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JOURNAL 



OF THE 



TDfto gopfe 6jnforaoIogiraI #orirtg< 



Vol. XXVIII. March, 1920. No. 1. 



RECORDS AND DESCRIPTIONS OF NEOTROPICAL 
CRANE-FLIES (TIPULID^, DIPTERA). II. 

By Charles P. Alexander, 
Urbana, III. 

The crane j flies discussed in the present article are almost without 
exception from South America. They are based largely on collect- 
ions made in 1919 near Para, Brazil, by Mr. Herbert S. Parish, the 
veteran insect collector of the American Tropics. Several other 
interesting species were sent me by Senor Charles Bruch, collected 
near La Plata, Argentina, and a few others taken by Dr. Witte in 
the Province of Santa Cruz, Southern Patagonia. The types are 
preserved in the collection of the writer; paratypes of several Ar- 
gentinian species in the Museum at La Plata. 

Genus Geranomyia Curtis. 
Geranomyia argentinensis new species. 

Head gray, the genae more buffy; mesonotum dull yellow, the prsescutum 
with three very broad brown stripes that are nearly confluent; femora with 
an indistinct, narrow, brown subterminal ring; wings subhyaline, the stigma 
dark brown, vein Se long, the basal deflection of Cu t before the fork of M. 

Male. — Length (excluding rostrum), 5.8-6 mm.; wing, 6.3-6.8 mm.; 
rostrum, 3.2-3.4' mm. 

Female. — Length (excluding rostrum), 7.5 mm.; wing, 7.5 mm.; rostrum, 
3.3 mm. 

1 



2 Journal New York Entomological Society. [Vol. xxviii. 

Rostrum dark brown, rather elongate, the tips of the paraglossas compara- 
tively short. Antenna with the scapal segments brown, the flagellum dark 
brown; basal flagellar segments oval, the terminal segments cylindrical. Head 
pale reddish, heavily gray pruinose, the gense buffy. Vertex between the eyes 
rather narrow. 

Mesonotum dull yellow, the prsescutum with three very broad brown 
stripes that are practically confluent; scutum with the lobes largely dark 
brown ; scutellum dark brown, broadly margined with yellow or, in some cases, 
entirely yellow ; postnotum dark brown. Pleura uniformly dull yellow, the 
mesonotum whitish pruinose. Halteres brown, the base of the stem paler. 
Legs with the coxae and trochanters yellow; femora pale yellowish brown, the 
tips a little brighter and with a very indistinct and narrow brown annulus; 
tibiae light brown, the tips narrowly darker; tarsi dark brown. Wings sub- 
hyaline ; stigma oval, dark brown ; veins dark brown. Venation : Sc long, Sc t 
extending to about opposite two thirds the length of the long sector, Sc 2 not 
far removed from the tip of Sc^ ; a supernumerary crossvein in cell Sc a short 
distance before the origin of the sector ; Rs long, about three times as long as 
the basal deflection of i? 4+B ; basal deflection of Cttj some distance before the 
fork of M. 

Abdominal tergites dark brown, the lateral margins of the tergites more 
yellowish ; sternites yellowish. Male hypopygium with the ventral pleural ap- 
pendage large, greatly exceeding the pleurite9; two acute needle-like points on 
either side at the base of the ventral pleural appendage. 

Habitat. — Argentina. 

Holotype, J 1 , Province of Buenos Aires (C. Bruch). 
Allotype, J. 
Paratopotypes, 5 $, 1$. 

Geranomyia argentinensis belongs _to the group of G. canadensis 
(Westwood) and is most nearly allied to G. guatetnalensis Alexander 
from which it may be told by the shorter rostrum, the different tho- 
racic and abdominal pattern, the less distinct brown femoral ring, 
and other details. 

Genus Dicranomyia Stephens. 
Dicranomyia amazonica new species. 

Size large (wing of $ over 8 mm.) ; general coloration black, the prsescu- 
tum reddish with a shiny black median stripe ; wings grayish brown, the costal 
region and narrow seams along the cord darker brown, Sc long, cell 1st M a 
small, subquadrate. 

Male. — Length, %8 mm. ; wing, 8.5 mm. 

Rostrum and palpi dark brown. Antennae black throughout, the last seg- 
ment elongate subulate. Head dark brown, gray pruinose, the vertex narrowed 
between the eyes, the latter with large coarse ommatidia. 



