conic. Seeds dark reddish-brown, r. I'j 
mm, fusiform to narrowly ovoid or narrowly 
cylindric, scarcely carinatc, with long terminal 
appendage soniclimcs expanded to form narrow 
wing, very shallowly ciongatc-rcticulatc. 

Distr. Taiwan, in Malf.siu: I'hilippincs (I u/on). 

licol. Stony places, river margins. 1000 1.5(K) m. 

Note. The I'hilippinc plants should clearly be 
included in //. geminiflorum, while I'rk r 688 


Flora Malesiana 

[sen I, vol. 8^ 

Fig. 14. Hypericum sewense N. Robson. a. Habit, nat. size, b. leaf, with black marginal dots, x4, c. 

flower, x2, d. pistil & stamens, x4, e. anther, with gland, xl6, f. pistil, x4, g. cross section of ovary, 

x8, h. sepal, x4, i. petal, x4, j. dehisced fruit, x4, k. seed, x 16 {a-k Savers NGF 21418). 


Hypericaceae (Robson) 


(from Taiwan, Prov. Taitung) and Hayata's and 
Klmura's figures indicate that H. ocutisepahun 
cannot be distinguished from it. H. trinerviuin 
seems rather distinct, at first glance, in having 
stouter shoots and pedicels and relatively broader 
leaves with a strong submarginal vein. As each of 
these characters appears in specimens of typical 
H. geminiflorum, although not in the same in- 
dividual, it seems best to regard H. trinervium as a 
local race of H. geminiflorum. 

5. Hypericum sewense N. Robson, Blumea 20 
(1972)254. — Fig. 14. 

Erect shrub, 0.6 m, branches divaricate-as- 
cending; branchlets 4-lined, flattened when young, 
soon 2-lined, eventually terete. Leaves sessile, 
8-12 by 2-4 mm, lanceolate to narrowly elliptic, 
apex rounded, base narrowly cuneate-amplexicaul, 
concolorous, plane, spreading or ascending; c. 5 
main lateral veins, mostly from basal loops, ± 
parallel, branched above on outer side only, 
without marked reticulate venation; laminar 
glands pale, linear near the base, sometimes with 
rows of streaks or dots on either side, becoming 
punctate towards apex and margin; intramarginal 
glands pale and black. Inflorescence 1 -flowered, 
without flowering shoots in uppermost axils or 
rarely with 1, branching profusely farther down; 
pedicels shorter than uppermost leaves, 4-8 mm 
in fruit. Flowers 2.2-2.8 cm 0, ± plane; buds 
narrowly ovoid, subacute. Sepals 5-7 by \M-2^A 
mm, free, imbricate, lanceolate, rounded to 
subacute, entire; laminar glands pale, mostly 
linear; submarginal glands pale or occasionally 
black. Petals bright yellow, 10-14 by 5-6'/^ mm, 
obovate, entire, persistent; apiculus almost absent; 
laminar glands pale, linear, sometimes interrupted 
distally; submarginal glands absent or 1-3, sessile, 
black, on or near apiculus. Stamens obscurely 
3-fascicled, c. 20, longest 8-9 mm, c. ^A as long as 
the petals, persistent; anthers bright yellow, gland 
black. Ovary 214-3 mm, ovoid; styles 3, 2!^-3 mm, 
equalling ovary, divergent; stigmas scarcely 
capitate; placentas 3, parietal. Capsule 7-9 by 
4-5 mm, ovoid, longitudinally vittate. Seeds 
yellow-brown, c. 0.8 mm, cylindric, scarcely 
carinatc, densely linear-foveolate. 

Distr. Malesia: New Guinea (Madang Distr.), 
one collection. Fig. 15. 

Ecol . Boggy tussock sedge-grassland, c. 2700 m. 

Fig. 15. Range of Hypericum scwensc N. RonsoN 

(A), H. hifurcalum N. Roi«V)S (■), //. macKrcnnrii 

F.v.M. ssp. macfircKorii (•) and (litln ssp. punc- 

latum N. Ro».v.»N ( j). 

Note. H. sewense, an apparently very local 
species, is the most primitive member of sect. 
Humifusoideum. Its nearest relatives belonging to 
another section are probably in sect. Campylo- 
sporus, from Africa, Madagascar and the Mas- 
carene Is. 

6. Hypericum macgregorii F.v.M. Trans. R. Soc. 

Vict. 1, 2 (1889) 12; Burk. Kew Bull. (1899) 97; 
Laut. Bot. Jahrb. 58 (1922) 4; Steen. Bull. Jard. 
Bot. Btzg III, 13 (1934) 219. — Fig. 18f. 

Erect shrub, 0.15-1 m, branches strict, creeping 
and rooting at the base; branchlets 2-lined when 
young, eventually terete. Leaves sessile, 4-9 by 
1-3 mm, ovate-lanceolate to elliptic or narrowly 
elliptic-oblong, apex subacute to rounded, base 
cuneate, concolorous, plane or ± carinate, ± 
imbricate-appressed; c. 6 main lateral veins, i 
parallel, not visibly branching, uniting near 
margin and apex, without visible reticulate vena- 
tion; laminar glands pale, linear, becoming ± 
interrupted or punctate towards margin; intra- 
marginal glands pale or black. Iiiflorescence 
1 -flowered, without shoots in uppermost axils, 
branching farther down stem; pedicels usually 
shorter than uppermost leaves, 2-6(-10) mm in 
fruit. Flowers 2-2'/2 cm o, ± plane; buds narrowly 
ovoid, subacute. Sepals 3-6!/i by M-lM mm, free, 
not imbricate, elliptic to linear-lanceolate, sub- 
acute to obtuse or rarely rounded, entire; laminar 
glands pale, linear to punctate; submarginal 
glands pale or black. Petals dark yellow to pale 
yellow, sometimes red-tinged below, 7-15 by 3-6 
mm, oblanceolate, entire, persistent; apiculus 
absent or almost so; laminar glands pale, linear, 
sometimes interrupted distally; submarginal glands 
absent. Stamens obscurely 3-fascicled, c. 17-24, 
longest 5-8 mm, c. ^/i as long as petals, persistent; 
anthers bright yellow, gland amber. Ovary 2 mm, 
ovoid; styles 3(^), 2 mm, equalling ovary, 
divergent; stigmas not capitate; placentas 3 (4) 
axile. Capsule 6-8 by 3-4 mm, ovoid, longitudinally 
vittate. Seeds yellow-brown, c. 1 mm. cylindric to 
cylindric-ellipsoid, slightly carinate, densely linear- 

ssp. macgregorii. 

Leaves with laminar glands linear, becoming 
interrupted or punctate towards margin; intra- 
marginal glands pale. Sepals with laminar glands 
all or mostly linear; submarginal glands always 

Distr. Malesia: New Guinea (Morobc, Central 
and Milne Bay Distr.). Fig. 15. 

Ecol. Open alpine grassland, usually in shallow 
soils, (1500 )27(K)39(K)m. 

Note. The population on Mt Dayman (Mancao 
Ra.) is aberrant in having (a) leaves relatively 
broader (leaf index 2.2 3.1 compared with 2'i, 3!» 
in the typical form) with the apex less narrowed 
and (b) longer pedicels (5 10 mm in fruit com- 
pared with 2 6 mm in the typical form). 

ssp. punctatum N. Roiison, Hlumca 20 (1972) 256. 

leaves with laminar glanils mostly dots or short 
streaks; intramarginal glands black. Sepals with 
laminar glands punctate; submarginal glands 
sometimes black. 

Distr. Malesia: New Guinea (lake Hahbcma). 
one collection. Fig. 15. 


Flora Malesiana 

[ser. I, vol. 8^ 

Note. Ssp. ptinctatiim is very similar in aspect 
to the Mt Maneao population. 

7. Hypericum saruwagedicum Diels, Bot. Jahrb. 62 
(1929) 482. — H. macgregorii {non F.v.M.) Hoogl. 
Blumea, Suppl. 4 (1958) 231. — Fig. 18e. 

Erect, bushy shrub, 0.1-1.2 m, branches strict, 
creeping and rooting at the base; branchlets 4- 
lined when young, soon 2-lined, eventually terete. 
Leaves sessile, 2-9 by li-3 mm, narrowly elliptic 
or lanceolate-elliptic to linear, apex rounded, base 
narrowly cuneate, concolorous, incurved and 
slightly carinate at the base, ± imbricate-ap- 
pressed; c. 6 main lateral veins, ± parallel, not 
visibly branching, uniting near margin and apex, 
without visible reticulate venation; laminar glands 
pale, linear, sometimes interrupted near margin; 
intramarginal glands pale only or pale and black. 
Inflorescence 1 -flowered, with shoots in uppermost 
axils, branching farther down stem; pedicels 
shorter than uppermost leaves, 2-A(-6) mm long in 
fruit. Flowers \-2M cm 0, plane or ± concave; 
buds narrowly ovoid-cylindric, subacute to rounded. 
Sepals 3H-8 by 1-3 mm, free, imbricate or not, 
ovate or lanceolate to elliptic or narrowly oblong, 
subacute to rounded, entire; laminar glands pale, 
all or mostly linear; submarginal glands pale only 
or pale or black. Petals dark yellow to pale 
yellow, sometimes red-tinged below, 7-15 by 3-6 
mm, oblanceolate, persistent; apiculus absent or 
almost so; margin entire, or rarely with a few i 
prominent marginal glands; laminar glands pale, 
linear, sometimes interrupted distally; marginal 
glands black or absent. Stamens not obviously in 
fascicles, 13-26, longest 4-7(-8) mm c. M-M as 
long as the petals, persistent; anthers bright yellow, 
gland amber or black. Ovary 2-3 mm, ovoid; 
styles 3, Wi-lMmm, M-M as long as the ovary, 
divergent; stigmas ± capitate; placentas 3, 
parietal, intrusive. Capsule {5\i-)6-9 by 3H-5 mm, 
ovoid, longitudinally vittate. Seeds orange-brown, 
Vi-y\ mm, cylindric, carinate, densely linear- 

Distr. Malesia: New Guinea (Mts Wilhelmina 
& Carstensz, Lake Habbema, E. Highlands, 
Madang, Morobe, Central & Milne Bay Distr.). 
Fig. 16. 

Ecol . Alpine grassland or open scrub, usually in 
well-drained soil, 2800-4300 m, but to c. 1800 m in 
Milne Bay District. 

Notes. H. saruwagedicum is a variable species 
in which the variation falls into four geographical, 
more or less morphological intergrading races. 

Fig. 16. Range of Hypericum saruwagedicum 

Although it does not appear desirable to name 

these formally, average members can be recognised 

by the following characters: 

Variant 1 (Mt Wilhelm): Leaves large. Flowers 
large. Black glands usually on leaves and sepals, 
not on petals or anthers. Shoot apex outcurving. 

Variant 2 (Territory of New Guinea except Mt 
Wilhelm, West Papua): Leaves medium. Flowers 
medium to large. Black glands on anthers only 
or absent. Shoot apex erect. 

Variant 3 (West New Guinea): Leaves small. 
Flowers large. Black glands on leaves, sepals, 
petals and anthers. Shoot apex sometimes 

Variant 4 (Papua, Milne Bay District): Leaves 
small. Flowers small. Black glands on anthers 
and (rarely) leaves. Shoot apex erect. 

8. Hypericum bifurcatum N. Robson, Blumea 20 
(1972)256. — Fig. ISd. 

Erect shrub (? or woody herb), Vi-\\4vn, 
branches ± strict, rooting at the base; branchlets 
2-lined when young, eventually terete. Leaves 
sessile, 7-13(-16) by lH-6 mm, narrowly ovate to 
narrowly elliptic-oblong, apex rounded, base 
cuneate to rounded, concolorous, plane, ascending 
or appressed ; c. 7 main lateral veins, mostly from 
basal loops, ± parallel, little branched, uniting 
near margin and apex, without marked reticulate 
venation; laminar glands pale, linear, sometimes 
with rows of dots on either side, becoming ± 
interrupted towards margin; intramarginal glands 
pale only. Inflorescence 1 -flowered, with strong 
flowering shoots in uppermost axils and often 
weaker ones in axils immediately below, the 
repeated branching producing an eff"ect of bifur- 
cation; pedicels equalling or exceeding uppermost 
leaves, 8-15 mm, long in fruit. Flowers \\i-2\4 cm 
0, plane; buds narrowly ovoid, rounded. Sepals 
4-6 by \ii-2 mm, free, imbricate, ovate-lanceolate, 
subacute, entire; laminar glands pale, all or mostly 
linear; submarginal glands pale or reddish. 
Petals bright yellow, orange- or red-tinged below, 
9-14 by 3-5 mm, obovate to oblanceolate, entire, 
persistent; apiculus absent or almost so; laminar 
glands pale, linear, sometimes interrupted distally ; 
marginal glands absent or one, sessile, reddish, on 
apiculus. Stamens obscurely 3-fascicled, 25-35, 
longest 6-8 mm, c. Vi as long as the petals, 
persistent; anthers bright yellow, gland black. 
Ovary 2 mm, ovoid; styles 3, 2 mm, equalling the 
ovary, divergent; stigmas narrowly capitate; 
placentas 3, parietal except at the very base. 
Capsule 6-9 by 7iM-AM mm, ± broadly to narrowly 
ovoid or ovoid-pyramidal, longitudinally vittate. 
Seeds yellow-brown, c. Vi mm, cylindric to cylindric- 
ellipsoid, slightly carinate, densely linear-foveolate. 

Distr. Malesia: New Guinea (E. Highlands & 
Morobe Distr.). Fig. 15. 

Ecol. Wet to dry grassland, c. 2200-3000 m. 

Note. The pair of flowering shoots in each 
uppermost axil distinguishes H. bifurcatum from 
all other members of sect. Humifusoideum except 
some forms of H. papuanum in which, however, 
the leaf gland pattern diff"ers and 4-5 styles often 

9. Hypericum papuanum Ridl. Trans. Linn. Soc. 
II, Bot. 9 (1916) 19; Laut. Bot. Jahrb. 58 (1922) 


Hypericaceae (Robson) 


5; Steen. Bull. Jard. Bot. Btzg III, 13 (1934) 219. — 
H.japonicum (non Thunb. ex Murr.) Warb. Bot. 
Jahrb. 16 (1893) 14. — H. luac^regorii {non F.v.M.) 
Lalt. Nova Guinea 8 (1912) 843. — H. hellwigii 
Lalt. Bot. Jahrb. 58 (1922) 4; Keller in E. & P. 
Nat. Pfl. Fam. ed. 2, 21 (1925) 181; Steen. Bull. 
Jard. Bot. Btzg III, 13 (1934) 219. — //. habbe- 
mense A. C. Smith, J. Arn. Arb. 22 (1941) 343. — 
Fig. 18c. 