March, 1920.] ALEXANDER : NEOTROPICAL CRANE-FLIES. 3 

Mesonotal praescutum reddish with a conspicuous black median stripe that 
is broadest and most distinct in front, becoming obliterated before the suture ; 
scutum, scutellum and postnotum black. Pleura mostly black, the mesepimera 
a little paler. Halteres dark brownish black. Legs with the coxae dull reddish, 
the fore coxae black; trochanters dull reddish; femora black, the bases nar- 
rowly reddish; remainder of the legs black. Wings grayish brown, the wing- 
base indistinctly brighter; the costal region, stigma, tip of the wing and narrow 
seams along the crossveins and deflections of veins dark brown ; veins dark 
brown, R, Cu and 2d A more yellowish. Venation : Sc long, extending to be- 
yond midlength of the long sector, Sc 2 at the tip of Sc t ; r at the tip of R lf 
longer than the apex of i? t beyond it ; cell 1st M 2 small, subquadrate ; basal 
deflection of C« a before the fork of M ; Cu 2 shorter than the deflection of C« t . 

Abdominal tergites black, the caudal margins of segments two to five 
broadly but indistinctly paler; sternites reddish brown. Hypopygium conspic- 
uously reddish orange ; pleurites elongate, much longer than the relatively small 
appendages ; peni9-guard conspicuous, about as long as the pleurites. 

Habitat. — Brazil. 

Holotype, J 1 , Igarape-Assu, Para, July 16, 1919 (H. S. Parish). 

Paratype, J 1 , Obidoo, September 10, 1919 (H. S. Parish). 

Dicranomyia sanctae-cruzae new species. 

General coloration reddish yellow ; antennal flagellum dark brown ; legs 
yellow, only the terminal tarsal segments darker; wings yellowish subhyaline 
with sparse dark brown marks at the stigma, along the cord, at crossvein m. 
and on the 2d Anal vein. 

Male. — Length, 8.4 mm.; wing, 12 mm. 

Female. — Length, 9-9.3 mm.; wing, 11. 5-12.6 mm. 

Rostrum and palpi light yellow. Antennas with the scapal segments brown- 
ish yellow, the flagellar segments dark brown, the intermediate flagellar seg- 
ments oval in shape. Head reddish brown with a microscopic dense white 
pubescence, the median line of the vertex darker. Eyes rather widely sepa- 
rated by the vertex. 

Thorax reddish yellow, the mesonotal praescutum with very indistinct 
darker reddish stripes. Pleura reddish brown. Halteres pale. Legs with the 
coxae and trochanters dull yellow; remainder of the legs pale yellow, the two 
terminal tarsal segments dark brown; last tarsal segment longer than the 
penultimate ; claws with three teeth. Wings subhyaline, the yellow veins sur- 
rounded by a pale yellow tinge ; stigma brown, connected with a brown seam 
at the fork of Rs; Cu 2 and the basal deflection of C« t seamed with brown; a 
large brown area on crossvein m; the distal half of vein 2d A with a large 
dark brown seam. Venation : Sc x ending opposite the origin of Rs, Sc 2 a short 
distance back from the tip so that Sc t alone is a little longer than r; basal 
deflection of i? 4+5 about one half the length of the feebly sinuous sector; cell 
1st M 2 closed ; basal deflection of Cu t at or some distance before the fork of M. 

Abdominal tergites reddish brown, the sternites yellowish, the subterminal 



4 Journal New York Entomological Society. t Vo1 - xxvin. 

sternites a little darker. Male hypopygium rather large, the pleurites slender, 
on the ventral inner face produced proximad into a prominent lobe that is 
covered with appressed hairs ; ventral pleural appendage large. Ovipositor 
with the valves nearly equal in length, or the tergal valves a little shorter than 
the sternal valves, the tips acute; the tergal valves are blackened and with the 
- dorsal edges minutely serrate. 

Habitat. — Argentina (Patagonia) ; Tierra del Fuego. 

Holotype, <$, Valle Tunel, Santa Cruz (Dr. Witte). ' 

Allotopotype, J. 

Paratopotype, <£; paratype, J, Tierra del Fuego (Ohlin). 

Dicranomyia sanctce-crusa is a large, vigorous species that re- 
sembles D. chlorotica (Philippi), but is readily told by the hand- 
somely spotted wings. 

The paratype is in the Reichsmuseum in Stockholm. 

Genus Rhipidia Meigen. 
Rhipidia (Rhipidia) costaloides new species. 

Antennae black; general coloration black, the anterior part of the praescu- 
tum more reddish brown ; wings pale gray with a grayish brown pattern along 
the veins. 

Female. — Length, 4.8 mm.; wing, 5.3 mm. 