Erect or ± straggling shrub or woody herb, 
0.1-1.3 m, branches strict or ± lax, creeping and 
rooting at the base; branchlets 2-4-lined when 
young, eventually terete. Leaves sessile, 0.6-2.5 by 
0.3-1.7 cm, narrowly ovate or ovate-triangular to 
elliptic or broadly ovate or subcircular, apex 
subacute or rarely acute to rounded, base rounded 
to cordate, concolorous, plane, spreading or 
ascending; 4-5(-6) main lateral veins, i parallel 
or diverging, much branched, uniting near margin 
and apex, v\ ith marked, usually ± dense, reticulate 
venation; laminar glands pale, linear towards the 
base or wholly punctate and striate or punctate 
only; intramarginal glands pale only or partly or 
wholly black. Inflorescence 1 -flowered, with or 
without flowering shoots in uppermost axils, or 
regularly dichasial or intermediate and irregular; 
pedicels usually exceeding uppermost leaves, 4-20 
mm in fruit. Flowers 1.8-2.6 cm 0, plane; buds 
ovoid to ellipsoid, subacute. Sepals 3-7(-8) by 
l-2li(-VA) mm, free, imbricate, ovate to lanceolate 
or narrowly oblong or sometimes broadly ovate 
and foliaceous, acute to rounded, entire; laminar 
glands pale, linear; submarginal glands black or 
absent. Petals bright yellow, 9-15 by 4-9 mm, 
obovate, entire, persistent; apiculus short or 
obsolete; laminar glands pale, linear, sometimes 
striate distally; marginal glands absent or black, 
sometimes only in apiculus, rarely few to numer- 
ous elsewhere on margin. Stamens not obviously in 
fascicles (15-)25-40(-50), longest 6-9 mm, c. V* as 
long as the petals, persistent; anthers bright 
yellow, gland amber or occasionally black. Ovary 
(2-)2'/i-3(-4) mm, narrowly or rarely broadly 
ovoid; styles 3 (4-5), 2-3 mm, M as long to as long 
as the ovary, divergent; stigmas narrowly to 
broadly capitate; placentas 3(-4-5), parietal. 
Capsule (5-)7-9(-10) by 3-4'A mm, narrowly ovoid 
or rarely broadly ovoid to ellipsoid, longitudinally 
vitiate. Seeds yellow-brown to dark brown, 0.7- 
0.8 mm, cylindric or subcylindric, scarcely carinate, 
densely lincar-foveolate to lincar-scalariform. 

Distr. Malesia: New Guinea (widespread in all 
mountainous regions). Fig. 17. 

Ecol. Wet to dry alpine grassland and bogs, 
screes, 1800-3800 m. 

Note. H. papuanwn is a very variable species in 
which the extreme forms, although quite distinct in 
appearance, are linked by intermediates with 
varying combinations of characters, so that the 
morphological trends are not coordinated. These 
trends are: 

(1) Leaves narrowly ovate and rt crowded with 
laminar glands mostly linear (in E., W. and S. 

Fig. 17. Range oi Hypericum papuanum Kidi 

I ig. 18. Hypericum pulogcnsc Ml kk. a. Habit, nat. 
size, b. leaf, > 4. — Leaves from underside of: 
c. //. papuanum RiDL.. > 4. d. //. hi/urcalum N. 
RoHsoN, /4, c. //. saruwanedinim Dni.s, - 10, 
f // macfircK»rii\- y.M., -4(</ /> Romiuns v./;.. dil. 
2V io 1965.f Sayik.s NGF 21419. d NCil- 15969. 
e IhKXii.ANU 10002. /Hartley NGF 12799). 


Flora Malesiana 

[sen I, vol. 8^ 

Highlands mainly) to broadly ovate or sub- 
circular, not crowded, with laminar glands all 
punctiform (constant in W. New Guinea and E. 
Papua, Northern and Milne Bay Districts). 

(2) Leaves, sepals, petals and anthers without black 
glands (mainly eastern) to with black glands, 
forming a continuous intramarginal row in the 
leaves (constant in W. New Guinea) and 
sepals and a continuous marginal row in the 
petals (rare). 

(3) Inflorescence 1 -flowered (mainly eastern) to 
regularly dichasial (mainly western). 

(4) Styles and placentae 3 with ovary and capsule 
narrowly ovoid (mainly eastern) to styles and 
placentae 4-5 with ovary and capsule ± 
broadly ovoid (mainly western). According to 
A. C. Smith, I.e., H . habbemense may have up 
to 6 styles but only up to 5 placentae. A re- 
examination of the syntype material has failed 
to reveal an ovary with 6 styles and 5 placentae, 
a combination of characters that is probably 
teratological in origin. 

(5) Habit dense with ± ascending branches 
(widespread) to lax with ± spreading branches 
(Madang and Morobe Districts). 

10. Hypericum pulogense Merr. Philip. J. So. 5 
(1910) Bot. 364; En. Philip. 3 (1923) 75. — Fig. 

Suffrutescent herb, 20-40 cm, branches strict, 
erect or ascending to decumbent from slender 
branching rhizome, 2(^1— 6)-lined or narrowly 
2-winged when young, sometimes eventually 
terete. Leaves sessile or very shortly petiolate, 
0.8-L2(-2) by 0.3-0.7 cm, elliptic to oblong or 
ovate, apex obtuse to rounded, base broadly 
cuneate to rounded, glaucous beneath, margin 
recurved (at least when dried), spreading or 
ascending; c. 4 main lateral veins, ± parallel, 
slightly branched, uniting near margin and apex, 
with scarcely visible reticulate venation; laminar 
glands pale, ± prominent, punctate and shortly 
striate; intramarginal glands pale or apparently 
absent. Inflorescence 1-flowered, with or without 
flowering shoots in uppermost axils, or regularly 
dichasial with 3-c. 10 flowers; pedicels shorter 
than uppermost leaves, 4-6 mm in fruit. Flowers 
2-2 '/^(-3) cm 0, plane; buds narrowly ovoid, 
subacute to obtuse. Sepals 4-6 by L8-2.4 mm, 
free, imbricate, lanceolate to oblong or elliptic- 
oblong, rounded or subapiculate, entire; laminar 
glands pale, linear and punctate; submarginal 
glands pale. Petals bright yellow, 10-12(-14) by 
4-6 mm, narrowly oblong-obovate, entire to 
subentire, persistent; apiculus small, glandular; 
laminar glands pale, linear, becoming striate and 
punctate distally; marginal glands pale. Stamens 
not obviously in fascicles (or in 5 fascicles, fide 
Merrill), c. 30-60, longest c. 9 mm, c. M as long 
as the petals, persistent; anthers bright yellow, 
gland amber. Ovary 3^ mm, broadly to narrowly 
ovoid; styles 3, c. 5 mm, 1.4-2 times as long as 
ovary, divergent; stigmas not capitate; placentas 3, 
axile. Capsule 5-8 by 4-5 mm, narrowly ovoid to 
ovoid, longitudinally and diagonally vittate. Seeds 
yellow-brown to dark brown, ^A~\ mm, cylindric, 
not carinate, densely linear-scalariform. 

Distr. Malesia: Philippines (Luzon). Fig. 20. 

Fig. 19. Hypericum beccarii N. Robson. a. Habit, 
nat. size, b. flower, x4, c. leaf, x4. — H. beccarii 
ssp. steenisii N. Robson. d. & f. Habit, nat. size, 
e. leaf, x4 (a-c Bunnemeijer 976a, d-e Van 
Steenis 85 14, /Van Steenis 9599). 


Hypericaceae (Robson) 


Ecol. Summit grasslands and open places in 
mossy forest, c. 2800 m. 

Note. H. pulogense is known only from Mt 
Pulog and Mt Tabayoc. Its nearest relatives (other 
than H. beccarii) are in New Guinea. The only 
specimen from Mt Tabayoc has slenderer, more 
decumbent stems than those from Mt Pulog. In 
these respects it approaches H. beccarii. 

11. Hypericum beccarii N. Robson, Blumea 20 
(1972) 260. — H. japonicum var. pinnatinervium 
Bakh./. in Back. & Bakh./. Fl. Java 1 (1963) 382, 
descr. angl. — Fig. 19. 

Weak perennial (? or annual) herb, c. 2^5 cm, 
stems wiry, decumbent or ascending (or erect fide 
Bakh. /.), creeping and branching irregularly, 
rooting at the base, narrowly 4-6-lined when 
young, often becoming 2-lined. Leaves 0.2-1 H mm 
petiolate, 2\i-\QM by M-6 mm, broadly oblong or 
elliptic-oblong to oblanceolate or linear, ape.x 
rounded to subacute, apiculate to muticous, base 
rounded to cuneate, glaucous beneath, margin not 
or slightly recurved, spreading; c. 3 main lateral 
veins, ± parallel, with reticulate branchings, 
uniting near margin and apex with dense con- 
spicuous reticulate venation; laminar glands pale, 
— prominent or not, irregularly punctate; intra- 
marginal glands dark or pale, irregular. Inflores- 
cence 1 -flowered, with flowering shoots in upper- 
most axil(s); pedicels usually exceeding uppermost 
leaves, (2-)5-17 mm in fruit. Flowers c. 7-10 mm 
0, plane; buds narrowly ovoid, obtuse. Sepals 
2li-5 by 0.6-1 .4 mm, free, ± broadly or not 
imbricate, elliptic-oblong to linear, rounded to 
subacute or apiculate; margin entire to irregularly 
glandular-ciliate; laminar glands pale, punctate to 
shortly striate; submarginal to marginal glands 
black. Petals yellow, 3-7 by 2-21-2 mm, oblong- 
oblanceolate, persistent; apiculus small, glandular 
or glandular-ciliate; margin entire or with few 
subscssile glands; laminar glands absent or few 
near ap)ex, pale or black, punctate ; marginal glands 
black. Stamens in 3 fascicles, c. 15-22, longest 
2'A-Slitnm, c. '/i-'/i as long as the petals, per- 
sistent; anthers yellow, gland black. Ovary li^mm 
long, narrowly ovoid; styles 3, c. \'A mm, about as 
long as the ovary, gradually divergent; stigmas 
slightly capitate; placentas 3, axile. Capsule 3-5'A 
by 2-i'A mm, -± narrowly ovoid, densely and ± 
prominently longitudinally vitiate or almost 
smooth. Seeds reddish brown, 0.7-0.9 mm, 
cylindric, not carinate, densely shallowly linear- 

ssp. beccarii. — Fig. 19a-c. 

Leaves 0.2-1 mm petiolate, 0.4-1 by 0.2-0.6 cm, 
broadly oblong or elliptic-oblong to narrowly 
obovatc, apex rounded, apiculate to muticous, 
base rounded to cuneate; laminar glands ! 
prominent. Sepals 3'A-5 by I -2 mm, ' broadly 
imbricate. Petals 5-7 mm; laminar glands (when 
present) black. Stamens c. 20 22. longest A'A-S'i 
mm. Capsule 4 5',i by 2A--VA mm, densely and i 
prominently vittatc. 

[Jistr. Malcsia: West Central Sumatra (Mis 
Singalang, Talang. Talamau. Kcrintji). W. Java 
(Mt i'apandajan, once collected). I ig. 20. 

tcol. Iktwccn 1 800 and KKM) m. 

Note. The specimens with broad leaves and 

Fig. 20. Range of Hypericum pulogense Merr. 

(n), H. beccarii N. Robson ssp. beccarii (#) and 

ditto ssp. steenisii N. Robson (O). 

large flowers (from Mts Talamau and Singalang) 
approach most closely to H. pulogense and other 
related species in this section. The Javanese 
specimens, although smaller in all parts, cannot 
otherwise be separated from those of Central 

ssp. steenisii N. Robson, Blumea 20 (1972) 261. — 
Fig. 19d-f. 

Leaves ^A-\ }A mm petiolate, 2'/i-10li by H-4 mm, 
oblanceolate to linear, apex rounded to subacute, 
i: apiculate, base cuneate; laminar glands not 
prominent. Sepals 2' 2-5 by 0.6-1.4 mm, not 
imbricate. Petals 3-6 mm; laminar glands (when 
present) pale. Stamens c. 15-20, longest 2;*^ls 
mm. Capsule 3^1-2 by 2-3 mm, sparsely and 
obscurely longitudinally vittate (± smooth). 

Distr. Malesia: N. Sumatra (Gajo Lands; Mts 
Losir & Kemiri). Fig. 20. 

Ecol. Along streamlets, in open vegetation, 
2700-3300 m. 

Note. Ssp. steenisii differs essentially from .isp. 
beccarii in having narrower leaves and smooth 
capsules with few vittac. 

12. Hypericum pseudopctiolatum R. Keller, Bull. 
Herb. Boiss. 5(1897)638; Y. Kimura. J. Jap. Bot. 
15 (1939) 295; in Nakai & Honda, Nova PI. Jap. 
10 (1951) 197, (V. 68, 69.— //. S. 
Suzuki, Trans. Nat. Hist. Soc. Formosa 20(19.10) 
239; Ann. Kcp. Taihoku Bot. Card. 1 (1931) 158; 
in Masamune, Short Fl. Formosa (1936) 141. — 
//. pseudopctiolatum var. taihc:anense (S. Suzuki) 
Y. KiMURA. J. Jap. Bot. 15 (1939) 296; Bot. Mag. 
Tokyo 54 (1940) 81 ; in Nakai & Honda, Nova II. 
Jap. 10(1951)204. 