Rostrum and palpi dark brownish black. Antennae black throughout. 
Head dark. 

Mesonotal praescutum reddish brown, becoming darker posteriorly, the re- 
mainder of the mesonotum likewise dark. Pleura dark, the sternum more yel- 
lowish. Halteres pale brown, the knobs darker. Legs with the coxae dark 
basally ; trochanters brown ; femora brown ; remainder of the legs broken. 
Wings pale gray subhyaline with a moderately heavy grayish brown pattern as 
follows : six costal blotches that are approximately as large as the interspaces 
between them, the first at the arculus, the third at the origin of Rs, the fifth 
subocellate, at the stigma, the sixth at the end of vein i? 2+3 ; broad paler gray 
seams along the cord and outer end of cell 1st Af a . Venation : Sc ending just 
beyond midlength oi Rs; r at the tip of i? t ; deflection of i? 1+5 about as long as 
cell 1st M 2 ; inner end of cell 1st M 3 not conspicuously arcuated ; basal deflec- 
tion of C% some distance before the fork of M. 

Abdominal tergites reddish brown, the caudal margin of the segments 
broadly dark brown ; sternites dull yellow, the segments conspicuously ringed 
with blackish brown as an annulus on the caudal margin of the segments and 
on the cephalic margin of the following segment ; genital segment dull yellow. 
Ovipositor with the valves short but heavily chitinized ; tergal valves slender, 
strongly upcurved at the acute tips ; sternal valves long, ending about on a 
level with the tergal valves, straight, the tips acute. 

Habita t. — Panama. 



March, 1920.] Alexander : Neotropical Crane-Flies. 5 

Holotype, J, Quebrada Richa, a branch of the Rio Tapaliza, a 
branch of the Rio Puero, altitude 1,300 feet, July 24, 1918 (Axel 
Olsson). 

This small species is closest to Rhipidia costalis Williston of Cen- 
tral America and the Lesser Antilles. In order to supplement Wil- 
liston's brief description, which applies equally well to more than 
one species in the Neotropical fauna, Mr. W. D. Lang, of the British 
Museum of Natural History, has very kindly sent me an enlarged 
accurate drawing of the wing of the type specimen of R. costalis. 
1 This figure agrees in all the essentials with the fly determined by the 
writer as costalis in an earlier paper (Journ. N. Y. Ent. Soc, Vol. 
22, pi. 3, fig. 2; 1914), the dark markings along the costal margin 
being considerably larger than the pale interspaces, the radial sector 
very long, the basal deflection of J? 4t5 comparatively short, basal de- 
flection of C«j just beyond the fork of M and other characters. In 
Mr. Lang's figures, cell first M„ is shorter and broader than in the 
specimen above mentioned, if. costaloides is readily told by the 
reduced costal pattern and the venational details. 

Genus Teucholabis Osten Sacken. 
Teucholabis mendax new species. 

General coloration reddish, the praescutum with three broad black stripes; 
head shining black ; pleura with large black blotches ; legs yellow throughout ; 
wings subhyaline, the apex and a broad seam along the cord slightly brownish. 

Male. — Length, 7 mm. ; wing, 6.4 mm. 

Rostrum pale brown, the palpi dark brown. Antennae dark brown, the 
basal segments only a little paler. Head intense shiny black. 

Pronotum shiny dark brown, the scutellum more reddish. Mesonotal prae- 
scutum reddish yellow with three intense black stripes that are very extensive, 
the interspaces restricted; median stripe ending before the suture; lateral 
stripes very broad, nearly circular in outline, continued caudad onto the scutal 
lobes; remainder of the mesonotum reddish, the postnotum with a distinct 
median and less distinct lateral stripes on the caudal half. Pleura orange- 
yellow with large jet black areas, smallest on the propleura ; two large blotches 
on the mesopleura, the largest on the mesepisternum, extending from the ster- 
num to the dorsopleural membranes; a smaller oval area immediately cephalad 
of the halteres. Halteres small, reddish, the stems a little darker. Legs dull 
yellow throughout, the femora somewhat thickened. Wings subhyaline with 
the apex and a broad diffuse seam along the cord very indistinctly darker; 
stigma dark brown; veins dark brown, the basal and costal regions more yel- 
lowish. Venation : Sc rather long, extending to about midlength of the sector ; 



6 Journal New York Entomological Society. t Vo1 - xxvm. 

r inserted a short distance beyond the line of the cord ; cell ist M 2 long and 
narrow, about as long as vein M 1+2 beyond it, longer than M 3 beyond it; basal 
deflection of C% at about one fourth the length of oell 1st Af 2 . 