Perennial herb, 15 40 cm, stems slender, 
ascending, branching above, not rooting, terete. 
Leaves sessile or up to »4 mm petiolate. 4 17 by 
2 6 mm. lanceolate or ovale to clliptic-oblong. 
apex rounded, base cuneate to roundcil (or the 
upper ones somclifucs cordalc-ainplcxicaul), paler 
beneath; 3 4 main lateral veins, with reticulate 
branchings, uniting near margin ami apex; reticu- 
late venation I- marked, dense; laminar glands 
pale, large, often prominent, dense, punctate; 


Flora Malesiana 

[ser. I, vol. 8^ 

intramarginal glands black, dense. Inflorescence 
\-c. lO-flowered, cymose, monochasial after 2nd 
grade, lax; pedicels sometimes exceeding upper- 
most leaves, 3-6 mm in fruit. Flowers 8-10 mm 0, 
concave; buds ellipsoid, obtuse. Sepals 3-5M by 
\-\\i mm, free, imbricate, lanceolate to elliptic- 
oblong, obtuse to rounded, entire; laminar glands 
pale, punctate or shortly striate, few; submarginal 
glands all black or basal ones pale. Petals yellow, 
5-6 by 2 mm, narrowly obovate or oblong-obovate, 
persistent; apiculus absent; margin entire or with 
1-2 subapical prominent glands; laminar glands 
absent or few near apex, pale, punctate; marginal 
glands black, few, sometimes prominent. Stamens 
in 3 (abnormally 4-5) fascicles, c. 30, longest c. 
4 mm, c. % as long as the petals, persistent; anthers 
yellow; gland black. Ovary c. 3 mm, ellipsoid; 
styles 3 (abnormally 4-5), VAmm, shorter than 
ovary, divergent; stigmas scarcely capitate; pla- 
centas 3 (abnormally 4-5), axile. Capsule 4-5 by 
3-4 mm, broadly ovoid to subglobose, longi- 
tudinally vittate. Seeds mid-brown, \A mm, 
cylindric-ellipsoid, not carinate, densely shallowly 

Distr. Taiwan (north and central), and Japan 
(north to southern Hokkaido), in Malesia: 
Philippines (Luzon, Pauai). 

Ecol. Occurs at 2100m in Luzon, but reaches 
2500 m near Arisan in Taiwan. 

Note. The Philippine plant resembles some of 
the specimens from Taiwan. The variation in that 
island, however, overlaps that of the Japanese H. 
pseudopetiolatum to such an extent that it is im- 
possible to recognise H. taihezanense, even at 
varietal level. 

13. Hypericum petiolulatum Hook./. & Thoms. ex 
Dyer, F1. Br. Ind. 1 (1874) 255; Ltv. Bull. Soc. 
Bot. Fr. 54 (1908) 594; Hand.-Mazz. Symb. Sin. 7 
(1931)402; Gagnep. F1. Gen. L-C. Suppl. 1 (1943) 
248. — H. thomsonii R. Keller, Bot. Jahrb. 33 
fl904) 552, quoad typum. — H. mutilutn (non L.) 
RiDL. J. Fed. Mai. Stat. Mus. 8 (1917) 17. — H. 
petiolatum (non L., nee L./., nee Walt., nee 
Lour.) R. Keller in E. & P. Nat. Pfl. Fam. ed. 2, 
21 (1925) 179. 

Annual herb, 10-60 cm, stems erect to ascending 
or procumbent, branching ± widely, rooting, 
terete. Leaves 1-3 mm petiolate, 0.9-3 by 0.5- 
1(-1 .6) cm, oblong or lanceolate-elliptic to obovate 
or subcircular, apex rounded or rarely obtuse, base 
cuneate or attenuate to rarely rounded, ± glaucous 
beneath; 3 main lateral veins, not or scarcely 
branched, uniting near margin and apex; reticulate 
venation obscure, rather dense to lax; laminar 
glands pale, very rarely with 1-2 black, large, 
usually dz prominent, ± dense, punctate; intra- 
marginal glands black, ± dense or irregular. 
Inflorescence (l-)3-9-flowered, cymose, with sub- 
sidiary inflorescences on often long lateral branches, 
monochasial after 1st grade, lax; pedicels ex- 
ceeding uppermost leaves, 4-13 mm in fruit. 
Flowers 5-7 mm o, plane; buds ellipsoid, obtuse. 
Sepals 2.3-3.4 by 0.5-0.9 mm, free, not imbricate, 
linear to very narrowly oblong-lanceolate, entire; 
very acute (or rarely (Nepal) narrowly elliptic- 
oblong and subacute); laminar glands pale or 
rarely black, punctate to linear, variable in size 
and number; marginal glands black, few, some- 

Fig. 21. Range of Hypericum petiolulatum Hook. 
/. & Thoms. ex Dyer ssp. petiolulatum (#) and 
ditto ssp. yunnanense (Franch.) N. Robson (O). 

times one apical, or all absent. Petals yellow, 4-5H 
by 1-1 '/i mm, oblanceolate, entire, persistent; 
apiculus small, acute, subapical; laminar glands 
absent or 1-2, pale, punctate; marginal glands 
black, 1-2 on or near the apiculus and occasionally 
also elsewhere. Stamens in 3 fascicles, 9-22, longest 
{2]4-)3-4\4. mm, c. Vi as long as the petals; anthers 
yellow, gland black. Ovary VA-2mm, ± broadly 
ovoid to subglobose; styles 3(-4), 1-2.1 mm, H~ 
\\i times as long as the ovary, divergent; stigrnas 
not or scarcely capitate; placentas 3(-4), axile. 
Capsule 3-5 by 3-5 mm, broadly ovoid to globose, 
longitudinally vittate. Seeds yellow-brown, 0.5- 
0.6 mm, cylindric, not carinate, densely shallowly 

Distr. Nepal to Burma and Yunnan, in 
Malesia: Sumatra (Mt Kerintji), N. Borneo (Mt 
Kinabalu). Fig. 21. 

Ecol. Stream banks in peaty or stony soil, 
2000-2400 m. 

Note. H. petiolulatum is variable, but the 
Malesian specimens fall within the range of 
variation of the type subspecies. On the other side 
of the Himalayan Range, in Yunnan and Szechuan, 
it is represented mostly by ssp. yunnanense 
(Franchet) N. Robson, in which the styles are 
longer than the ovary, the capsule is ± broadly 
ovoid, the leaf lamina is usually broadest at or 
below the middle and often larger than in ssp. 
petiolulatum, the branching is more regular, and 
the main inflorescence usually occupies 2-3 nodes. 

14. Hypericum gramineum G. Forster, F1. Ins. 
Austr. Prodr. (1786) 53; Vahl, Symb. Bot. 2 
(1791) 86; Bailey, Rep. Austr. Ass. Adv. Sc. 7 
(1898) 429; Queensl. Fl. 1 (1899) 100; Hochr. 
Candollea 2 (1925) 437; R. Keller in E. & P. Nat. 
Pfl. Fam. ed. 2, 21 (1925) 183; Guillaumin, F1. 
Nouv. Caled. (1948) 217. — Ascyrum involutum 
Labill. Nov. Holl. PI. Spec. 2 (1806) 32, t. 174. — 
H. involutum (Labill.) Choisy, Prod. Monogr. 
Hyper. (1821) 50; in DC. Prod. 1 (1824) 549. — 
Brathys billardieri Spach, Ann. Sc. Nat. II, Bot. 5 
(1836) 367. — Brathys forsteri Spach, I.e. — H. 
pedicellare Endl. En. PI. Hueg. (1837) \2. — H. 
aureum Banks & Sol. ex Hook. /. Fl. Nov. Zel. 1 


Hypericaceae (Robson) 