Abdomen reddish. Hypopygium weakly armed; an outer flattened pale 
blade that bears a few long setae ; a long-curved appendage that terminates in 
a blackened needle-like point ; an inner, bifid heavily chitinized appendage with 
the anterior blade flattened, the edge indistinctly two-toothed, truncated or 
slightly concave; penis-guard stout, the tip blackened. 

Habitat. — Brazil. 

Holotype, <$, Prata, Para, June 30, 1919 (H. S. Parish). 
Teucholabis mendax is readily told from all its relatives by the 
unusually pale banded wings and the shining black head. 

Teucholabis persimilis new species. 

General coloration light brown, the pleura yellow, wings yellow, crossvein 
r lacking, veins issuing from cell 1st M 2 long. 

Female. — Length, 4.6 mm.; wing, 4.5 mm. 

Rostrum pale brown, the palpi darker. Antennae unusually long for this 
genus of flies, if bent backward extending to beyond the wing-root ; brown, the 
basal segment a little paler j flagellar segments long-cylindrical with long ver- 
ticils and a dense white pubescence. Head brownish yellow, sparsely, white 
pruinose ; vertex between the eyes broad. 

Pronotum yellowish medially, the scutellum more whitish, the pleural 
regions more brownish. Mesonotum light brown, sparsely pollinose, the 
median area of the scutum and postnotum more yellowish. Pleura yellow, the 
dorsal pleurites somewhat darker. Halteres brown, the knobs dark. Legs dull 
yellow, the terminal three or four tarsal segments dark brown; claws small, 
untoothed, empodia conspicuous. Wings with a strong yellowish tinge ; stigma 
indistinct, grayish ; veins brownish yellow. Venation : Sc rather long, Sc t very 
long, extending to beyond midlength of the long sector, Sc 2 far from the tip 
of Sc ± and slightly proximad of the origin of Rs; r lacking; veins R^., and 
i? 4t5 long, curved caudad and running parallel at their ends; cell ist M s nar- 
row, shorter than the veins issuing from it ; basal deflection of Cu t a short dis- 
tance beyond the fork of M. 

Abdominal tergites dark brown, the sternites more yellowish. Ovipositor 
with the valves long and slender, strongly upcurved, the tips acute. 

Habitat. — Brazil. 

Holotype, J, Igarape-Assu, Para, July 15, 1919 (H. S. Parish). 

This curious little fly is referred to the genus Teucholabis with 
considerable doubt. The radial cross-vein is completely lacking and 
vein Sc 1 is very long for a member of this genus of flies. T. per- 
similis bears a great resemblance to T. parishi Alexander (Psyche, 



March, 1920.] ALEXANDER : NEOTROPICAL CrANE-FlIES. 7 

Vol. 20, pp. 46, 47; 1913) which must likewise be considered a doubt- 
ful member of the genus. The present species may be told by the 
long antennas and the details of venation, especially the shorter sub- 
costa, the lack of a spur of vein i? M and the shorter cell first M 2 with 
the longer veins issuing from it. That the spur on J? M in T. parishi 
is a normal condition for this species is shown by the fact that it 
bears a macrotrichia on this spur in both wings of the unique type, 
this undoubtedly representing the origin of vein R 2 . 

Genus Gonomyia Meigen. 
Gonomyia (Gonomyella) paraensis new species. 

Antenna with the basal segments light yellow, the flagellum brown ; head 
yellow, silvery pollinose ; mesonotum lead-colored, the lateral margins narrowly 
light yellow; wings strongly tinged with brownish yellow, cell i? 2 very short, 
cell M 2 very deep. 

Male. — Length, about 4 mm.; wing, 4.5 mm. 

Female. — Length, 4.5 mm. ; wing, 4.7 mm. 

Rostrum and palpi dark brown. Antenna with the first scapal segment 
light yellow, stout, moderately elongated ; second segment tumid, dull yellow ; 
flagellum slender, brown, the segments with a dense white pubescence, in the 
male with long, secund verticils. Head light yellow, darker on the occiput, 
covered with a sparse silvery white pollen that is most distinct on the anterior 
part of the vertex. 