(1853) 36, in svnon. — H. lalandii (non Choisy) 
Dyer. F1. Br. Ind. 1 (1874) 256; Lev. Bull. Soc. 
Bot. Fr. 54 (1908) 589, 593; R. Keller, Bull. 
Herb. Boiss. II, 8 (1908) 187, pro parte; in E. & P. 
Nat. Pfl. Fam. ed. 2, 21 (1925) 181, pro parte; 
Hand.-Mazz. Symb. Sin. 7 (1931) 404; Kanjilal 
& Das, F1. Assam 1, 1 (1935) 100. — H.foetidum 
Hook./. & Thoms. ex Dyer, FI. Br. Ind. 1 (1874) 
257, in svnon. — H. japonicum var. oust rale R. 
Keller, Bull. Herb. Boiss. II, 8 (1908) 186. — H. 
japonicum var. lanceolatum Y. Kimura, Bot. Mag. 
Tokyo 54 (1940) 88. — Sarothra saginoides Y. 
Kimura in Nakai & Honda, Nova Fl. Jap. 10 
(1951) 246, t. 81. — Sarothra graminea (G. 
Forster) Y. Kimura, I.e. 232. 

Perennial or annual herb, (2!^-)5-72 cm, with 
stems erect or decumbent, branching strictly from 
the base (rarely elsewhere) or unbranched below 
the inflorescence, not rooting, 4-lined. Leaves 
sessile, 0.4-2.5 by 0.12-0.8 cm, lanceolate to linear 
or oblong or rarely ovate-lanceolate, apex obtuse 
to rounded, base cordate to rounded or sometimes 
cuneate, amplexicaul, ± glaucous beneath; 1-3 
main basal veins, not or scarcely branched, with 1 
or more lateral veins often visible; reticulate 
venation absent; laminar glands pale, not promin- 
ent, small and dense above, larger and laxer 
beneath; intramarginal glands absent. Inflorescence 
(l-2-)3-c. 30-flowered, cymose and regularly 
dichasial to monochasial or wholly monochasial, 
or with branches in uppermost leaf axil, or mixed, 
sometimes with subsidiary inflorescences from 
upper 1-3 leaf pairs, lax; pedicels usually exceeding 
uppermost leaves, 2-30 mm in fruit. Flowers 
(5-)6-12(-15) mm 0, plane; buds ellipsoid, sub- 
acute. Sepals 2.8-7. 5(-9) by 0.8-2 mm, free, imbri- 
cate, lanceolate to narrowly elliptic, entire, acute 
to subacute, 3-5-nerved, often with ± prominent 
midrib, especially in fruit; laminar glands pale, 
linear towards the base, becoming punctate 
distally; marginal glands absent. Petals pale or 
bright yellow or orange, 5-10 by 2-5 mm, c. 1.3 
times as long as the sepals, obovate to oblanceo- 
late, entire, persistent; apiculus small, mucronate 
to rounded, lateral; laminar and marginal glands 
absent. Stamens not in apparent fascicles, c. 30-50, 
longest 2!^-4 mm, c. M as long as petals; anthers 
yellow, gland amber. Ovary 1 1/^-2.3 mm, narrowly 
ovoid-conic; styles 3, 0.7-1 .8 mm, 'A-*Ao as long as 
ovary, ± divergent; stigmas capitate; placentas 3, 
parietal. Capsule 2\4-S by ]-3'A(-4) mm, narrowly 
ovoid to cyiindric, not vittate. Seeds yellow-brown, 
Vi mm, cyiindric, not carinate, longitudinally ribbed 
with fine transverse striae. 

Oistr. Vietnam, Taiwan, China (Yunnan), 
India (Assam), Bhutan (?), in Malesia: New 
Guinea (Morobc, E., W. & S. Highlands, Western 
Distr.), Australia, New Zealand, New Caledonia. 

Hcol. Cirows in wet to dry, but open and well- 
drained habitats at 10-2600 m. 

Note. //. gramincum appears to remain quite 
distinct from //. japonicum except in Bhutan, where 
hybrid (?) intermediates occur. It can be dis- 
tinguished from //.japonicum by its larger flowers 
and floral parts, longer petals relative to sepals, 
more numerous stamens and, usually, its narrower 
leaves and lanceolate to narrowly elliptic acute 
sepals. In //.Japonicum the sepals rarely show this 
combination of characters. In addition, //. 

gramineum never has the straggling to prostrate 
habit often found in H. japonicum. 

15. Hypericum japonicum Thunb. ex Murray, 
Syst. Veg. ed. 14 (July 1784) 702; Thunb. Fl. Jap. 
(Aug. 1784) 295, t. 31; Lamk, Encycl. M6th. 4 
(1797) 163; Choisy, Prod. Monogr. Hyper. (1821) 
48; in DC. Prod. 1 (1824) 548, ind. var. ramosum 
Choisy, I.e. 549; Bl. Bijdr. 1 (1825) 143; D. Don, 
Prod. Fl. Nepal. (1825) 219; Royle, III. Bot. 
Himal. (1834) 131, t. 24, f. 2; Hook./. Fl. Nov. 
Zel. 1 (1853) 37, incl. var. humifusum (Labill.) 
Hook./.; Choisy in Zoll. Syst. Verz. (1854) 151; 
MiQ. Ann. Mus. Bot. Lugd.-Bat. 2 (1866) 259, 
incl. f. microphylla MiQ. et f. tenuior MiQ.; Prol. 
Fl. Jap. (1866) 147; Dyer, Fl. Br. Ind. 1 (1874) 256; 
Kurz, J. As. Soc. Beng. 43, ii (1874) 84; Franch. 
&. Sav. En. PI. Jap. 2 (1878) 300; Forb. & Hemsl. 
J. Linn. Soc. Bot. 23 (1886) 73; Trim. FI. Ceylon 
1 (1893) 93; R. Keller, Bull. Herb. Boiss. 5 (1897) 
641; Bailey, Queensl. Fl. 1 (1899) 101; Lev. Bull. 
Soc. Bot. Fr. 53 (1906) 501; ibid. 54 (1908) 593; 
Gagnep. Fl. Gen. I.-C. 1 (1908) 286, f. 28, 15- 
21 ; R. Keller, Bull. Herb. Boiss. II, 8 (1908) 185, 
186, incl. var. maximowiczii R. Keller, var. 
thunbergii (Franch. & Sav.) R. Keller, var. 
calyculatum R. Keller et var. simplicius R. 
Keller; Pampanini, Nuov. Giom. Bot. Ital. n.s. 
17 (1910) 670, incl. var. accumbens (Bl.) Pamp.; 
Lev. in Fedde, Rep. 8 (1910) 451, incl. var. 
plurinervium Lev.; Ridl. J. Str. Br. R. As. Soc. 54 
(1910) 16; Back. Schoolfl. Java (1911) 86; Hayata, 
Ic. PI. Formos. 1 (1911) 78; Gamble, FI. Pres. 
Madras 1 (1915) 70; Ridl. Trans. Linn. Soc. 
Lond. II, Bot. 9 (1916) 20; Fl. Mai. Pen. 1 (1922) 
152; Laut. Bot. Jahrb. 58 (1922) 5; Baker/. J. 
Bot. 62 (1924) Suppl. 7; Craib, Fl. Siam. En. 1 
(1925) 111; R. Keller in E. & P. Nat. Pfl. Fam. 
cd. 2, 21 (1925) 181 ; Makino, Syokubutu Dzukan 
(1925) 553, t. 628, cum f. yabei (Lev. & Van.) 
Making; Hochr. Candollea 2 (1925) 436, 437, 
incl. var. typicum Hochr.; Koidz. Bot. Mag. 
Tokyo 40 (1926) 344, incl. var. robusta MiQ. ex 
Koidz. I.e. 435: Ridl. Kew Bull. (1926) 59; Hind. 
Ciard. Bull. S. S. 4 (1928) 222; Koidz. Fl. Symb. 
Or. As. (1930) 92, incl. var. cavalcriei (Liv.) 
Koidz.; Hand.-Mazz. Symb. Sin. 7 (1 93 1) 404; 
Fyson, Fl. S. India Hill Stat. 1 (1932) 49, incl. var. 
major Fyson; Burk. Diet. (1935) 1217; Kanjii al 
& Das, FI. Assam I, I (1935) 102; Y. Kimura, Bot. 
Mag. Tokyo 54 (1940) 87; Corner, Wayside Trees 
(1940) 324; Gagnfp. FI. G6n. I.-C. Suppl. I (1^43) 
250; Masamune, Trans. Nat. Hist. Soc. I orniosa 
33 (1943) 168, incl. var. kainantcnsc Masam.; 
Banerji. J. Homb. Nat. Hist. Soc. 51 (1953) 774; 
Hknd. Malayan Wild H. (1954) 34, f. 24; Hundi lY 
& Kg Ko, List trees etc. Burma cd. 3 (l')6l) 19; 
Back. & Baku./ Fl. Java I (1963) 382; Oiiwi. II. 
Jap. (Engl, transl.) (1965) 631 ; Banirji, Rcc. Hot. 
Surv. Ind. 19(1966) 27; Siirf.stha, Hull. Dcp. Med. 
PI. Nepal 1 (1967) 7; .Sti-fn. Mt I"I. Java (1972) t. 
23: 7. — Ascvrum humifusum I, Attn i. Nov. I loll. 
I'l. Sp. 2 (1806) 33. t. 175. - - //. pusillum ( iioisv. 
Prod. Monogr. Hyp<ir. (1821) 50; in DC. Prod. 1 
( I 824) 549. -- //. campestrc MfX)N, ( at. Ind. Ixol. 
PI. Ceylon (1824) 56. nomen. — //. dichotomum 
Biicm.-Ham. ex I). Don. Prod. M. Nepal. (IK25) 
219 in svnon. /iralhvs humifusa (I. a mil.) 