Pronotum yellow medially, the sides dark. Mesonotal prsescutum dark 
lead color, sparsely pollinose, the lateral margin narrowly but conspicuously 
light yellow, two indistinct brownish stripes occupy the region of the inter- 
spaces; remainder of the mesonotum dark plumbeous. Pleura dull yellow, 
with a broad, ill-defined, dusky, dorsal stripe. Halteres dark brown. Legs dull 
yellow, darkening on the tibiae and tarsi, the latter dark brown. Wings strongly 
brownish yellow, the costal region more saturated ; stigma indistinct ; veins 
brown, the costal veins more yellowish. Venation : Sc t ending slightly before 
the fork of Rs; fusion of R i+S very long, the cell i? 2 being shorter than in any 
other described American species of Gonomyella; r at about two fifths the 
length of i? a+3 ; cell M 2 very deep; basal deflection of Cu t just beyond the 
fork of M. 

Abdominal tergites dull yellowish brown, the sternites more yellowish ; 
hypopygium dull brownish yellow. In the female the antennas are shorter, 
the abdomen dark brown. Ovipositor horn-colored, the tergal valves very long, 
slender, upcurved at the tip. 

Habitat. — B razil. 

Holotype, <£ Prata, Para, June 30, 1919 (H. S. Parish). 

Allotype, 2- 



8 Journal New York Entomological Society. C Vo1 - xxviii. 

The holotype male has the wings very much darker brown than 
in the female, the costal margin being especially suffused; in addi- 
tion, the venational details are somewhat different, r being placed 
before one third the length of J? M ; R a considerably longer^ slightly 
more than one half the length of R 3 ; basal deflection of Cu t some 
distance beyond the fork of M. With the exception of these rather 
conspicuous alar differences, the two flies are very similar to one 
another and until further material becomes available they must be 
considered as representing a single variable species. 

Genus Erioptera Meigen. 
■Erioptera (Erioptera) micromyia new species. 

Size very small (wing of 5 under 3 mm.) ; general coloration pale brown, 
the thoracic dorsum unstriped ; wings grayish; ovipositor very long and slender. 

Female. — Length, 3.3 mm. ; wing, 2.8-2.9 mm. 

Rostrum yellowish brown ; palpi dark brown. Antennae moderately long 
and slender, brown. Head light brown, sparsely gray pruinose. 

Mesonotum light brown, the praescutum without stripes ; tuberculate pits 
black, separated by a distance about equal to one and one half times the diam- 
eter of one. Pleura light brown, heavily light gray pruinose. Halteres with 
the knobs large, dark brown, the base of the stems pale. Legs with the coxae 
small, pale testaceous, sparsely gray pruinose ; trochanters testaceous ; re- 
mainder of the legs brown. Wings grayish subhyaline; veins dark brown. 
Venation : As in the subgenus Erioptera ; 2d anal vein moderately sinuate on 
its outer third. 

Abdomen with the tergites dark brown ; sternites more yellowish. Ovi- 
positor with the valves very long and slender, acicular, gently upcurved ; 
sternal valves similar in appearance to the tergal valves but more elongate, 
ending but a short distance before the tips of the tergal valves. 

Habitat. — Brazil. 

Holotype, J, Prata, Para, June 30, 1919 (H. S. Parish). 

Paratopotypes, 2 J's. 

Genus Ozodicera Macquart. 
Ozodicera attenuata new species. 

Antennal segments four to nine with a single short pectination; segments 
ten to thirteen very long and attenuated ; dorsum of the praescutum with four 
brown stripes ; lateral margins of the abdominal tergites brown. 

Male. — Length, about 20 mm.; wing, 16.5 mm. 

Frontal prolongation of the head reddish, shiny, narrowly darker dorso- 
medially; nasus short; palpi black. Antennae rather long for this genus of 



March, 1920.] ALEXANDER : NEOTROPICAL CrANE-FlIES. 9 

flies, the three basal segments orange, thence passing into black ; segments four 
to nine each bears a single short pectination which is shorter than the seg- 
ment that bears it; the last four flagellar segments are greatly elongated, 
filiform, their combined length considerably exceeding the remainder of the 
antenna. Head yellowish tawny, the vertex very narrow between the large eyes. 

Mesonotal praescutum yellowish tawny with four dark reddish brown 
stripes; remainder of the mesonotum brown, sparsely pruinose, especially the 
postnotum. Pleura light reddish brown, sparsely white pruinose. Halteres 
brown, the knobs a little darker. Legs with the coxae rtddish brown, gray 
pruinose ; trochanters and femora dull brownish yellow ; tibia: yellowish brown ; 
tarsi dark brown, very long and slender. Wings light gray, the base of the 
wing and the costal region more brownish yellow; stigma elongate, grayish; 
veins yellowish brown, C, Sc and Cu more yellowish. Venation: tip of R t 
pale; r-m very short to almost obliterated by the approximation of veins 
R t +s and M 1+2 ; cell M t broadly sessile; m-cu obliterated by the punctiform 
contact of C«j on M 3+i . 

Abdominal tergites yellow, the lateral margins of the segments broadly 
brown, this mark widening on the fourth and succeeding segments to include 
most of the segments; sternites reddish yellow, segments seven and eight 
darker. Hypopygium reddish, the pleurites long, slender, bearing the com- 
pressed pleural appendages at their tips. 

Habitat. — Brazil. 

Holotype, J 1 , Prata, Para, July 5, 1919 (H. S. Parish). 

Ozodicera attenuata is closest to O. gracilis (Westwood) and 
O. griseipennis Loew. It is characterized by the very short antennal 
pectinations, the attenuated apical flagellar segments, the quadrivit- 
tate thorax and the dark lateral margins to the abdominal tergites. 
0. gracilis has the apical flagellar segments less elongated and the 
thorax subvittate; 0. griseipennis has the median prsescutal stripe 
entire and the abdomen with a dark brown dorso-median stripe. 

Pectinotipula new genus. 

Antennas 13-segmented, in the male with the terminal ten seg- 
ments each with two long pectinations that exceed in length the seg- 
ments that bear them; flagellar segments verticillate. Venation 
with cell first M 2 small as in Tipula; cell M, long-petiolate. Male 
hypopygium as in Tipula, the ninth tergite, pleurite and sternite being 
entirely distinct; eighth sternite with a long, conspicuous median 
appendage. Coloration as in species of the monilifera group of the 
genus Tipula, the body with a capillary brown dorso-median line, the 
praescutum with the interspaces set with tiny brown setigerous 
punctures. 



10 Journal New York Entomological Society. [Vol. xxviii. 

Genotype. — Ozodicera argentina van der Wulp (Argentina). 

Enderlein (Zool. Jahrb., Vol. 32, pt. i, p. 27; 1912) places argen- 
tina in his restricted genus Ozodicera under the mistaken belief 
that the antennae are unipectinate. In reality, the antenna in the 
male sex, at least, are long-bipectinate and of a structure that di- 
verges widely from all species of the genus Ozodicera. Likewise in 
its wing-venation which is almost identical with the normal type of 
the genus Tipula and the simple male hypopygium this species de- 
parts from the compact type of Ozodicera. The presence of ten 
pectinate flagellar segments and the petiolate cell M 1 are ample char- 
acters upon which to remove this fly from Ozodicera to a position 
nearer the genus Tipula, to which it is obviously more nearly allied. 
The genitalia and color of the body closely resemble those of species 
of the monilifera group of the genus Tipula and it seems probable 
that the true affinities of this fly are not far distant from this group. 

I am indebted to Sefior Jorgensen for a specimen of this interest- 
ing fly, taken at Esquina Grande, Catamarca, Argentina, April 7, 

I9I5- 

Genus Tipula Linnaeus. 
Tipula bruchi new species. 

Antennae bicolorous ; mesonotal praescutum with dark brown stripes ; legs 
very long and slender, the tips of the femora and tibiae black; wings with the 
costal margin yellowish, the membrane clouded with gray and with four dark 
brown blotches in the subcostal cell. 

Male. — Length about 17-18 mm.; wing, 18-19.2 mm. 

Hind leg, femur, 14.2 mm.; tibia, 16 mm.; metatarsus, 21.6 mm. 

Frontal prolongation of the head slender, buffy, the sides darker; nasus 
slender with long yellow hairs. Antenna: short, the three basal segments dull 
yellow, the remaining segments yellow with the basal enlargement black; ter- 
minal segments more uniformly darkened ; verticils longer than the segments 
that bear them. Head yellowish buff with a long dark brown median mark on 
the vertex; vertical tubercle rather conspicuous, brown medially, with an im- 
pressed median furrow. 