Spacii. Ann. Sc. Nat. II. 5 (1836) 367. — India 


Flora Malesiana 

[ser. I, vol. 81 

frankenioides Korth, Tijd. Nat. Gesch. Phys. 3 
(1836) 17, t. 1. — Brathys japonica (Thunb. ex 
Murray) Wight, 111. Ind. Bot. 1 (1840) 113; Bl. 
Mus. Bot. Lugd. Bat. 2 (1856) 19, incl. var. 
accumbens Bl.; Miq. F1. Ind. Bat. 1, 2 (1859) 513, 
514, incl. var. inucronisepala Miq. et var. acuti- 
sepala MiQ. — H. nervatum Hance in Walp. Ann. 
2 (1851) 188. — H. sumatranum Miq. PI. Jungh. 
(1855) 395. — Brathys laxa Bl. Mus. Bot. Lugd. 
Bat. 2 (1856) 19. — Brathys nepalensis Bl. I.e. — 
Brathys debilis Bl. I.e. 20. — Brathys radicans Bl. 
I.e. — Brathys caespitosa Bl. I.e., inel. var. pusilla 
Bl. — Brathys oryzetum Bl. I.e. — H. ealycatum 
Jacquem. ex Dyer, F1. Br. Ind. 1 (1874) 256, in 
synon. — H. thunbergii Franch. & Sav. En. PI. 
Jap. 2 (1878) 300; Lev. Bull. Soc. Bot. Fr. 53 
(1906)498,501. —H.mutilum(nonh.)MAxm. Mel. 
Biol. 11 (1882) 171; Bull. Ac. Sc. St. Petersb. 27 
(1882) 436; Staff, Trans. Linn. Soc. Lond. II, 
Bot. 4 (1894) 132; Gibbs, J. Linn. Soc. Lond. 42 
(1914) 59; Arfak (1917) 149; Merr. J. Mai. Br. R. 
As. Soc. 1 (1923) 53; Steen. Bull. Gard. Bot. Btzg 
III, 13 (1934) 220. — H. yabei Lev. & Vant. Bull. 
Soc. Bot. Fr. 53 (1906) 498, 501. — H. taquetii 
Lev. & Van. in Fedde, Rep. 5 (1908) 279. — H. 
cavaleriei Lev. Bull. Soc. Bot. Fr. 54 (1908) 593. — 
H. dominii Lev. I.e. — H. pseudo-japonieiun 
Nakai, Bot. Mag. Tokyo 27 (1913) 130, nomen. — 
H. laxum (Bl.) Koidz. Bot. Mag. Tokyo 40 (1926) 
344; Masamune, F1. & Geobot. Stud. Ins. Yakusi- 
ma (1934) 305, inel. var. hananoegoense Masam. ; 
Y. KiMURA, Bot. Mag. Tokyo 51 (1937) 737; 
Hatusima, Bot. Mag. Tokyo 56 (1942) 571, inel. 
var. novo-guineense Hatus.; Ohwi, F1. Jap. (Engl, 
trans!.) (1965) 630. — Sarothra japoniea (Thunb. 
ex Murray) Y. Kimura in Nakai & Honda, Nova 
Fl. Jap. 10 (1951) 235, t. 78, inel. f. vulgaris Y. 
Kimura, I.e. 240, f. microphylla (Miq.) Y. Kimura, 
I.e., et f. robusta (Miq. ex Koidz.) Y. Kimura, 
I.e. — Sarothra laxa (Bl.) Y. Kimura in Nakai & 
Honda, Nova Fl. Jap. 10 (1951) 241, t. 79, incl. f. 
repens Y. Kimura, I.e. 244, /. simplex Y. Kimura, 
l.c.,f. ramosa Y. Kimura, l.c.,f. erecta Y. Kimura, 
I.e. 245, /. hananoegoensis (Masam.) Y. Kimura, 
I.e., et f. ramosissima Y. Kimura, I.e. 