Pronotum pale yellowish brown with three dark brown marks. Mesonotal 
praescutum light buff with three dark brown stripes, the median stripe divided 
by a broad pale line which, in turn, is split by a capillary dark brown median 
line ; humeral region dark ; scutum buffy gray, each lobe with two large dark 
brown marks; scutellum light brown with a dark brown median line; post- 
notum grayish, the posterior margin and median area more grayish, the latter 
with a capillary dark brown line. Pleura light buff, indistinctly spotted with 



March, 1920.] ALEXANDER: NEOTROPICAL CRANE-FLIES. 11 

darker. Mesosternum indistinctly dark gray. Halteres rather long, the knobs 
dark brown with the apices dull yellow. Legs with the coxae and trochanters 
brownish yellow i femora yellowish brown, the tips broadly black ; tibiae dark 
brown, the tips narrowly blackened ; tarsi very long, dark brown, the metatarsi 
longer than the tibiae. Wings subhyaline, with four dark brown subcostal 
blotches and a heavy grayish brown clouding in all the cells ; cells C and Sc 
strongly yellowish ; the first dark brown subcostal mark is at h and also in- 
cludes the bases of cells R and M and extends into the costal cell ; the re- 
maining three marks do not reach the costal cell but send paler clouds caudad 
into cell R; the third mark, at the origin of Rs, is the largest, stigma rather 
pale brown ; the grayish brown clouds include the apical half of cells R 2 and 
R 3 ; most of cells i? 5 to C« x , conspicuous zigzag areas in cells M, Cu and 1st A ; 
cell sd A more uniformly darkened. Venation: Veins R 2 and R 3 strongly di- 
vergent, the base of cell R 2 being very narrow ; cell 1st M„ much longer than 
broad ; m-cu distinct. 

Abdomen with the first tergite buffy with a dark brown median mark ; sec- 
ond tergite dark with only the basal half pale laterally; remaining tergites 
darker brown, indistinctly ringed caudally with paler; the lateral margins are 
marked with a brown streak; sternites similar to the tergites; ninth segment 
pale. Male hypopygium with the ninth tergite extensive, pale, with a deep 
and narrow V-shaped' median notch, the broad lateral lobes irregularly notched 
and toothed, the most lateral tooth largest. Ninth pleurite complete, the 
pleural appendages being long and slender ; outer appendage cylindrical, clothed 
with long, coarse yellow hairs ; inner appendage with the posterior margin 
beautifully fringed with long flattened hairs. Ninth sternite with the caudal 
inner angle with a pale, roughly oval lobe which is covered with a pale ap- 
pressed pubescence and whose ventral angle bears a pencil of long reddish 
hairs that are decussate with those of the opposite side across the genital 
chamber. Eighth sternite bearing a broad, flattened, heart-shaped median lobe 
that is provided with long, coarse, yellow hairs. 

Habitat. — Argentina. 

Holotype, £, Province of Buenos Aires (C. Bruch). 

Paratopotypes, 2 ^. 

Tipula bruchi is named in honor of Sefior Charles Bruch, to 
whom I am indebted for many favors. The species agrees fairly 
well with the description of T. nubifera van der Wulp, but the pat- 
tern of the thorax and wings is very different in the two species. 

Tipula wittei new species. 

Antennal flagellum indistinctly bicolorous ; mesonotum dark gray with five 
brown stripes, the three intermediate stripes capillary, the lateral stripes 
broader ; wings grayish, the stigma pale yellowish brown ; abdominal tergites 
brownish yellow with three dark brown stripes. 

Female. — Length, about 26 mm. ; wing, 19.2 mm. 



12 Journal New York Entomological Society. t Vo1 - xxvm. 

Frontal prolongation of the head pale brownish yellow, more pruinose 
above; palpi dark brown. Antennae with the first segment gray pruinose; sec- 
ond segment yellow; flagellar segments with the basal enlargement dark brown, 
the remainder of each segment a trifle paler to produce a very indistinct bi- 
colorous effect; flagellar verticils long. Head brownish gray with a very in- 
distinct brown line on the vertical tubercle. 

Pronotum brownish with an indistinct capillary dark brown median line. 
Mesonotal praescutum dark gray with five brown stripes, the three intermediate 
very narrow, being formed by the lateral margins and capillary median vitta 
of the usual broad median stripe ; lateral stripes broad ; humeral region yellow ; 
praescutal interspaces with a few dark setigerous punctures; scutum dark 
gray ; scutellum dark testaceous brown ; postnotum light gray with a capillary 
black median line. Pleura light yellow, gray pruinose; dorsopleural mem- 
branes dull yellow. Halteres long, dark brown, the base of the stem paler. 
Legs with the coxae pale reddish, sparsely gray pruinose ; trochanters dull yel- 
low ; femora reddish brown, the tips indistinctly darker ; tibiae yellowish brown, 
the tips narrowly dark brown ; metatarsi yellowish brown, the tips darkened ; 
remainder of the tarsi dark brown; metatarsi shorter than the tibiae; claws 
small, simple. Wings grayish, the apical cells grayish brown, the costal and 
subcostal cells more yellowish; stigma pale yellowish brown; an obliterative 
streak before and beyond the stigma, the latter in the bases of cells R, and R t . 
Venation: Cell R, rather broad at the base; basal deflection of R i+S short; 
cell ist M i small, almost regularly pentagonal ; petiole of cell M 1 short, about 
as long as cell Ist M 2 ; m-cu punctiform. 