Annual (? or perennial) herb, 2^5(-50) cm, 
with stems erect to decumbent or prostrate, un- 
branched below the inflorescence or also branched 
from the base or from various parts of the stem, 
with branches strict or ascending, rooting at the 
base, 4-lined. Leaves sessile, 2-18 by 1-10 mm, 
ovate to oblong or elliptic or subcircular or oblong- 
lanceolate or more rarely oblanceolate or obovate- 
spathulate, apex obtuse to rounded, base cordate- 
amplexicaul to cuneate or more rarely attenuate, 
sometimes glaucous beneath; 1-3 main basal veins, 
not or scarcely branched, often with 1 or more 
lateral veins; reticulate venation absent; laminar 
glands pale, not prominent, small, denser above 
than beneath; intramarginal glands absent. 
Inflorescence \-c. 30-flowered,cymose and regularly 
dichasial to monochasial or with branches in 
uppermost leaf axils, sometimes with subsidiary 
inflorescences from the next leaf pairs or with 
branching becoming sympodial and flowers 
apparently axillary, lax; pedicels exceeding upper- 
most leaves or not, (1.4-)2-14mm in fruit. 
Flowers 4-8 mm 0, plane; buds cylindric-ellipsoid, 
± obtuse. Sepals 2-5'/^ by M-2 mm, free, imbricate. 

narrowly oblong or (rarely) lanceolate to elliptic 
or obovate, acute or obtuse to rounded, entire, 
sometimes shortly mucronate, 3-5-nerved, often 
with ± prominent midrib, especially in fruit; 
laminar glands pale, linear towards the base, 
becoming punctate distally; marginal glands absent. 
Petals pale or bright yellow to orange, 1 .7-5 by 
0.8-1.8 mm, e. 0.9-1.3 as long as the sepals, 
elliptic or oblong to obovate, entire, persistent; 
apiculus absent or vestigial, lateral; laminar and 
marginal glands absent. Stamens not in apparent 
fascicles but forming 5 irregular groups when few, 
5-30, longest 1 [4-2.8 mm, H-H as long as the 
petals; anthers yellow, gland amber. Ovary 1- 
l'/^(-1.8) mm, ± broadly ovoid to subglobose; 
styles (2-)3, 0.4-0.8(-l) mm, -A-^^ as long as ovary, 
± divergent, gradually broadening towards 
capitate stigmas; placentas (2-)3, parietal. Capsule 
(2-)2^2-6 by 1.3-2.8 mm, cylindric to globose, not 
vittate. Seeds yellow-brown, c. ^A mm, cylindric, 
not carinate, longitudinally ribbed with fine 
transverse striae. 

Distr. Japan, S. Korea and SE. China to 
Ceylon, Australia, New Zealand and Hawaii; 
throughout Malesia. 

Ecol. Wet or marshy habitats (rice-fields, 
ditches, stream margins) to dry localities (short 
grassland, roadsides), but always in exposed places, 
0-3400 m. 

Notes. H. japonicum is an extremely variable 
species; but the variation appears to be continuous, 
so that, despite the long list of published segregate 
species or infraspecific taxa, it seems impossible to 
distinguish infraspecific categories. The following 
numbered variants, however, will give some idea of 
the nature of this variation: 

(1) Stems erect, simple. Leaves ± broadly ovate, 
subacute to obtuse, cordate-amplexicaul. 
Inflorescence terminal only, regularly dichasial 
at first. Bracts linear. Sepals oblong, acute. 
Petals as long as sepals. Stamens numerous 
(25-30). Capsule ± cylindric, shorter than 
sepals (Korea, China, Philippines). 

(2) As type 1, but with subsidiary inflorescences 
and branches, and often somewhat decumbent 
habit (S. Japan, Korea, China, India (Assam, 
Madras), Ceylon, Thailand, Malaya, Sumatra, 
Java, Celebes, Borneo, Philippines). 

(3) As type 1, but inflorescence branching wholly 
monochasial and stems always ± decumbent 
(Vietnam, Malaya, Sumatra, Java, Borneo, 

(4) Differs from type 1 in having some or all the 
bracts foliar and the habit nearly always 
decumbent to prostrate, often with diffuse 
branching. This is found throughout the area 
of the species. The shape of all parts is very 
variable, but one characteristic form with 
obovate outer sepals {var. 'calyeulatum') occurs 
in the Himalaya. Another characteristic form — 
prostrate with radiating branches, small, 
obovate to circular leaves and obovate or 
broadly elliptic sepals — occurs in the western 
half of Java (Priangan Mts & Dieng). 

(5) The stem-branching in this decumbent to 
prostrate delicate diffuse variant is sympodial 
(resulting in pseudo-axillary flowers) and the 
sepals narrowly elliptic to narrowly oblong or 
oblanceolate (var. humifuswri'); the size of all 


Hypericaceae (Robson) 


parts is small and the stamens few in number 
(5-10), and the subglobose capsules exceed 
the sepals (New Guinea, Tasmania, New 

H. japonicum has been confused with H. 
mutilum L., a species mainly of eastern North 
America that differs from H. japonicum in several 
characters, e.g. the branches are more widely 
spreading (40-70^ from vertical rather than 20- 
40"), those of the inflorescence being more slender 
than the stem and main branches (not equally 
stout), whilst those at the uppermost pair of nodes 
are usually more or less congested and often sub- 
umbellate (not roughly equally spaced); also, the 
bracts and bracteoles are nearly always linear- 
subulate and about 1-1 H mm long (not broader 
and 'or 2-7 mm long). 

Hypericum chinense Osbeck, Dagbok Ostind. 
Resa (1757) 244. 
This name predates H. chinense L. (1759), as 

Merrill (Am. J. Bot. 3, 1916, 588) pointed out. 
Merrill, however, thought that Linnaeus and 
OsBECK had described the same species, whereas 
Osbeck's description is of a herb with quadrangular 
stems, and therefore his name cannot be applied to 
the Linnaean species. If H. chinense Osbeck is a 
Hypericum, as seems likely, then it is almost 
certainly an earlier name for H. japonicum Thunb. 
ex Murray. In the absence of a specimen, however, 
and in the interest of nomenclatural stability, it 
should be regarded as a nomen dubium. 

Hypericum consimile R. Keller, Bot. Jahrb. 33 
(1904) 552. 

This species was described as coming from Java 
and belonging to sect. Androsaemum (Dun.) 
GoDR., which is otherwise confined to Europe, 
W. Asia, N. Africa and Macaronesia. It is said to 
be close to H. elatum Ait. {i.e. H. inodorum 
Mill.), which is the hybrid H. hircinum L. x H. 
androsaemum L. The type specimen has not been 
located, but, if it does come from Java — and is 
correctly described — then it is almost certain to be 
an escape from cultivation. 

New York Botanical Garden Library 

3 5185 00275 8199 


under the Auspices of 

Lembaga Biologi Nasional (Botanic Gardens of 

Indonesia), Bogor, Indonesia, 

and the Rijksherbarium, Leyden, Holland, 

executed by Foundation Flora Malesiana 

Scientific Communications 

concerning the Flora Malesiana should be addressed to 

the General Editor, Dr. C. G. G. J. van Steenis 




P.O. Box 26, Leyden, The Netherlands 

For sale only - Not for exchange 

Series I - Spermatophyta (Flowering Plants) 

Volume 1. Cyclopaedia of collectors & collections. 1950. pp. clii + 639 
Volume 2. Malesian vegetation (In preparation) 

Volume 3. Malesian plant geography (In preparation) 

Volume 4. General chapters and revisions. 1948-1954. pp. ccix + 631 
Volume 5. Bibliography, specific delimitation, & revisions. 1955-1958. pp. 

cccxlii + 596 
Volume 6. Systematic revisions. 6 parts. 1960-1972. pp. 20 + 1023. 
Volume 7. Systematic revisions. Parts 1-3. 1971-1974. pp. 753 
Volume 8. Cyclopaedia of Collectors, Suppl. 2. Systematic revisions. Part 1 



ISBN 90 286 0164 3 
LCCC No. 72-175112