Abdominal tergites brownish yellow with three broad dark brown stripes, 
the lateral margins of the segments rather broadly pale, the caudal margins 
indistinctly ringed with yellowish, most evident on segments six and seven; 
sternites pale yellow, the terminal segments indistinctly pruinose. Ovipositor 
with the long dorsal shield and the valves light chestnut brown, the straight, 
slender tergal valves greatly exceeding the sternal valves, along the ventral 
margin with scattered erect setae. 

Habitat.— Argentina (Patagonia). 

Holotype, <j>, Valle Tunel, Santa Cruz (Dr. Witte). 

Tipula philippiana new species. 

Antennae indistinctly bicolorous ; mesonotal praescutum dark brown with 
three broad bluish gray stripes that are narrowly and incompletely margined 
with pale gray and dark brown ; pleura with a broad brown dorsal stripe ; 
wings white, sparsely streaked with brown, the costal and subcostal cells more 
yellowish ; abdomen dull yellow, trivittate with dark brown. 

Female. — Length, about 23 mm.; wing, 19.5 mm. 

Frontal prolongation of the head reddish testaceous; palpi dark brown. 
Antenna; with the basal flagellar segments indistinctly bicolorous, the basal 
swelling brown, the terminal flagellar segments more unicolorous brown; ver- 



March, i 9 20.] ALEXANDER: NEOTROPICAL CrANE-FlIES. 13 

ticils conspicuous. Head light gray, more brownish adjoining the inner margin 
of the eyes. 

Mesonotal praescutum dark brown, the prasscutum stripes clear bluish gray, 
margined with dull whitish gray ; the median stripe is split by a capillary dark- 
brown line that broadens out behind, just before the suture enlarging into a 
blotch; immediately laterad of this and occupying the praescutal interspaces a 
similar short brownish line extending from the suture cephalad; interspaces 
sparsely dotted with brown setigerous punctures ; lateral margins of the median 
stripe at about midlength of the sclerite narrowly and indistinctly margined 
with brown ; lateral margins and the humeral region of the dark brown ground 
color; scutum dull gray, the lobes with a linear dark brown stripe; median 
lobe of the scutellum and the postnotum gray pruinose, the latter with an 
indistinct brown median line. Pleura brown, grayish pruinose; a dark brown 
longitudinal stripe on the dorsal pleurites ; a dark brown velvety spot imme- 
diately dorsad of the base of the halteres on the sides of the postnotum and a 
small white area immediately before the halteres ; dorso-pleural membranes 
dull yellowish. Halteres dark brown, the base of the stem and the base of the 
knob paler. Legs with the coxae pale, grayish pruinose ; trochanters and femora 
dull brownish yellow, the latter with an indistinct brownish ring immediately 
before their apices; tibiae yellowish brown, the tips darker; tarsi brown; meta- 
tarsi considerably shorter than the tibiae. Wings white, sparsely streaked and 
lined with brown ; cells C and Sc strongly yellowish ; stigma brown ; small dark 
brown marks at h, at the origin of Rs, midway between the two latter, and at 
the tip of Sc ; larger but slightly paler clouds at the fork of Rs, connecting 
with the stigmal spot, a large blotch in the end of cell R 2 and the middle of cell 
R 3 ; a large blotch at the base of cells R and M and along vein Cu in cells M 
and Cu for almost the whole length of this vein ; grayish clouds in the ends of 
the anal cells ; the ends of veins M t , M,, M 3 , Cu t and 1st A are seamed with 
brown. Venation : Base of cell i? 2 broad ; cell 1st M 2 subpentagonal ; m-cu 
obliterated by the fusion of Cu ± on M 3+1 ; cells R 2 and jl^ are somewhat broader 
than in T. wittei. 

Abdomen with the first tergite gray pruinose ; remaining tergites dull yel- 
low with three diffuse dark brown stripes that almost obliterate the ground 
color except at the base of the sclerites ; sternites dull yellow, sparsely 
pruinose. Ovipositor almost as in T. wittei but the dorsal basal shield less 
elongate. 

Habitat. — Argentina ( Patagonia ) . 

Holotype, J, Valle Tunel, Santa Cruz (Dr. Witte).